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<document ID-DOI="10.11646/zootaxa.5256.5.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="7758872" ID-ZooBank="E5B9708E-D626-4923-825A-A9FD59AE4024" checkinTime="1679468565563" checkinUser="plazi" docAuthor="Matalin, Andrey V. &amp; Wiesner, Jürgen" docDate="2023" docId="050B87D78D5B2470E1B819E1FA5C75E3" docLanguage="en" docName="zootaxa.5256.5.1.pdf" docOrigin="Zootaxa 5256 (5)" docSource="http://dx.doi.org/10.11646/zootaxa.5256.5.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Therates sauteri W. Horn 1912" docType="treatment" docVersion="1" lastPageNumber="431" masterDocId="F932FFAF8D40246EE12F1976FFC07601" masterDocTitle="Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)" masterLastPageNumber="433" masterPageNumber="401" pageNumber="428" updateTime="1679505860750" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title>Revision of the Therates fruhstorferi complex (Coleoptera, Cicindelidae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Matalin, Andrey V.</mods:namePart>
<mods:nameIdentifier type="ORCID">0000-0002-7790-8709</mods:nameIdentifier>
<mods:affiliation>Moscow State Pedagogical University, Department of Zoology &amp; Ecology, Kibalchicha str. 6, build. 3, Moscow 129164, Russia. andrei-matalin @ yandex. ru; https: // orcid. org / 0000 - 0002 - 7790 - 8709 &amp; Pirogov National Research Medical University, Department of Biology, Ostrovitianova str. 1, Moscow 117997, Russia.</mods:affiliation>
<mods:nameIdentifier type="email">andrei-matalin@yandex.ru</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Wiesner, Jürgen</mods:namePart>
<mods:affiliation>Dresdener Ring 11, D- 38444 Wolfsburg, Germany.</mods:affiliation>
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<mods:title>Zootaxa</mods:title>
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<mods:date>2023</mods:date>
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<taxonomicName authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" box="[151,537,151,178]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis bold="true" box="[151,347,151,178]" italics="true" pageId="27" pageNumber="428">Therates sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[396,537,151,177]" pageId="27" pageNumber="428" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
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, bona sp.
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<figureCitation box="[151,279,187,213]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="27" pageNumber="428">Figs 26–28</figureCitation>
,
<figureCitation box="[290,363,187,213]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="27" pageNumber="428">48, 49</figureCitation>
,
<figureCitation box="[374,447,187,213]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="27" pageNumber="428">77–80</figureCitation>
,
<figureCitation box="[458,558,187,213]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="27" pageNumber="428">109–114</figureCitation>
,
<figureCitation box="[569,670,187,213]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="27" pageNumber="428">173, 174</figureCitation>
.
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<treatmentCitation author="Horn, W." box="[151,699,223,250]" page="133" pageId="27" pageNumber="428" year="1912">
<taxonomicName ID-CoL="5F7VN" authority="W. Horn, 1912: 133" authorityName="W. Horn" authorityPageNumber="133" authorityYear="1912" box="[151,699,223,250]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="sauteri">
<emphasis box="[151,470,223,249]" italics="true" pageId="27" pageNumber="428">Therates fruhstorferi sauteri</emphasis>
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W.
<bibRefCitation author="Horn, W." box="[514,699,223,250]" pageId="27" pageNumber="428" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912: 133</bibRefCitation>
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(
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locality—
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,
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<taxonomicName ID-CoL="5F7VN" authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" box="[151,596,292,316]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="sauteri">
<emphasis box="[151,440,292,316]" italics="true" pageId="27" pageNumber="428">Therates fruhstorferi sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[482,596,292,316]" pageId="27" pageNumber="428" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
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—
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<bibRefCitation author="Horn, W." box="[622,786,291,316]" pageId="27" pageNumber="428" pagination="16 - 23" refId="ref25821" refString="Horn, W. (1915) Coleoptera Adephaga. Fam. Carabidae. Subfam. Cicindelinae. In: Wytsman, Ph. (Ed.), Genera Insectorum. 82 c. Bruxelles, pp. 209 - 487, Taf. 16 - 23." type="book chapter" year="1915">Horn 1915: 439</bibRefCitation>
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<treatmentCitation author="Horn, W." box="[798,902,291,316]" page="114" pageId="27" pageNumber="428" year="1926">
<bibRefCitation author="Horn, W." box="[798,902,291,316]" pageId="27" pageNumber="428" pagination="1902 - 1970" refId="ref25870" refString="Horn, W. (1926) Carabidae: Cicindelinae. In: Junk, W. &amp; Schenkling, S. (Eds.), Coleopterorum Catalogus. Pr. 86. L. DesmetVerteneuil [etc.], 1902 - 1970, Berlin, 345 pp." type="book chapter" year="1926">1926: 114</bibRefCitation>
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<treatmentCitation author="Miwa, Y." box="[913,1071,291,316]" page="25" pageId="27" pageNumber="428" year="1936">
<bibRefCitation author="Miwa, Y." box="[913,1071,291,316]" pageId="27" pageNumber="428" pagination="1 - 98" refId="ref26431" refString="Miwa, Y. (1936) Family Cicindelidae. In: Uchida, T., Miwa, Y. &amp; Chujo M. (Eds.), Fauna Nipponica. Sanseido Co., Ltd. Co., Ltd. Tokyo, 10 (4), pp. 1 - 98. [in Japanese]" type="journal article" year="1936">Miwa 1936: 25</bibRefCitation>
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<treatmentCitation author="Wiesner, J." box="[1082,1265,292,316]" page="40" pageId="27" pageNumber="428" year="1988">
<bibRefCitation author="Wiesner, J." box="[1082,1265,292,316]" pageId="27" pageNumber="428" pagination="5 - 107" refId="ref27536" refString="Wiesner, J. (1988) Die Gattung Therates Latr. und ihre Arten. 15. Beitrag zur Kenntnis der Cicindelidae (Coleoptera). Mitteilungen der Munchner Entomologischen Gesellschaft, 78, 5 - 107." type="journal article" year="1988">Wiesner 1988: 40</bibRefCitation>
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<bibRefCitation author="Wiesner, J." box="[1275,1368,292,316]" pageId="27" pageNumber="428" refId="ref27575" refString="Wiesner, J. (1992) Verzeichnis der Sandlaufkafer der Welt, Checklist of the Tiger Beetles of the World. Vol. 27. Beitrag zur Kenntnis der Cicindelidae. Verlag Erna Bauer, Keltern, 364 pp." type="book" year="1992">1992: 92</bibRefCitation>
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<treatmentCitation author="Wiesner, J." page="95" pageId="27" pageNumber="428" year="2020">
<bibRefCitation author="Wiesner, J." pageId="27" pageNumber="428" refId="ref27678" refString="Wiesner, J. (2020) Checklist of the tiger beetles of the world. 2 nd Edition. Edition winterwork, Borsdorf, 540 pp." type="book" year="2020">2020: 95</bibRefCitation>
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;
<treatmentCitation author="Cassola, F." box="[239,431,322,346]" page="510" pageId="27" pageNumber="428" year="1985">
<bibRefCitation author="Cassola, F." box="[239,431,322,346]" pageId="27" pageNumber="428" pagination="499 - 514" refId="ref25207" refString="Cassola, F. (1985) &quot; Memoire sur l'Eurychile &quot; di Franco Andrea Bonelli e attuali conoscere sul genre Therates Latreille (Coleoptera, Cicindelidae). Bollettino del Museo Regionale di Scienze Naturali, Torino, 3 (2), 499 - 514." type="journal article" year="1985">Cassola 1985: 510</bibRefCitation>
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;
<treatmentCitation author="Lorenz, W." box="[442,613,322,346]" page="38" pageId="27" pageNumber="428" year="1998">
<bibRefCitation author="Lorenz, W." box="[442,613,322,346]" pageId="27" pageNumber="428" refId="ref26095" refString="Lorenz, W. (1998) Systematic list of extant ground beetles of the word (Insects Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). First Edition. Published by the author, Tutzing, Germany, 503 pp." type="book" year="1998">Lorenz 1998: 38</bibRefCitation>
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<treatmentCitation author="Lorenz, W." box="[625,722,322,346]" page="117" pageId="27" pageNumber="428" year="2005">
<bibRefCitation author="Lorenz, W." box="[625,722,322,346]" pageId="27" pageNumber="428" refId="ref26145" refString="Lorenz, W. (2005) Systematic list of extant ground beetles of the Word (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). 2 nd Edition. Published by the author, Tutzing, 530 pp." type="book" year="2005">2005:117</bibRefCitation>
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<treatmentCitation author="Hua, L. - Z." box="[734,863,322,346]" page="5" pageId="27" pageNumber="428" year="2002">
<bibRefCitation author="Hua, L. - Z." box="[734,863,322,346]" pageId="27" pageNumber="428" refId="ref25990" refString="Hua, L. - Z. (2002) List of Chinese Insects. Volume II. Zhongshau (Sun Yat - sen) University Press - Huayu Nature Book Trade Co. Ltd, Guangzhou China, 612 pp." type="book" year="2002">Hua 2002: 5</bibRefCitation>
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<treatmentCitation author="Werner, K. &amp; Chen, K. - M. &amp; Yang, M. - M." box="[874,1107,322,346]" page="41" pageId="27" pageNumber="428" year="2002">
<bibRefCitation author="Werner, K. &amp; Chen, K. - M. &amp; Yang, M. - M." box="[874,1107,322,346]" pageId="27" pageNumber="428" pagination="35 - 52" refId="ref27480" refString="Werner, K., Chen, K. - M. &amp; Yang, M. - M. (2002) Contribution to the knowledge of the tiger beetles of Taiwan with notes to the species of Lanyu (Coleoptera: Cicindelidae). Collection and Research, 15, 35 - 52." type="journal article" year="2002">
Werner
<emphasis box="[955,1007,322,346]" italics="true" pageId="27" pageNumber="428">et al.</emphasis>
2002: 41
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;
<treatmentCitation author="Puchkov, A. V. &amp; Matalin, A. V." box="[1119,1432,322,346]" page="115" pageId="27" pageNumber="428" year="2003">
<bibRefCitation author="Puchkov, A. V. &amp; Matalin, A. V." box="[1119,1432,322,346]" pageId="27" pageNumber="428" pagination="1802" refId="ref26492" refString="Puchkov, A. V. &amp; Matalin, A. V. (2003) Subfamily Cicindelinae Latreille, 1802. In: Lobl, I &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 1. Archostemata - Myxophaga - Adephaga. Appolo Book, Stenstrup, pp. 99 - 118." type="book chapter" year="2003">Puchkov &amp; Matalin 2003: 115</bibRefCitation>
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<treatmentCitation author="Shook, G &amp; Wiesner, J." box="[199,478,353,377]" page="20" pageId="27" pageNumber="428" year="2006">
<bibRefCitation author="Shook, G &amp; Wiesner, J." box="[199,478,353,377]" pageId="27" pageNumber="428" refId="ref26913" refString="Shook, G &amp; Wiesner, J. (2006) A list of the tiger beetles of China (Coleoptera: Cicindelidae). In: Zhang, Z. - Q. (Ed.), Fauna of China, Vol. 5. Magnolia Press, Auckland, pp. 5 - 26." type="book" year="2006">Shook &amp; Wiesner 2006: 20</bibRefCitation>
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<treatmentCitation author="Wu, X. - Q. &amp; Shook, G." box="[488,719,353,377]" page="90" pageId="27" pageNumber="428" year="2010">
<bibRefCitation author="Wu, X. - Q. &amp; Shook, G." box="[488,719,353,377]" pageId="27" pageNumber="428" pagination="73 - 94" refId="ref27929" refString="Wu, X. - Q. &amp; Shook, G. (2010) Common English and Chinese Names for Tiger Beetles of China. Journal of the Entomological Research Society, 12 (1), 73 - 94." type="journal article" year="2010">Wu &amp; Shook 2010: 90</bibRefCitation>
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<treatmentCitation author="Wu, X. - Q." box="[729,862,353,377]" page="31" pageId="27" pageNumber="428" year="2011">
<bibRefCitation author="Wu, X. - Q." box="[729,862,353,377]" pageId="27" pageNumber="428" pagination="21 - 43" refId="ref27836" refString="Wu, X. - Q. (2011) A revised list of the tiger beetles of China (Coleoptera: Cicindelidae). Cicindela, 43 (2), 21 - 43." type="journal article" year="2011">Wu 2011: 31</bibRefCitation>
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; Putchkov &amp; Matalin 2017: 245.
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</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="428" type="materials_examined">
<paragraph blockId="27.[151,1437,423,2033]" pageId="27" pageNumber="428">
<emphasis bold="true" box="[151,329,423,449]" pageId="27" pageNumber="428">Type material.</emphasis>
<typeStatus box="[340,466,423,449]" pageId="27" pageNumber="428">LECTOTYPE</typeStatus>
, ♂ (designated here)—“Taihorin,
<collectingCountry box="[868,969,424,450]" name="Taiwan" pageId="27" pageNumber="428">Formosa</collectingCountry>
, H. Sauter
<collectionCode box="[1104,1138,423,449]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="428">VI</collectionCode>
.11“ [printed white label]; “Type!, coll. W Horn” [printed white label with black border]; “
<typeStatus box="[888,997,459,485]" pageId="27" pageNumber="428">Syntypus</typeStatus>
” [typewritten red label]; “
<emphasis box="[1300,1436,459,485]" italics="true" pageId="27" pageNumber="428">
<taxonomicName box="[1300,1432,459,485]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="fruhstorferi">fruhstorferi</taxonomicName>
,
</emphasis>
<taxonomicName authority="W. Horn" authorityName="W. Horn" authorityYear="1912" box="[151,341,496,522]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis box="[151,229,496,521]" italics="true" pageId="27" pageNumber="428">sauteri</emphasis>
W. Horn
</taxonomicName>
, det. Wiesner 86” [typewritten white label]; “Coll. W. Horn,
<collectionCode box="[1063,1110,496,522]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Eberswalde” [printed white label]; “
<collectionCode box="[241,290,531,557]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04835” [printed light green label], “
<typeStatus box="[936,1101,531,557]" pageId="27" pageNumber="428">LECTOTYPE</typeStatus>
,
<taxonomicName authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="sauteri">
<emphasis box="[1114,1436,531,557]" italics="true" pageId="27" pageNumber="428">Therates fruhstorferi sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[191,324,567,594]" pageId="27" pageNumber="428" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
</taxonomicName>
, design. A. Matalin et J.
<bibRefCitation author="Wiesner, J. &amp; Geiser, M." box="[612,777,567,593]" pageId="27" pageNumber="428" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner, 2016</bibRefCitation>
” [printed red label] (
<collectionCode box="[1020,1088,567,593]" pageId="27" pageNumber="428">SDEI</collectionCode>
).
<typeStatus box="[1109,1298,567,593]" pageId="27" pageNumber="428">PARALECTOTYPES</typeStatus>
(designated here)—1♂ (
<collectionCode box="[285,333,603,629]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04833)
<specimenCount box="[646,680,603,629]" pageId="27" pageNumber="428" type="female">1♀</specimenCount>
(
<collectionCode box="[694,742,603,629]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04829)—labelled as
<typeStatus box="[1200,1308,603,629]" pageId="27" pageNumber="428">Holotype</typeStatus>
; 3♂♂ (
<collectionCode box="[1388,1436,603,629]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04822, 04825, 04827)
<specimenCount box="[622,678,639,665]" pageId="27" pageNumber="428" type="female">4♀♀</specimenCount>
(
<collectionCode box="[692,740,639,665]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04823, 04824, 04826, 04828)—“Taihorin,
<collectingCountry box="[151,252,676,702]" name="Taiwan" pageId="27" pageNumber="428">Formosa</collectingCountry>
, H. Sauter
<collectionCode box="[377,411,675,701]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="428">VI</collectionCode>
.11“ [printed white label]; “
<typeStatus box="[716,825,675,701]" pageId="27" pageNumber="428">Syntypus</typeStatus>
” [printed red label]; “Coll.
<collectionCode box="[1136,1185,676,702]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
, Eberswalde” [printed white label], “
<typeStatus box="[309,547,711,737]" pageId="27" pageNumber="428">PARALECTOTYPE</typeStatus>
,
<taxonomicName authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" box="[559,1057,711,738]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="sauteri">
<emphasis box="[559,880,711,737]" italics="true" pageId="27" pageNumber="428">Therates fruhstorferi sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[925,1057,711,738]" pageId="27" pageNumber="428" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
</taxonomicName>
, design. A. Matalin et J.
<bibRefCitation author="Wiesner, J. &amp; Geiser, M." pageId="27" pageNumber="428" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner, 2016</bibRefCitation>
” [printed red label]; 3♂♂ (
<collectionCode box="[525,573,747,773]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04830, 04831, 04832)—“Taihorin,
<collectingCountry box="[1214,1315,748,774]" name="Taiwan" pageId="27" pageNumber="428">Formosa</collectingCountry>
, H. Sauter
<collectionCode box="[151,185,783,809]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="428">VI</collectionCode>
.11“ [printed white label]; “CoType!, coll. W Horn” [handwritten and printed white label with black border]; “Coll. W. Horn,
<collectionCode box="[339,386,820,846]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Eberswalde” [printed white label]; “
<typeStatus box="[804,913,819,845]" pageId="27" pageNumber="428">Syntypus</typeStatus>
” [typewritten red label]; “
<emphasis box="[1214,1350,819,845]" italics="true" pageId="27" pageNumber="428">
<taxonomicName box="[1214,1346,819,845]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="fruhstorferi">fruhstorferi</taxonomicName>
,
</emphasis>
<taxonomicName authority="W. Horn" authorityName="W. Horn" authorityYear="1912" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis box="[1358,1436,820,845]" italics="true" pageId="27" pageNumber="428">sauteri</emphasis>
W. Horn
</taxonomicName>
, det. Wiesner 86” [typewritten white label]; “
<collectionCode box="[765,814,855,881]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04830” [printed light green label], “
<typeStatus box="[161,399,891,917]" pageId="27" pageNumber="428">PARALECTOTYPE</typeStatus>
,
<taxonomicName authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" box="[409,898,891,918]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="sauteri">
<emphasis box="[409,726,891,917]" italics="true" pageId="27" pageNumber="428">Therates fruhstorferi sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[768,898,891,918]" pageId="27" pageNumber="428" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
</taxonomicName>
, design. A. Matalin et J.
<bibRefCitation author="Wiesner, J. &amp; Geiser, M." box="[1169,1331,891,917]" pageId="27" pageNumber="428" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner, 2016</bibRefCitation>
” [printed red label];
<specimenCount box="[274,297,927,953]" pageId="27" pageNumber="428" type="female">1♀</specimenCount>
—“Taihorin,
<collectingCountry box="[459,560,928,954]" name="Taiwan" pageId="27" pageNumber="428">Formosa</collectingCountry>
, H. Sauter
<collectionCode box="[686,720,927,953]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="428">VI</collectionCode>
.11“ [printed white label]; “Type!, coll. W Horn” [printed white label with black border]; “Coll. W. Horn,
<collectionCode box="[607,654,964,990]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Eberswalde” [printed white label]; “
<typeStatus box="[1057,1166,963,989]" pageId="27" pageNumber="428">Syntypus</typeStatus>
” [typewritten red label]; “
<emphasis box="[164,300,999,1025]" italics="true" pageId="27" pageNumber="428">
<taxonomicName box="[164,296,999,1025]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="fruhstorferi">fruhstorferi</taxonomicName>
,
</emphasis>
<taxonomicName authority="W. Horn" authorityName="W. Horn" authorityYear="1912" box="[307,491,1000,1026]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis box="[307,385,1000,1025]" italics="true" pageId="27" pageNumber="428">sauteri</emphasis>
W. Horn
</taxonomicName>
, det. Wiesner 86” [typewritten white label]; “
<emphasis box="[1005,1025,1000,1025]" italics="true" pageId="27" pageNumber="428">F.</emphasis>
ssp.,
<emphasis box="[1090,1171,999,1025]" italics="true" pageId="27" pageNumber="428">Sauteri</emphasis>
, m.” [handwritten light yellow label with black border]; “
<collectionCode box="[523,572,1035,1061]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="27" pageNumber="428" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg, Col – 04834” [printed light green label], “
<typeStatus box="[1194,1432,1035,1061]" pageId="27" pageNumber="428">PARALECTOTYPE</typeStatus>
,
<taxonomicName authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" box="[151,645,1071,1098]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="sauteri">
<emphasis box="[151,470,1071,1097]" italics="true" pageId="27" pageNumber="428">Therates fruhstorferi sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[514,645,1071,1098]" pageId="27" pageNumber="428" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
</taxonomicName>
, design. A. Matalin et J.
<bibRefCitation author="Wiesner, J. &amp; Geiser, M." box="[921,1085,1071,1097]" pageId="27" pageNumber="428" pagination="61 - 117" refId="ref27796" refString="Wiesner, J. &amp; Geiser, M. (2016) Faunistic survey of the Tiger Beetles (Coleoptera, Carabidae, Cicindelinae) of Laos. Entomologica Basiliensia et Collectionis Frey, 35, 61 - 117." type="journal article" year="2016">Wiesner, 2016</bibRefCitation>
” [printed red label] (
<collectionCode box="[1319,1387,1071,1097]" pageId="27" pageNumber="428">SDEI</collectionCode>
).
</paragraph>
<paragraph blockId="27.[151,1437,423,2033]" pageId="27" pageNumber="428">
<emphasis bold="true" box="[199,441,1107,1133]" pageId="27" pageNumber="428">Additional material.</emphasis>
<materialsCitation collectingDate="1997-06-13" collectingDateMax="2004-05-29" collectingDateMin="1997-06-13" collectionCode="JW" collectorName="L. M. Yaung" country="China" county="Ilan Co." location="Taiwan" municipality="Fushan" pageId="27" pageNumber="428" specimenCount="1" specimenCount-female="1">
<collectingCountry box="[446,522,1107,1133]" name="China" pageId="27" pageNumber="428">CHINA</collectingCountry>
,
<emphasis box="[532,613,1108,1133]" italics="true" pageId="27" pageNumber="428">
<location LSID="urn:lsid:plazi:treatment:050B87D78D5B2470E1B819E1FA5C75E3:887D601A8D5B2475E33B1D22FDA5726C" box="[532,613,1108,1133]" country="China" county="Ilan Co." municipality="Fushan" name="Taiwan" pageId="27" pageNumber="428">Taiwan</location>
</emphasis>
: 1♂—
<location LSID="urn:lsid:plazi:treatment:050B87D78D5B2470E1B819E1FA5C75E3:887D601A8D5B2475E3801D23FCE7726C" box="[687,807,1107,1136]" country="China" county="Ilan Co." municipality="Fushan" name="Ilan Hsien" pageId="27" pageNumber="428">Ilan Hsien</location>
,
<location LSID="urn:lsid:plazi:treatment:050B87D78D5B2470E1B819E1FA5C75E3:887D601A8D5B2475E21F1D25FC35726C" box="[816,1013,1107,1133]" country="China" county="Ilan Co." municipality="Fushan" name="Yuanshan Hsiang" pageId="27" pageNumber="428">Yuanshan Hsiang</location>
,
<location LSID="urn:lsid:plazi:treatment:050B87D78D5B2470E1B819E1FA5C75E3:887D601A8D5B2475E2D01D25FADB726C" box="[1023,1307,1107,1133]" country="China" county="Ilan Co." municipality="Fushan" name="Fushan Botanical Garden" pageId="27" pageNumber="428">Fushan Botanical Garden</location>
,
<date box="[1317,1432,1107,1134]" pageId="27" pageNumber="428" value="2004-05-29">
<collectingDate box="[1317,1432,1107,1134]" pageId="27" pageNumber="428" value="2004-05-29">29.5.2004</collectingDate>
</date>
, leg.
<collectorName box="[203,391,1143,1170]" pageId="27" pageNumber="428">Tzong-Yeng Lin</collectorName>
(
<collectionCode box="[410,453,1143,1169]" pageId="27" pageNumber="428">JW</collectionCode>
); 1♂—
<collectingMunicipality box="[536,623,1145,1172]" pageId="27" pageNumber="428">Fushan</collectingMunicipality>
,
<collectingCounty box="[638,734,1143,1169]" pageId="27" pageNumber="428">Ilan Co.</collectingCounty>
,
<collectorName box="[749,834,1143,1169]" pageId="27" pageNumber="428">Taiwan</collectorName>
,
<date box="[848,983,1143,1169]" pageId="27" pageNumber="428" value="1997-06-13">
<collectingDate box="[848,983,1143,1169]" pageId="27" pageNumber="428" value="1997-06-13">VI-13-1997</collectingDate>
</date>
,
<collectorName box="[998,1136,1144,1170]" pageId="27" pageNumber="428">L.M. Yaung</collectorName>
(
<collectionCode box="[1154,1197,1143,1169]" pageId="27" pageNumber="428">JW</collectionCode>
);
</materialsCitation>
<materialsCitation collectingDate="1997-06-13" collectingDateMax="2004-05-29" collectingDateMin="1997-06-13" collectionCode="JW" collectorName="L. M. Yaung" country="China" county="Ilan Co." location="Taiwan" municipality="Fushan" pageId="27" pageNumber="428" specimenCount="1" specimenCount-female="1">
<specimenCount box="[1221,1243,1143,1169]" pageId="27" pageNumber="428" type="female">1♀</specimenCount>
—
<collectorName box="[1274,1432,1143,1169]" pageId="27" pageNumber="428">Taipei Hsien</collectorName>
,
<collectorName box="[151,238,1179,1205]" pageId="27" pageNumber="428">Ssuling</collectorName>
,
<collectorName box="[249,348,1180,1206]" pageId="27" pageNumber="428">Taoyuan</collectorName>
,
<date box="[359,410,1179,1206]" pageId="27" pageNumber="428" value="1987-06-05">
5.
<collectionCode box="[376,410,1179,1206]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="27" pageNumber="428">VI</collectionCode>
</date>
</materialsCitation>
<materialsCitation box="[409,664,1179,1206]" collectionCode="JW" collectorName="K. La" country="China" pageId="27" pageNumber="428" specimenCount="1">
.1987, leg.
<collectorName box="[534,599,1180,1206]" pageId="27" pageNumber="428">K. La</collectorName>
(
<collectionCode box="[614,657,1179,1205]" pageId="27" pageNumber="428">JW</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="428" type="reference_group">
<paragraph blockId="27.[151,1437,423,2033]" pageId="27" pageNumber="428">
<emphasis bold="true" box="[199,336,1215,1241]" pageId="27" pageNumber="428">References.</emphasis>
<collectingCountry box="[347,423,1215,1241]" name="China" pageId="27" pageNumber="428">CHINA</collectingCountry>
,
<emphasis box="[437,528,1216,1241]" italics="true" pageId="27" pageNumber="428">Taiwan:</emphasis>
1♂—near Kuangfu, ca.
<quantity box="[818,895,1215,1242]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="27" pageNumber="428" unit="m" value="250.0">250 m</quantity>
,
<date box="[910,1027,1215,1241]" pageId="27" pageNumber="428" value="2001-05-14">14.V.2001</date>
, leg. U. Buchsbaum (
<bibRefCitation author="Werner, K. &amp; Chen, K. - M. &amp; Yang, M. - M." pageId="27" pageNumber="428" pagination="35 - 52" refId="ref27480" refString="Werner, K., Chen, K. - M. &amp; Yang, M. - M. (2002) Contribution to the knowledge of the tiger beetles of Taiwan with notes to the species of Lanyu (Coleoptera: Cicindelidae). Collection and Research, 15, 35 - 52." type="journal article" year="2002">
Werner
<emphasis box="[1377,1437,1215,1241]" italics="true" pageId="27" pageNumber="428">et al.</emphasis>
2002
</bibRefCitation>
); Wulai (
<bibRefCitation author="Wiesner, J." box="[320,484,1251,1277]" pageId="27" pageNumber="428" pagination="5 - 107" refId="ref27536" refString="Wiesner, J. (1988) Die Gattung Therates Latr. und ihre Arten. 15. Beitrag zur Kenntnis der Cicindelidae (Coleoptera). Mitteilungen der Munchner Entomologischen Gesellschaft, 78, 5 - 107." type="journal article" year="1988">Wiesner 1988</bibRefCitation>
); Hualien Co, Jungyang Mountains, Butterfly Valley, ca.
<quantity box="[1154,1231,1251,1278]" metricMagnitude="2" metricUnit="m" metricValue="3.5" pageId="27" pageNumber="428" unit="m" value="350.0">350 m</quantity>
, S Kuangfu, near Fuiuan, ca.
<geoCoordinate box="[282,375,1287,1314]" degrees="23" direction="north" minutes="35" orientation="latitude" pageId="27" pageNumber="428" precision="925" value="23.583334">
23
<superScript attach="none" box="[310,318,1287,1301]" fontSize="6" pageId="27" pageNumber="428">o</superScript>
35’N
</geoCoordinate>
<geoCoordinate box="[383,489,1287,1313]" degrees="121" direction="east" minutes="21" orientation="longitude" pageId="27" pageNumber="428" precision="925" value="121.35">
121
<superScript attach="none" box="[425,433,1287,1301]" fontSize="6" pageId="27" pageNumber="428">o</superScript>
21’E
</geoCoordinate>
,
<date box="[501,660,1287,1314]" pageId="27" pageNumber="428" value="2001-05-13" valueMax="2001-05-15" valueMin="2001-05-13">13–15.V.2001</date>
, leg. K. Werner (K. Werner, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection pageId="27" pageNumber="428" type="diagnosis">
<paragraph blockId="27.[151,1437,423,2033]" pageId="27" pageNumber="428">
<emphasis bold="true" box="[199,322,1323,1349]" pageId="27" pageNumber="428">Diagnosis.</emphasis>
From related
<taxonomicName authorityName="W. Horn" authorityYear="1913" box="[483,588,1323,1349]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis box="[483,588,1323,1349]" italics="true" pageId="27" pageNumber="428">T. vitalisi</emphasis>
</taxonomicName>
,
<taxonomicName authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[603,746,1323,1349]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis box="[603,746,1323,1349]" italics="true" pageId="27" pageNumber="428">T. biserratus</emphasis>
</taxonomicName>
and
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[802,957,1323,1349]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="428" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. n.">
<emphasis box="[802,957,1323,1349]" italics="true" pageId="27" pageNumber="428">T. hunanensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[965,1030,1323,1349]" pageId="27" pageNumber="428">
<taxonomicNameLabel box="[965,1030,1323,1349]" pageId="27" pageNumber="428" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
this species is clearly distinguished by the shape of antennomeres 9–11 (
<figureCitation box="[557,682,1359,1385]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="27" pageNumber="428">Figs 48, 49</figureCitation>
<emphasis box="[688,719,1360,1385]" italics="true" pageId="27" pageNumber="428">vs.</emphasis>
<figureCitation box="[725,854,1359,1385]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="27" pageNumber="428">Figs 42–47</figureCitation>
), by colouration of hind femora (
<figureCitation box="[1220,1345,1359,1385]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="27" pageNumber="428">Figs 26, 28</figureCitation>
<emphasis box="[1351,1382,1360,1385]" italics="true" pageId="27" pageNumber="428">vs.</emphasis>
<figureCitation captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="27" pageNumber="428">Figs 15, 17–19, 21, 22, 24, 25</figureCitation>
), by elytral pattern with only well-developed central dot (
<figureCitation box="[1099,1230,1395,1421]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="27" pageNumber="428">Figs 26, 28</figureCitation>
,
<figureCitation box="[1244,1341,1395,1421]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="27" pageNumber="428">109–114</figureCitation>
<emphasis box="[1349,1380,1396,1421]" italics="true" pageId="27" pageNumber="428">vs.</emphasis>
<figureCitation captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="27" pageNumber="428">Figs 15, 17–19, 21, 22, 24, 25</figureCitation>
,
<figureCitation box="[452,539,1431,1457]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="27" pageNumber="428">99–108</figureCitation>
), as well as by the longer aedeagus with long thin apex (
<figureCitation box="[1185,1341,1431,1457]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="27" pageNumber="428">Figs 173, 174</figureCitation>
<emphasis box="[1350,1380,1432,1457]" italics="true" pageId="27" pageNumber="428">vs.</emphasis>
<figureCitation captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="27" pageNumber="428">Figs 152–171</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection lastPageId="28" lastPageNumber="429" pageId="27" pageNumber="428" type="description">
<paragraph blockId="27.[151,1437,423,2033]" pageId="27" pageNumber="428">
<emphasis bold="true" box="[199,373,1503,1529]" pageId="27" pageNumber="428">Redescription.</emphasis>
TL =
<quantity box="[444,608,1503,1530]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.29" metricValueMin="1.0699999999999998" pageId="27" pageNumber="428" unit="mm" value="11.8" valueMax="12.9" valueMin="10.7">10.7–12.9 mm</quantity>
(mean =
<quantity box="[716,832,1503,1530]" metricMagnitude="-2" metricUnit="m" metricValue="1.155" pageId="27" pageNumber="428" unit="mm" value="11.55">11.55 mm</quantity>
, n = 10) in males,
<quantity box="[1044,1207,1503,1530]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" metricValueMax="1.29" metricValueMin="1.15" pageId="27" pageNumber="428" unit="mm" value="12.2" valueMax="12.9" valueMin="11.5">11.5–12.9 mm</quantity>
(mean =
<quantity box="[1315,1432,1503,1530]" metricMagnitude="-2" metricUnit="m" metricValue="1.216" pageId="27" pageNumber="428" unit="mm" value="12.16">12.16 mm</quantity>
, n = 7) in females.
</paragraph>
<paragraph blockId="27.[151,1437,423,2033]" pageId="27" pageNumber="428">
Head black with light blue-violet reflection; orbital plates indistinctly striated in posterior third; frons slightly elevated in anterior third (see above), slightly downward anteriorly (see at side), frontal sulci deep and slightly convergent in anterior half but shallow and widely divergent in basal half; occiput flat. Mandibles pale-yellow or yellow-reddish with brown teeth and apical molar. Labial palpi pale except brown apical palpomere; two basal palpomeres of maxillary palpi pale, palpomeres 3 and 4 light brown or brown. Antennae barely extend shoulders; scape pale on anterior side and brown-black on posterior side; antennomere 2 brown or dark-brown with lighter apex; antennomeres 3–6 dark-brown with light-brown apical third or brown-black with brown apices; antennomeres 9 and
<quantity box="[151,207,1827,1853]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="27" pageNumber="428" unit="in" value="10.0">10 in</quantity>
males with protruding downward anterior lower margins, in females barely dilated anteriorly; antennomere
<quantity metricMagnitude="-1" metricUnit="m" metricValue="2.794" pageId="27" pageNumber="428" unit="in" value="11.0">11 in</quantity>
males slightly dilated with abrupt rough anterior margin (
<figureCitation box="[823,907,1863,1889]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="27" pageNumber="428">Fig. 49</figureCitation>
), in females elongated with wide rounded apex (
<figureCitation box="[159,243,1899,1925]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="27" pageNumber="428">Fig. 48</figureCitation>
). Labrum as long as wide, LW/LL = 0.93–1.08 (mean = 1.01, n = 17), black-brown with large apical yellow spot unseparated from central apical teeth; apical and lateral teeth red-brown (
<figureCitation box="[1024,1156,1935,1961]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="27" pageNumber="428">Figs 77–80</figureCitation>
).
</paragraph>
<paragraph blockId="27.[151,1437,423,2033]" pageId="27" pageNumber="428">Pronotum black with bright blue-violet reflection; indistinctly transverse, PW/PL = 1.0–1.1 (mean = 1.04, n = 17), apical lobe practically equal wide throughout; thorax black with violet reflection.</paragraph>
<paragraph blockId="28.[151,1437,151,573]" pageId="28" pageNumber="429">Fore femora pale with dark brown strip on posterior side of knees; middle and hind femora pale with dark brown knees; fore and middle tibiae dark brown with brown basal third or light brown with yellow-brown basal half, hind tibiae black or brown-black with violet tinge; tarsi black with violet tinge, in some specimens tarsomeres 1 and 2 brown-black with black apices; HTbL/HTaL = 1.10–1.20 (mean = 1.15, n = 13).</paragraph>
<paragraph blockId="28.[151,1437,151,573]" pageId="28" pageNumber="429">
Elytra black with bright blue-violet reflection, sub-parallel or indistinctly divergent toward apex, EL/EW = 1.86–2.08 (mean = 1.95, n = 17); punctuation deep and regular between humeral hump and central dot, shallow and sparse in apical third; scutellum black with violet tinge; apical margin in males shortly cut, in females angularly rounded. Elytral pattern presented by short slightly oblique pale central dot as well as by very small and indistinct brownish basal portion of humeral lunule (see in front) in some specimens (
<figureCitation box="[999,1128,439,465]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="28" pageNumber="429">Figs 26, 28</figureCitation>
,
<figureCitation box="[1140,1241,439,465]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="28" pageNumber="429">109–114</figureCitation>
).
</paragraph>
<paragraph blockId="28.[151,1437,151,573]" pageId="28" pageNumber="429">
Aedeagus with long straight thin apex as well as with short lateral carinae (
<figureCitation box="[1052,1213,475,501]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="28" pageNumber="429">Figs 173, 174</figureCitation>
); AL =
<quantity box="[1301,1437,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="28" pageNumber="429" unit="mm" value="3.05" valueMax="3.2" valueMin="2.9">2.9–3.2 mm</quantity>
(mean = 3.03, n = 7), EL/AL =
<quantity box="[503,666,510,537]" metricMagnitude="-3" metricUnit="m" metricValue="2.41" metricValueMax="2.55" metricValueMin="2.27" pageId="28" pageNumber="429" unit="mm" value="2.41" valueMax="2.55" valueMin="2.27">2.27–2.55 mm</quantity>
(mean = 2.41, n = 7).
</paragraph>
</subSubSection>
<subSubSection box="[199,655,546,573]" pageId="28" pageNumber="429" type="distribution">
<paragraph blockId="28.[151,1437,151,573]" box="[199,655,546,573]" pageId="28" pageNumber="429">
<emphasis bold="true" box="[199,353,546,572]" pageId="28" pageNumber="429">Distribution.</emphasis>
<collectingCountry box="[359,437,547,573]" name="China" pageId="28" pageNumber="429">CHINA</collectingCountry>
: Taiwan (
<figureCitation box="[544,642,547,573]" captionStart="FIGURES 184" captionStartId="28.[152,270,1579,1603]" captionTargetBox="[151,1435,597,1556]" captionTargetId="figure-226@28.[151,1435,597,1556]" captionTargetPageId="28" captionText="FIGURES 184. Distribution of Therates spp.: dark blue solid circle—T. vitalisi vitalisi; dark blue half-solid circle—T. vitalisi ida; lilac solid circle—T. hunnanensis sp. n.; orange solid circle—T. biserratus; pink solid circle—T. motoensis; red open circle—T. sauteri; red solid quadrat—T. fruhstorferi fruhstorferi; red half-solid quadrat—T. fruhstorferi australis ssp. n.; blue solid quadrat—T. ziyardamensis sp. n.; green solid quadrat—T. pseudovitalisi sp. n. (nominotypical subspecies); green halfsolid quadrat—T. pseudovitalisi tibetana ssp. n.; L in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png." figureDoi="http://doi.org/10.5281/zenodo.7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="28" pageNumber="429">Fig. 184</figureCitation>
).
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7758916" ID-Zenodo-Dep="7758916" httpUri="https://zenodo.org/record/7758916/files/figure.png" pageId="28" pageNumber="429" startId="28.[152,270,1579,1603]" targetBox="[151,1435,597,1556]" targetPageId="28">
<paragraph blockId="28.[151,1437,1579,1783]" pageId="28" pageNumber="429">
<emphasis bold="true" box="[152,321,1579,1603]" pageId="28" pageNumber="429">FIGURES 184.</emphasis>
Distribution of
<taxonomicName box="[484,624,1579,1603]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis box="[484,573,1579,1603]" italics="true" pageId="28" pageNumber="429">Therates</emphasis>
spp.
</taxonomicName>
: dark blue solid circle—
<taxonomicName authorityName="W. Horn" authorityYear="1913" box="[873,1042,1579,1603]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="vitalisi">
<emphasis box="[873,1042,1579,1603]" italics="true" pageId="28" pageNumber="429">T. vitalisi vitalisi</emphasis>
</taxonomicName>
; dark blue half-solid circle—
<taxonomicName authorityName="Mandl" authorityYear="1954" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="ida">
<emphasis italics="true" pageId="28" pageNumber="429">T. vitalisi ida</emphasis>
</taxonomicName>
; lilac solid circle—
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[394,550,1615,1639]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="hunnanensis" status="sp. n.">
<emphasis box="[394,550,1615,1639]" italics="true" pageId="28" pageNumber="429">T. hunnanensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[559,619,1615,1639]" pageId="28" pageNumber="429">
<taxonomicNameLabel box="[559,619,1615,1639]" pageId="28" pageNumber="429" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; orange solid circle—
<taxonomicName authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[855,987,1615,1639]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis box="[855,987,1615,1639]" italics="true" pageId="28" pageNumber="429">T. biserratus</emphasis>
</taxonomicName>
; pink solid circle—
<taxonomicName authorityName="Tan" authorityYear="1981" box="[1199,1328,1616,1639]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis box="[1199,1328,1616,1639]" italics="true" pageId="28" pageNumber="429">T. motoensis</emphasis>
</taxonomicName>
; red open circle—
<taxonomicName authorityName="W. Horn" authorityYear="1912" box="[233,328,1652,1675]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis box="[233,328,1652,1675]" italics="true" pageId="28" pageNumber="429">T. sauteri</emphasis>
</taxonomicName>
; red solid quadrat—
<taxonomicName authorityName="W. Horn" authorityYear="1902" box="[540,807,1651,1675]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
<emphasis box="[540,807,1651,1675]" italics="true" pageId="28" pageNumber="429">T. fruhstorferi fruhstorferi</emphasis>
</taxonomicName>
; red half-solid quadrat—
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[1065,1304,1651,1675]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="australis">
<emphasis box="[1065,1304,1651,1675]" italics="true" pageId="28" pageNumber="429">T. fruhstorferi australis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1311,1378,1651,1675]" pageId="28" pageNumber="429">ssp. n.</emphasis>
; blue solid quadrat—
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[310,481,1687,1711]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="ziyardamensis" status="sp. n.">
<emphasis box="[310,481,1687,1711]" italics="true" pageId="28" pageNumber="429">T. ziyardamensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[489,548,1687,1711]" pageId="28" pageNumber="429">
<taxonomicNameLabel box="[489,548,1687,1711]" pageId="28" pageNumber="429" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
; green solid quadrat—
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[786,954,1687,1711]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="species" species="pseudovitalisi" status="sp. n.">
<emphasis box="[786,954,1687,1711]" italics="true" pageId="28" pageNumber="429">T. pseudovitalisi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[962,1021,1687,1711]" pageId="28" pageNumber="429">
<taxonomicNameLabel box="[962,1021,1687,1711]" pageId="28" pageNumber="429" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(nominotypical subspecies); green halfsolid quadrat—
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[311,571,1723,1747]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="429" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" subSpecies="tibetana">
<emphasis box="[311,571,1723,1747]" italics="true" pageId="28" pageNumber="429">T. pseudovitalisi tibetana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[580,649,1723,1747]" pageId="28" pageNumber="429">ssp. n.</emphasis>
;
<emphasis box="[665,679,1724,1747]" italics="true" pageId="28" pageNumber="429">L</emphasis>
in the symbols—according only to the literature sources. Map source— https://commons.wikimedia.org/wiki/File:East_Asia_topographic_map.png.
</paragraph>
</caption>
<subSubSection lastPageId="30" lastPageNumber="431" pageId="29" pageNumber="430" type="key">
<paragraph blockId="29.[151,733,151,178]" box="[151,733,151,178]" pageId="29" pageNumber="430">
<emphasis bold="true" box="[151,733,151,178]" pageId="29" pageNumber="430">
Key to species of
<taxonomicName box="[366,621,151,178]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="fruhstorferi">
<emphasis bold="true" box="[366,621,151,178]" italics="true" pageId="29" pageNumber="430">Therates fruhstorferi</emphasis>
</taxonomicName>
complex
</emphasis>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,213,235]" pageId="29" pageNumber="430">1(2) Male............................................................................................... 3</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,243,265]" pageId="29" pageNumber="430">2(1) Female............................................................................................ 25</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,272,294]" pageId="29" pageNumber="430">
3(4) Apex of aedeagus with lateral carinae (
<figureCitation box="[600,734,272,294]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="29" pageNumber="430">Figs 152–174</figureCitation>
)........................................................ 5
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,301,323]" pageId="29" pageNumber="430">
4(3) Apex of aedeagus with lateral carinae (
<figureCitation box="[600,733,301,323]" captionStart-0="FIGURES 115–135" captionStart-1="FIGURES 136–151" captionStartId-0="22.[152,270,1114,1138]" captionStartId-1="23.[152,270,1334,1358]" captionTargetBox-0="[155,1431,193,1084]" captionTargetBox-1="[175,1413,195,1300]" captionTargetId-0="figure-28@22.[151,1435,181,1089]" captionTargetId-1="figure-17@23.[151,1435,181,1309]" captionTargetPageId-0="22" captionTargetPageId-1="23" captionText-0="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." captionText-1="FIGURES 136–151. Therates fruhstorferi fruhstorferi, variability of aedeagus, left lateral view: 136–143—Vietnam, Phu Tho Province, ~90 km W of Hanoi, Xuan Son Nat. Park, h = 400–700 m, 21o07’01’’N 104o56’39’’E, 6–15.VI.2014; 144–151— Vietnam, Vinh Phu Province, Tam Dao, different years (144, 145—3–11.VI.1985; 146—27.V–2.VI.1986; 147—2–11.VI.1989; 148—6–10.V.1990; 149—12.VI.1994; 150, 151—9–19.V.1996). Scales in mm." figureDoi-0="http://doi.org/10.5281/zenodo.7758900" figureDoi-1="http://doi.org/10.5281/zenodo.7758904" httpUri-0="https://zenodo.org/record/7758900/files/figure.png" httpUri-1="https://zenodo.org/record/7758904/files/figure.png" pageId="29" pageNumber="430">Figs 115–151</figureCitation>
)....................................................... 15
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
5(6) Elytra shining green, with very small (see in front of) yellow basal portion of humeral lunule invisible dorsally and small trapezoid yellow central dot (
<figureCitation box="[504,578,360,382]" captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="29" pageNumber="430">Figs 13</figureCitation>
,
<figureCitation box="[588,615,360,381]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="29" pageNumber="430">97</figureCitation>
). Labrum yellowish with black margins and large black basal spot (
<figureCitation box="[1255,1327,360,382]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="29" pageNumber="430">Fig. 62</figureCitation>
). Aedeagus clearly dilated in apical third middle portion (
<figureCitation box="[663,745,389,411]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="29" pageNumber="430">Fig. 172</figureCitation>
).....................................
<taxonomicName authority="Tan, 1981" authorityName="Tan" authorityYear="1981" box="[1215,1436,390,411]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis box="[1215,1335,390,411]" italics="true" pageId="29" pageNumber="430">T. motoensis</emphasis>
<bibRefCitation author="Tan, J. - J." box="[1341,1436,390,411]" pageId="29" pageNumber="430" pagination="331 - 335" refId="ref27058" refString="Tan, J. - J. (1981) Cicindelidae. In: Insects of Xizang (Tibet). Vol. I. Comprehensive Scientific Expedition to the Qinghai-Xizang Plateau, Chinese Academy of Sciences. Science Press, Beijing, pp. 331 - 335. [in Chinese, with English summary]" type="book chapter" year="1981">Tan, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
6(5) Elytra shining violet-black or bluish-black with purple lustre, basal portion of humeral lunule large or medium-sized (
<figureCitation captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="29" pageNumber="430">Figs 17–19, 24–26, 28</figureCitation>
,
<figureCitation box="[405,681,448,470]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="29" pageNumber="430">100, 103, 106, 107, 109–111</figureCitation>
). Labrum black or brown with large or medium-sized pale apical spot (
<figureCitation captionStart="FIGURES 65–72" captionStartId="17.[152,270,1944,1968]" captionTargetBox="[283,1300,190,1909]" captionTargetId="figure-17@17.[267,1320,181,1920]" captionTargetPageId="17" captionText="FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype." figureDoi="http://doi.org/10.5281/zenodo.7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="29" pageNumber="430">Figs 65, 69–71</figureCitation>
,
<figureCitation box="[298,433,477,499]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="29" pageNumber="430">74, 75, 77, 79</figureCitation>
). Aedeagus not dilated in apical third (
<figureCitation box="[803,1033,477,499]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="29" pageNumber="430">Figs 152–171, 173, 174</figureCitation>
)............................... 7
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,507,529]" pageId="29" pageNumber="430">
7(8) Elytra with four pale dots including apical one (
<figureCitation box="[684,795,507,529]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="29" pageNumber="430">Figs 24, 25</figureCitation>
,
<figureCitation box="[805,890,507,529]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="29" pageNumber="430">106, 107</figureCitation>
)........................................... 9
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,536,558]" pageId="29" pageNumber="430">
8(7) Elytra with one–three pale dots, apical dot undeveloped (
<figureCitation box="[773,956,536,558]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="29" pageNumber="430">Figs 17–19, 26, 28</figureCitation>
,
<figureCitation box="[966,1145,536,557]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="29" pageNumber="430">100, 103, 109–111</figureCitation>
)..................... 11
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
9(10) Antennomeres 9 and 10 sharply serrated, antennomere 11 very wide and sharply abrupt anteriorly (
<figureCitation box="[1214,1289,565,587]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="29" pageNumber="430">Fig. 44</figureCitation>
). Labrum with medium-sized apical yellow spot narrowly separated from apical teeth (
<figureCitation box="[905,976,595,617]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="29" pageNumber="430">Fig. 74</figureCitation>
). Elytra wider, EL/EW = 1.87; apical lunule thin and not extend the suture (
<figureCitation box="[482,567,624,646]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="29" pageNumber="430">Fig. 106</figureCitation>
). Hind tibiae as well as middle and hind tarsi entirely black (
<figureCitation box="[1155,1227,624,646]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="29" pageNumber="430">Fig. 24</figureCitation>
). Aedeagus with very small blunt apex (
<figureCitation box="[398,482,653,675]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="29" pageNumber="430">Fig. 168</figureCitation>
); EL/AL = 2.84......................................
<taxonomicName authority="Tan, Mo, Liang, 1991" authorityName="Tan, Mo, Liang" authorityYear="1991" box="[1101,1436,653,675]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis box="[1101,1221,653,675]" italics="true" pageId="29" pageNumber="430">T. biserratus</emphasis>
<bibRefCitation author="Tan, J. - J. &amp; Mo, W. &amp; Liang, C." box="[1227,1436,653,675]" pageId="29" pageNumber="430" pagination="243 - 245" refId="ref27368" refString="Tan, J. - J., Mo, W. &amp; Liang, C. (1991) A new species of genus Therates Latreille from Guizhou province, China (Coleoptera: Cicindelidae). Sinozoologia, 5 (8), 243 - 245. [in Chinese, with English summary]" type="journal article" year="1991">Tan, Mo, Liang, 1991</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
10(9) Antennomeres 9 and 10 with protruding downward anterior lower margins, antennomere 11 oval with slightly elongated apex (
<figureCitation box="[233,305,712,734]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="29" pageNumber="430">Fig. 43</figureCitation>
). Labrum with very large apical yellow spot unseparated from apical teeth (
<figureCitation box="[1036,1108,712,734]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="29" pageNumber="430">Fig. 75</figureCitation>
). Elytra narrower, EL/EW = 2.03; apical lunule wide and extend the suture (
<figureCitation box="[628,713,741,763]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="29" pageNumber="430">Fig. 107</figureCitation>
). Hind tibiae yellow or brown except brown-black apices, middle and hind tarsi black except yellow-brown or brown basal tarsomeres (
<figureCitation box="[803,874,771,793]" captionStart="FIGURES 15–28" captionStartId="9.[152,270,1641,1665]" captionTargetBox="[156,1418,712,1612]" captionTargetId="figure-217@9.[151,1435,709,1617]" captionTargetPageId="9" captionText="FIGURES 15–28. Therates spp., habitus: 15–18—T. vitalisi vitalisi (15, 16—Vietnam, Lao Cai; 17—Vietnam, Fan Si Pan Mnt; 18—Vietnam, Sa Pa); 19–21—T. vitalisi ida (China, Kuatun); 22–24—T. biserratus (22, 23—China, Yinjiang County; 24—China, Fanjing Shan-Kuaichang); 25—T. hunnanensis sp. n. (China, Shunhuangshan); 26–28—T. sauteri (26, 27—China, Tiawan, Taihorin; 28—China, Taiwan, Ilan Hsien); 17–20, 24–28—males; 15, 16, 21–23 —females; 15, 16, 19, 20, 25— holotypes; 22, 23—paratype; 26, 27—lectotype; 15, 17–19, 21, 22, 24–26, 28—habitus; 16, 20, 23, 27—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758876" httpUri="https://zenodo.org/record/7758876/files/figure.png" pageId="29" pageNumber="430">Fig. 25</figureCitation>
). Aedeagus with slightly elongate and slightly curved apex (
<figureCitation box="[233,369,800,822]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="29" pageNumber="430">Figs 169–171</figureCitation>
); EL/AL = 2.74............................................................
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1251,1382,800,822]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="hunanensis" status="sp.n.">
<emphasis box="[1251,1382,800,822]" italics="true" pageId="29" pageNumber="430">T. hunanensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1388,1437,800,821]" pageId="29" pageNumber="430">
<taxonomicNameLabel box="[1388,1437,800,821]" pageId="29" pageNumber="430" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
11(12) Antennomeres 9 and 10 with downward protruding lower anterior margins, antennomere 11 slightly dilated with abrupt rough anterior margin (
<figureCitation box="[389,461,859,881]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="29" pageNumber="430">Fig. 49</figureCitation>
). Labrum with large apical yellow spot unseparated from apical teeth (
<figureCitation box="[1148,1261,859,881]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="29" pageNumber="430">Figs 77, 79</figureCitation>
). Elytra with very small indistinct basal portion of humeral lunule and relatively short and wide central dot, basal dot undeveloped (
<figureCitation box="[1292,1426,888,910]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="29" pageNumber="430">Figs 109–111</figureCitation>
). Hind femora pale with dark brown knees; HTbL/HTaL = 1.15. Aedeagus with long straight thin apex and short lateral carinae (
<figureCitation box="[233,368,947,969]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="29" pageNumber="430">Figs 173, 174</figureCitation>
); EL/AL = 2.41.......................................................
<taxonomicName authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" box="[1199,1436,947,968]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis box="[1199,1289,947,968]" italics="true" pageId="29" pageNumber="430">T. sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,947,968]" pageId="29" pageNumber="430" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
12(11) Antennomeres 9 and 10 sharply serrated, antennomere 11 sharply dilated with abrupt straight anterior margin (
<figureCitation box="[1273,1344,976,998]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="29" pageNumber="430">Fig. 46</figureCitation>
). Labrum with small or medium-sized apical yellow spot clearly separated from apical teeth (
<figureCitation box="[1022,1170,1005,1027]" captionStart="FIGURES 65–72" captionStartId="17.[152,270,1944,1968]" captionTargetBox="[283,1300,190,1909]" captionTargetId="figure-17@17.[267,1320,181,1920]" captionTargetPageId="17" captionText="FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype." figureDoi="http://doi.org/10.5281/zenodo.7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="29" pageNumber="430">Figs 65, 69–71</figureCitation>
). Elytra with clearly-visible basal portion of humeral lunule, small basal dot entirely reduced in some specimens and slightly oblique central dot (
<figureCitation captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="29" pageNumber="430">Figs 100, 103</figureCitation>
). Hind femora black except pale-yellow basal third; HTbL/HTaL = 1.21. Aedeagus with short sharply curved apex and long lateral carinae (
<figureCitation box="[374,511,1093,1115]" captionStart="FIGURES 152–174" captionStartId="24.[152,270,1210,1234]" captionTargetBox="[163,1428,181,1165]" captionTargetId="figure-28@24.[151,1435,181,1186]" captionTargetPageId="24" captionText="FIGURES 152–174. Therates spp., aedeagus: 152–166—T. vitalisi vitalisi (152, 153—Vietnam, Xuan Son N.P.; 154— Vietnam, Sa Pa; 155, 156—Laos, Ban Nape—Kaev Nua pass; 157–163—Laos, 15–20 km NW Louang Namtha; 164, 165— Laos, 8 km NE Ban Nape; 166—Laos, Thong Khan); 167—T. vitalisi ida (China, Kuatun); 168—T. biserratus (China, Fanjing Shan-Kuaichang); 169–171—T. hunnanensis sp. n. (China, Shunhuangshan); 172—T. motoensis (China, Hanmi); 173, 174—T. sauteri (173—China, Taiwan, Taihorin; 174—China, Taiwan, Ilan Hsien); 167, 169—holotypes; 170, 171—paratypes; 173— lectotype; 152—dorsal view; 153–174—left lateral view; ac—apical carinae; sp—spring; fl—flagellum. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758906" httpUri="https://zenodo.org/record/7758906/files/figure.png" pageId="29" pageNumber="430">Figs 152–167</figureCitation>
); EL/AL = 2.58.......................................
<taxonomicName authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[1154,1390,1093,1115]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis box="[1154,1243,1093,1115]" italics="true" pageId="29" pageNumber="430">T. vitalisi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1281,1390,1094,1115]" pageId="29" pageNumber="430" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
(13)
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
13(14) Labrum indistinctly transverse, LW/LL = 1.04 (
<figureCitation box="[682,830,1123,1145]" captionStart="FIGURES 65–72" captionStartId="17.[152,270,1944,1968]" captionTargetBox="[283,1300,190,1909]" captionTargetId="figure-17@17.[267,1320,181,1920]" captionTargetPageId="17" captionText="FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype." figureDoi="http://doi.org/10.5281/zenodo.7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="29" pageNumber="430">Figs 65, 69, 70</figureCitation>
). Fore and middle femora yellow brown............................................................................................
<taxonomicName authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[1129,1436,1152,1174]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="vitalisi">
<emphasis box="[1129,1289,1152,1174]" italics="true" pageId="29" pageNumber="430">T. vitalisi vitalisi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,1152,1174]" pageId="29" pageNumber="430" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1181,1203]" pageId="29" pageNumber="430">
14(13) Labrum clearly transverse, LW/LL = 1.14 (
<figureCitation box="[640,712,1181,1203]" captionStart="FIGURES 65–72" captionStartId="17.[152,270,1944,1968]" captionTargetBox="[283,1300,190,1909]" captionTargetId="figure-17@17.[267,1320,181,1920]" captionTargetPageId="17" captionText="FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype." figureDoi="http://doi.org/10.5281/zenodo.7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="29" pageNumber="430">Fig. 71</figureCitation>
). Fore and middle femora red brown........
<taxonomicName authority="Mandl, 1954" authorityName="Mandl" authorityYear="1954" box="[1182,1436,1181,1203]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="ida">
<emphasis box="[1182,1308,1181,1203]" italics="true" pageId="29" pageNumber="430">T. vitalisi ida</emphasis>
<bibRefCitation author="Mandl, K." box="[1314,1436,1181,1203]" pageId="29" pageNumber="430" pagination="157 - 161" refId="ref26194" refString="Mandl, K. (1954) Zur Kenntnis der Cicindeliden (Col.) Sud-Chinas. Bonner Zoologische Beitrage, 5, 157 - 161." type="journal article" year="1954">Mandl, 1954</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
15(16) Hind tibiae and tarsi partly pale. Apical dot wide and expanded apical margin and nearest apical portion of elytra in
<specimenCount pageId="29" pageNumber="430" type="generic">one specimens</specimenCount>
or only as very narrow yellow or brown apical margin, humeral lunule complete or separated on basal and apical portions, in some specimens apical portion virtually invisible or absent, basal dot presented in
<specimenCount box="[1139,1281,1269,1291]" pageId="29" pageNumber="430" type="generic">one specimens</specimenCount>
or undeveloped in other............................................................
<taxonomicName authority="W. Horn, 1902" authorityName="W. Horn" authorityYear="1902" box="[1038,1436,1299,1320]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
<emphasis box="[1038,1289,1299,1320]" italics="true" pageId="29" pageNumber="430">T. fruhstorferi fruhstorferi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,1299,1320]" pageId="29" pageNumber="430" pagination="65 - 75" refId="ref25658" refString="Horn, W. (1902) Neue Cicindeliden gesammelt von Fruhstorfer in Tonkin 1900. Deutsche Entomologische Zeitschrift, 1902, 65 - 75." type="journal article" year="1902">Horn, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1328,1350]" pageId="29" pageNumber="430">16(15) Hind tibiae and tarsi entirely black...................................................................... 17</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1357,1379]" pageId="29" pageNumber="430">
17(18) Middle tibiae partly pale.......................................................
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[1143,1367,1357,1379]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="australis">
<emphasis box="[1143,1367,1357,1379]" italics="true" pageId="29" pageNumber="430">T. fruhstorferi australis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1373,1436,1358,1379]" pageId="29" pageNumber="430">ssp. n.</emphasis>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1387,1409]" pageId="29" pageNumber="430">18(17) Middle tibiae entirely black or black-brown............................................................... 19</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
19(20) Basal dot absent, central dot relatively short virtually transverse; elytral punctuation deep and regular in basal 3/4 except humeral area, in apical quarter shallow and tenuous (
<figureCitation box="[734,805,1445,1467]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="29" pageNumber="430">Fig. 96</figureCitation>
). Aedeagus with thin clearly sinuate elongate apex (
<figureCitation box="[1305,1387,1445,1467]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="29" pageNumber="430">Fig. 131</figureCitation>
)...................................................................................
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1216,1376,1475,1496]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="ziyardamensis" status="sp. n.">
<emphasis box="[1216,1376,1475,1496]" italics="true" pageId="29" pageNumber="430">T. ziyardamensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1382,1436,1475,1496]" pageId="29" pageNumber="430">
<taxonomicNameLabel box="[1382,1436,1475,1496]" pageId="29" pageNumber="430" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
20(19) Basal dot small, in some specimens virtually invisible or entirely absent, central dot elongate and oblique; elytral punctuation deep and regular between basal humps and central dot then shallow and sparse, apical third virtually smooth (
<figureCitation captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="29" pageNumber="430">Figs 81, 83, 84, 87, 88, 90, 91, 94</figureCitation>
). Aedeagus with relatively wide elongate apex (
<figureCitation box="[856,1085,1563,1585]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="29" pageNumber="430">Figs 115–130, 132–135</figureCitation>
).......................... 21
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
21(22) Basal dot small, in some specimens virtually invisible or entirely absent. Hind tarsi longer, HTbL/HTaL = 1.18. Aedeagus with shorter indistinctly sinuate apex (
<figureCitation box="[545,678,1621,1643]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="29" pageNumber="430">Figs 126–130</figureCitation>
).....................................
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[1143,1367,1621,1643]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="australis">
<emphasis box="[1143,1367,1621,1643]" italics="true" pageId="29" pageNumber="430">T. fruhstorferi australis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1373,1436,1622,1643]" pageId="29" pageNumber="430">ssp. n.</emphasis>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
22(21) Basal dot absent. Hind tarsi shorter, HTbL/HTaL = 1.32. Aedeagus with longer gradually sinuate apex (
<figureCitation box="[1218,1351,1651,1673]" captionStart="FIGURES 115–135" captionStartId="22.[152,270,1114,1138]" captionTargetBox="[155,1431,193,1084]" captionTargetId="figure-28@22.[151,1435,181,1089]" captionTargetPageId="22" captionText="FIGURES 115–135. Therates spp., aedeagus: 115–125—T. fruhstorderi fruhstorferi (115, 116—Vietnam, Vietnam, Montes Mouson; 117–121—Vietnam, Xuan Son N.P.; 122, 123—Vietnam, Tam Dao; 124—Laos, Ban Nape; 125—Laos, Ban Nape— Kaev Nua pass); 126–130—T. fruhstorferi australis ssp. n. (126—Vietnam, Kon Ka Kinh N.P.; 127—Vietnam, Gia Lai; 128— Vietnam, Ngok Boc I Mt.; 129, 130—Vietnam, song Thanh N.P.); 131—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 132– 134—T. pseudovitalisi pseudovitalisi ssp. n. (132—Laos, Ban Song Cha; 133, 134—Laos, 17 km NEE Oudom Xai); 135—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 115—lectotype; 116—paralectotype; 126, 131, 132, 135—holotypes; 127–130, 133, 134—paratypes; 117—dorsal view; 115, 116, 118–135—left lateral view. Scale bars in mm." figureDoi="http://doi.org/10.5281/zenodo.7758900" httpUri="https://zenodo.org/record/7758900/files/figure.png" pageId="29" pageNumber="430">Figs 132–135</figureCitation>
)...................................................................................
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1173,1330,1680,1702]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="pseudovitalisi" status="sp. n.">
<emphasis box="[1173,1330,1680,1702]" italics="true" pageId="29" pageNumber="430">T. pseudovitalisi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1336,1390,1680,1701]" pageId="29" pageNumber="430">
<taxonomicNameLabel box="[1336,1390,1680,1701]" pageId="29" pageNumber="430" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(23)
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
23(24) Antennomere 10 not wider than 11 (
<figureCitation box="[566,637,1709,1731]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="29" pageNumber="430">Fig. 37</figureCitation>
). Elytra black-violet with purple lustre; central dot narrower, distinctly oblique (
<figureCitation captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="29" pageNumber="430">Figs 90–91</figureCitation>
); EL/AL = 2.62...................................................
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1071,1367,1739,1760]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" status="ssp. n." subSpecies="pseudovitalisi">
<emphasis box="[1071,1367,1739,1760]" italics="true" pageId="29" pageNumber="430">T. pseudovitalisi pseudovitalisi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1373,1436,1739,1760]" pageId="29" pageNumber="430">
<taxonomicNameLabel box="[1373,1436,1739,1760]" pageId="29" pageNumber="430" rank="subSpecies">ssp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" pageId="29" pageNumber="430">
24(23) Antennomere 10 wide than 11 (
<figureCitation box="[526,598,1768,1790]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="29" pageNumber="430">Fig. 39</figureCitation>
). Elytra black-blue with light violet reflection; central dot wider, transverse with dilated lateral portion (
<figureCitation box="[374,446,1797,1819]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="29" pageNumber="430">Fig. 94</figureCitation>
); EL/AL = 2.48...........................................
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[1125,1367,1797,1819]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" subSpecies="tibetana">
<emphasis box="[1125,1367,1797,1819]" italics="true" pageId="29" pageNumber="430">T. pseudovitalisi tibetana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1373,1436,1798,1819]" pageId="29" pageNumber="430">ssp. n.</emphasis>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1827,1849]" pageId="29" pageNumber="430">
25(26) Humeral lunule complete or separated on basal and apical portions............
<taxonomicName authority="W. Horn, 1902" authorityName="W. Horn" authorityYear="1902" box="[1038,1436,1827,1848]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
<emphasis box="[1038,1289,1827,1848]" italics="true" pageId="29" pageNumber="430">T. fruhstorferi fruhstorferi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,1827,1848]" pageId="29" pageNumber="430" pagination="65 - 75" refId="ref25658" refString="Horn, W. (1902) Neue Cicindeliden gesammelt von Fruhstorfer in Tonkin 1900. Deutsche Entomologische Zeitschrift, 1902, 65 - 75." type="journal article" year="1902">Horn, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1856,1878]" pageId="29" pageNumber="430">26(25) Humeral lunule without apical portion................................................................... 27</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1885,1907]" pageId="29" pageNumber="430">27(28) Elytra with pale apical dot............................................................................. 29</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1915,1937]" pageId="29" pageNumber="430">28(27) Elytra without pale apical dot or with only narrow pale or light brown apical margin............................... 35</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1944,1966]" pageId="29" pageNumber="430">29(30) Hind tibiae and tarsi entirely black...................................................................... 31</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,1973,1995]" pageId="29" pageNumber="430">30(29) Hind tibiae and tarsi partly pale or brownish............................................................... 33</paragraph>
<paragraph blockId="29.[151,1437,213,2025]" box="[151,1436,2003,2025]" pageId="29" pageNumber="430">
31(32) Larger, TL = 14.5. Labrum transverse, LW/LL = 1.16..............................
<taxonomicName authority="Tan, Mo, Liang, 1991" authorityName="Tan, Mo, Liang" authorityYear="1991" box="[1101,1436,2003,2025]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="430" phylum="Arthropoda" rank="species" species="biserratus">
<emphasis box="[1101,1221,2003,2024]" italics="true" pageId="29" pageNumber="430">T. biserratus</emphasis>
<bibRefCitation author="Tan, J. - J. &amp; Mo, W. &amp; Liang, C." box="[1227,1436,2003,2025]" pageId="29" pageNumber="430" pagination="243 - 245" refId="ref27368" refString="Tan, J. - J., Mo, W. &amp; Liang, C. (1991) A new species of genus Therates Latreille from Guizhou province, China (Coleoptera: Cicindelidae). Sinozoologia, 5 (8), 243 - 245. [in Chinese, with English summary]" type="journal article" year="1991">Tan, Mo, Liang, 1991</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,151,173]" pageId="30" pageNumber="431">
32(31) Smaller, TL = 9.8–12.8. Labrum virtually as wide as length, LW/LL = 1.05......
<taxonomicName authority="W. Horn, 1902" authorityName="W. Horn" authorityYear="1902" box="[1038,1436,151,173]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
<emphasis box="[1038,1289,151,173]" italics="true" pageId="30" pageNumber="431">T. fruhstorferi fruhstorferi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,152,173]" pageId="30" pageNumber="431" pagination="65 - 75" refId="ref25658" refString="Horn, W. (1902) Neue Cicindeliden gesammelt von Fruhstorfer in Tonkin 1900. Deutsche Entomologische Zeitschrift, 1902, 65 - 75." type="journal article" year="1902">Horn, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
33(34) Apical dot approximately as wide as central dot (
<figureCitation box="[702,787,181,203]" captionStart="FIGURES 99–114" captionStartId="21.[152,273,1418,1442]" captionTargetBox="[155,1421,210,1377]" captionTargetId="figure-17@21.[151,1435,181,1394]" captionTargetPageId="21" captionText="FIGURES 99–114. Therates spp., left elytron: 99–102—T. vitalisi vitalisi (99 —Vietnam, Lao Cai; 100, 101—Vietnam, Sa Pa; 102—Laos, 15 km NW Louang Namtha); 103, 104—T. vitalisi ida (China, Kuatun); 105, 106—T. biserratus (105— China, Yinjiang County; 106—China, Fanjing Shan-Kuaichang); 107, 108—T. hunnanensis sp. n. (China, Shunhuangshan); 109–114—T. sauteri (109, 110, 112, 113—China, Taiwan, Taihorin; 111—China, Taiwan, Ilan Hsien; 114—China, Taiwan, Taipei Hsien); 100, 103, 106, 107, 109–111—males; 99, 101, 102, 104, 105, 108, 112–114—females; 99, 103, 107— holotypes; 105, 108—paratypes; 109, 110, 112, 113—paralectotypes" figureDoi="http://doi.org/10.5281/zenodo.7758896" httpUri="https://zenodo.org/record/7758896/files/figure.png" pageId="30" pageNumber="431">Fig. 108</figureCitation>
). Labrum with very large pale apical spot and narrower apex (
<figureCitation captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="30" pageNumber="431">Fig. 76</figureCitation>
)..................................................................................
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1245,1376,210,232]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="hunanensis" status="sp. n.">
<emphasis box="[1245,1376,210,232]" italics="true" pageId="30" pageNumber="431">T. hunanensis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1382,1436,210,231]" pageId="30" pageNumber="431">
<taxonomicNameLabel box="[1382,1436,210,231]" pageId="30" pageNumber="431" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
34(33) Apical dot clearly narrower than central dot (
<figureCitation box="[651,762,239,261]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="30" pageNumber="431">Figs 82, 85</figureCitation>
). Labrum with medium-sized or small pale apical spot and wider apex (
<figureCitation box="[233,378,269,291]" captionStart="FIGURES 50–57" captionStartId="15.[152,270,1909,1933]" captionTargetBox="[320,1268,195,1860]" captionTargetId="figure-17@15.[303,1285,181,1885]" captionTargetPageId="15" captionText="FIGURES 50–57. Therates fruhstorferi ssp., labrum: 50–55—T. fruhstorferi fruhstorferi (50, 51—Vietnam, Montes Mauson; 52—Vietnam, Na Hang; 53—Vietnam, Tam Dao; 54, 55—Laos, Ban Nape); 56, 57—T. fruhstorferi australis ssp. n. (56—Vietnam, Kon Ka Kinh N.P.; 57—Vietnam, Pak Khe River); 50, 52, 54, 56—males; 51, 53, 55, 57—females; 50, 51—paralectotypes; 56—holotype; 57—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758882" httpUri="https://zenodo.org/record/7758882/files/figure.png" pageId="30" pageNumber="431">Figs 51, 53, 55</figureCitation>
).....................................................
<taxonomicName authority="W. Horn, 1902" authorityName="W. Horn" authorityYear="1902" box="[1038,1436,269,291]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
<emphasis box="[1038,1289,269,291]" italics="true" pageId="30" pageNumber="431">T. fruhstorferi fruhstorferi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,269,290]" pageId="30" pageNumber="431" pagination="65 - 75" refId="ref25658" refString="Horn, W. (1902) Neue Cicindeliden gesammelt von Fruhstorfer in Tonkin 1900. Deutsche Entomologische Zeitschrift, 1902, 65 - 75." type="journal article" year="1902">Horn, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
35(36) Elytra shining green, without humeral lunule and basal dot (
<figureCitation box="[801,873,298,320]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="30" pageNumber="431">Fig. 98</figureCitation>
). Hind tarsi pale except dark apices of all tarsomeres (
<figureCitation captionStart="FIGURES 1–14" captionStartId="3.[152,270,1637,1661]" captionTargetBox="[159,1427,717,1605]" captionTargetId="figure-307@3.[151,1435,705,1613]" captionTargetPageId="3" captionText="FIGURES 1–14. Therates spp., habitus: 1–6—T. fruhstorferi fruhstorferi (1–3—Vietnam, Montes Mouson; 4—Vietnam, Na Hang; 5—Vietnam, Tam Dao; 6—Laos, Ban Nape); 7, 8—T. fruhstorferi australis ssp. n. (Vietnam, Kon Ka Kinh); 9, 10—T. pseudovitalisi sp. n. (9—Laos, Bang Son Cha; 10—Laos, Ban Bun Neua); 11—T. pseudovitalisi tibetana ssp. n. (China, Xia- Cha-Yu); 12—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 13, 14—T. motoensis (China, Hanmi); 1, 2, 4, 5, 7, 9–13— males; 3, 6, 8, 14—females; 1, 2—lectotype; 3—paralectotype; 7, 9, 11, 12—holotypes; 8, 10—paratypes; 1, 3–14—habitus; 2—type labels." figureDoi="http://doi.org/10.5281/zenodo.7758874" httpUri="https://zenodo.org/record/7758874/files/figure.png" pageId="30" pageNumber="431">Fig. 14</figureCitation>
)..............................................................................
<taxonomicName authority="Tan, 1981" authorityName="Tan" authorityYear="1981" box="[1215,1436,328,349]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="motoensis">
<emphasis box="[1215,1335,328,349]" italics="true" pageId="30" pageNumber="431">T. motoensis</emphasis>
<bibRefCitation author="Tan, J. - J." box="[1341,1436,328,349]" pageId="30" pageNumber="431" pagination="331 - 335" refId="ref27058" refString="Tan, J. - J. (1981) Cicindelidae. In: Insects of Xizang (Tibet). Vol. I. Comprehensive Scientific Expedition to the Qinghai-Xizang Plateau, Chinese Academy of Sciences. Science Press, Beijing, pp. 331 - 335. [in Chinese, with English summary]" type="book chapter" year="1981">Tan, 1981</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,357,379]" pageId="30" pageNumber="431">36(35) Elytra shining violet-black or blue-black with purple lustre, at least with basal portion of humeral lunule.............. 37</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,386,408]" pageId="30" pageNumber="431">
37(38) Hind tibiae and tarsi partly pale or brown.................................
<taxonomicName authority="W. Horn, 1902" authorityName="W. Horn" authorityYear="1902" box="[1038,1436,386,408]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
<emphasis box="[1038,1289,386,408]" italics="true" pageId="30" pageNumber="431">T. fruhstorferi fruhstorferi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,386,408]" pageId="30" pageNumber="431" pagination="65 - 75" refId="ref25658" refString="Horn, W. (1902) Neue Cicindeliden gesammelt von Fruhstorfer in Tonkin 1900. Deutsche Entomologische Zeitschrift, 1902, 65 - 75." type="journal article" year="1902">Horn, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,415,437]" pageId="30" pageNumber="431">38(37) Hind tibiae and tarsi entirely black, hind tarsi in some specimens with one or two brownish basal tarsomeres........... 39</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,445,467]" pageId="30" pageNumber="431">39(40) Middle tibiae partly pale or brownish in proximal portion. Elytra with narrow pale or brown apical margins............ 41</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,474,496]" pageId="30" pageNumber="431">40(39) Middle tibiae black or brown-black. Elytra with black apical margins........................................... 47</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,503,525]" pageId="30" pageNumber="431">
41(42) Antennomeres 9–10 slightly dilated and just a little wider than 8 one (
<figureCitation box="[897,1114,503,525]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="30" pageNumber="431">Figs 29, 31, 33, 36, 38</figureCitation>
)....................... 45
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,533,555]" pageId="30" pageNumber="431">
42(41) Antennomeres 9–10 distinctly dilated and clearly wider than 8 one (
<figureCitation box="[878,987,533,555]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="30" pageNumber="431">Figs 45, 47</figureCitation>
)............
<taxonomicName authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[1154,1390,533,555]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="vitalisi">
<emphasis box="[1154,1243,533,555]" italics="true" pageId="30" pageNumber="431">T. vitalisi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1281,1390,533,554]" pageId="30" pageNumber="431" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
(43)
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
43(44) Labrum with relatively small yellow apical spot (
<figureCitation box="[710,825,562,584]" captionStart="FIGURES 65–72" captionStartId="17.[152,270,1944,1968]" captionTargetBox="[283,1300,190,1909]" captionTargetId="figure-17@17.[267,1320,181,1920]" captionTargetPageId="17" captionText="FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype." figureDoi="http://doi.org/10.5281/zenodo.7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="30" pageNumber="431">Figs 66, 68</figureCitation>
), in some specimens entirely black (
<figureCitation box="[1183,1257,562,584]" captionStart="FIGURES 65–72" captionStartId="17.[152,270,1944,1968]" captionTargetBox="[283,1300,190,1909]" captionTargetId="figure-17@17.[267,1320,181,1920]" captionTargetPageId="17" captionText="FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype." figureDoi="http://doi.org/10.5281/zenodo.7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="30" pageNumber="431">Fig. 67</figureCitation>
). Fore and middle femora pale-yellow..........................................................
<taxonomicName authority="W. Horn, 1913" authorityName="W. Horn" authorityYear="1913" box="[1129,1436,591,613]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="vitalisi">
<emphasis box="[1129,1289,591,613]" italics="true" pageId="30" pageNumber="431">T. vitalisi vitalisi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,592,613]" pageId="30" pageNumber="431" pagination="362 - 366" refId="ref25772" refString="Horn, W. (1913) Materiaux pour servir a l'etude de la faune entomologique de l'Indochine. Cicindelinae. Annales de la Societe Entomologique de Belgique, 57, 362 - 366. https: // doi. org / 10.5962 / bhl. part. 4597" type="journal article" year="1913">Horn, 1913</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" box="[151,1436,621,643]" pageId="30" pageNumber="431">
44(43) Labrum with medium-sized yellow apical spot (
<figureCitation box="[686,758,621,643]" captionStart="FIGURES 65–72" captionStartId="17.[152,270,1944,1968]" captionTargetBox="[283,1300,190,1909]" captionTargetId="figure-17@17.[267,1320,181,1920]" captionTargetPageId="17" captionText="FIGURES 65–72. Therates vitalisi ssp. labrum: 65–70—T. vitalisi vitalisi (65, 67—Vietnam, Sa Pa; 66—Vietnam, Lao Cai; 68—Laos, 15 km NW of Louang Namtha; 69—Vietnam, Pu Hoat N.P.; 70—Vietnam, Xuan Lien N.P.); 71, 72—T. vitalisi ida (China, Fukien); 65, 69–71—males; 66–68, 72—females; 66, 71—holotypes; 72 – paratype." figureDoi="http://doi.org/10.5281/zenodo.7758888" httpUri="https://zenodo.org/record/7758888/files/figure.png" pageId="30" pageNumber="431">Fig. 72</figureCitation>
). Fore and middle femora red-brown.....
<taxonomicName authority="Mandl, 1954" authorityName="Mandl" authorityYear="1954" box="[1182,1436,621,643]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="vitalisi" subSpecies="ida">
<emphasis box="[1182,1308,621,643]" italics="true" pageId="30" pageNumber="431">T. vitalisi ida</emphasis>
<bibRefCitation author="Mandl, K." box="[1314,1436,621,643]" pageId="30" pageNumber="431" pagination="157 - 161" refId="ref26194" refString="Mandl, K. (1954) Zur Kenntnis der Cicindeliden (Col.) Sud-Chinas. Bonner Zoologische Beitrage, 5, 157 - 161." type="journal article" year="1954">Mandl, 1954</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
45(46) Basal portion of humeral lunule and basal dot present, in some specimens basal dot small and poorly visible, rarely entirely absent. Labrum with wider apex (
<figureCitation box="[548,694,679,701]" captionStart="FIGURES 50–57" captionStartId="15.[152,270,1909,1933]" captionTargetBox="[320,1268,195,1860]" captionTargetId="figure-17@15.[303,1285,181,1885]" captionTargetPageId="15" captionText="FIGURES 50–57. Therates fruhstorferi ssp., labrum: 50–55—T. fruhstorferi fruhstorferi (50, 51—Vietnam, Montes Mauson; 52—Vietnam, Na Hang; 53—Vietnam, Tam Dao; 54, 55—Laos, Ban Nape); 56, 57—T. fruhstorferi australis ssp. n. (56—Vietnam, Kon Ka Kinh N.P.; 57—Vietnam, Pak Khe River); 50, 52, 54, 56—males; 51, 53, 55, 57—females; 50, 51—paralectotypes; 56—holotype; 57—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758882" httpUri="https://zenodo.org/record/7758882/files/figure.png" pageId="30" pageNumber="431">Figs 51, 53, 55</figureCitation>
)..........................
<taxonomicName authority="W. Horn, 1902" authorityName="W. Horn" authorityYear="1902" box="[1038,1436,679,701]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="fruhstorferi">
<emphasis box="[1038,1289,679,701]" italics="true" pageId="30" pageNumber="431">T. fruhstorferi fruhstorferi</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,680,701]" pageId="30" pageNumber="431" pagination="65 - 75" refId="ref25658" refString="Horn, W. (1902) Neue Cicindeliden gesammelt von Fruhstorfer in Tonkin 1900. Deutsche Entomologische Zeitschrift, 1902, 65 - 75." type="journal article" year="1902">Horn, 1902</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
46(45) Basal portion of humeral lunule and basal dot absent, rarely shoulders with very small indistinct brownish area. Labrum with narrower apex (
<figureCitation box="[376,486,738,760]" captionStart="FIGURES 73–80" captionStartId="18.[152,270,1909,1933]" captionTargetBox="[302,1276,200,1868]" captionTargetId="figure-28@18.[296,1292,181,1885]" captionTargetPageId="18" captionText="FIGURES 73–80. Therates spp. labrum: 73, 74—T. biserratus (73—China, Yinjiang County; 74—China, Fanjing Shan- Kuaichang); 75, 76—T. hunnanensis sp. n. (China, Shunhuangshan); 77–80—T. sauteri (77, 78 —China, Taiwan, Taihorin; 79—China, Taiwan, Ilan Hsien; 80—China, Taiwan, Taipei Hsien); 74, 75, 77, 79—males; 73, 76, 78, 80—females; 75— holotype; 73, 76—paratype; 77, 78—paralectotypes." figureDoi="http://doi.org/10.5281/zenodo.7758890" httpUri="https://zenodo.org/record/7758890/files/figure.png" pageId="30" pageNumber="431">Figs 78, 80</figureCitation>
)..........................................................
<taxonomicName authority="W. Horn, 1912" authorityName="W. Horn" authorityYear="1912" box="[1199,1436,738,760]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="sauteri">
<emphasis box="[1199,1289,739,760]" italics="true" pageId="30" pageNumber="431">T. sauteri</emphasis>
W.
<bibRefCitation author="Horn, W." box="[1327,1436,738,760]" pageId="30" pageNumber="431" pagination="129 - 139" refId="ref25746" refString="Horn, W. (1912) H. Sauter's Formosa-Ausbeute: Cicindelinae. Entomologische Mitteilungen, 1 (5), 129 - 139." type="journal article" year="1912">Horn, 1912</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
47(48) Antennomere 9 with rounded while antennomere 10 with indistinctly protruding downward lower anterior margin (
<figureCitation box="[1352,1425,767,789]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="30" pageNumber="431">Fig. 33</figureCitation>
). Elytra with narrower apices (
<figureCitation box="[505,577,797,819]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="30" pageNumber="431">Fig. 89</figureCitation>
). Hind tibiae longer, HTbL/HTaL = 1.19.............
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[1143,1367,797,819]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="fruhstorferi" subSpecies="australis">
<emphasis box="[1143,1367,797,819]" italics="true" pageId="30" pageNumber="431">T. fruhstorferi australis</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1373,1436,797,818]" pageId="30" pageNumber="431">ssp. n.</emphasis>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
48(47) Antennomeres 9 and 10 with lower anterior margins clearly protruding downward (
<figureCitation box="[1021,1132,826,848]" captionStart="FIGURES 29–49" captionStartId="11.[152,270,1754,1778]" captionTargetBox="[250,1341,191,1691]" captionTargetId="figure-17@11.[240,1347,181,1730]" captionTargetPageId="11" captionText="FIGURES 29–49. Therates spp., left antennomeres 6–11: 29–32—T. fruhstorferi fruhstorferi (29, 30 – Vietnam, Na Hang; 31, 32 – Laos, Ban Nape); 33, 34—T. fruhstorferi australis ssp. n. (33—Vietnam, Kon Ka Kinh N.P.; 34—Vietnam, Pak Khe River); 35—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 36, 37—T. pseudovitalisi pseudovitalisi ssp. n. (36—Laos, Ban Song Cha; 37—Laos, Ban Bun Neua); 38, 39—T. pseudovitalisi tibetana ssp. nov. (China, Xia-Cha-Yu); 40, 41—T. motoensis (China, Hanmi); 42, 43—T. hunnanensis sp. n. (China, Shunhuangshan); 44—T. biserratus (China, Fanjing Shan-Kuaichang); 45, 46—T. vitalisi vitalisi (Vietnam, Xuan Son N.P.); 47—T. vitalisi ida (China, Fukien); 48, 49—T. sauteri (48—China, Taiwan, Taipei Hsien; 49—China, Taiwan, Ilan Hsien); 30, 32, 34, 35, 37, 39, 41, 43, 44, 46, 49—males; 29, 31, 33, 36, 38, 40, 42, 45, 47, 48—females; 33, 35, 37, 39, 41—holotypes; 34, 36, 38, 42—paratypes. Without scales." figureDoi="http://doi.org/10.5281/zenodo.7758880" httpUri="https://zenodo.org/record/7758880/files/figure.png" pageId="30" pageNumber="431">Figs 36, 38</figureCitation>
). Elytra with wider apices (
<figureCitation captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="30" pageNumber="431">Figs 92, 93, 95</figureCitation>
). Hind tibiae shorter, HTbL/HTaL = 1.29.....................................
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1173,1330,855,877]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="species" species="pseudovitalisi" status="sp. n.">
<emphasis box="[1173,1330,855,877]" italics="true" pageId="30" pageNumber="431">T. pseudovitalisi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1336,1390,856,877]" pageId="30" pageNumber="431">
<taxonomicNameLabel box="[1336,1390,856,877]" pageId="30" pageNumber="431" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(49)
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
49(50) Labrum entirely black (
<figureCitation box="[452,524,885,907]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="30" pageNumber="431">Fig. 59</figureCitation>
). Central dot narrow and distinctly oblique (
<figureCitation box="[949,1059,885,907]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="30" pageNumber="431">Figs 92, 93</figureCitation>
).................................................................................................
<taxonomicName authority="Matalin &amp; Wiesner, 2023" authorityName="Matalin &amp; Wiesner" authorityYear="2023" box="[1048,1344,914,936]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" status="ssp. nov." subSpecies="pseudovitalisi">
<emphasis box="[1048,1344,914,936]" italics="true" pageId="30" pageNumber="431">T. pseudovitalisi pseudovitalisi</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1350,1436,914,935]" pageId="30" pageNumber="431">
<taxonomicNameLabel box="[1350,1436,914,935]" pageId="30" pageNumber="431" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph blockId="30.[151,1437,151,995]" pageId="30" pageNumber="431">
50(49) Labrum black-brown with small indistinctly visible brown spot in anterior third (
<figureCitation box="[1021,1095,943,965]" captionStart="FIGURES 58–64" captionStartId="16.[152,270,1909,1933]" captionTargetBox="[356,1241,203,1829]" captionTargetId="figure-28@16.[314,1272,181,1885]" captionTargetPageId="16" captionText="FIGURES 58–64. Therates spp., labrum: 58, 59—T. pseudovitalisi pseudovitalisi ssp. n. (58—Laos, Ban Song Cha; 59— Laos, Ban Bun Neua); 60, 61—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 62, 63—T. motoensis (China, Hanmi); 64—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 58, 60, 62, 64—males; 59, 61, 63—females; 58, 60, 64—holotypes; 59, 61—paratypes." figureDoi="http://doi.org/10.5281/zenodo.7758884" httpUri="https://zenodo.org/record/7758884/files/figure.png" pageId="30" pageNumber="431">Fig. 61</figureCitation>
). Central dot practically transverse with clearly dilated lateral portion (
<figureCitation box="[566,635,973,995]" captionStart="FIGURES 81–98" captionStartId="19.[152,270,1318,1342]" captionTargetBox="[156,1427,198,1278]" captionTargetId="figure-17@19.[151,1435,181,1294]" captionTargetPageId="19" captionText="FIGURES 81–98. Therates spp., left elytron: 81–87—T. fruhstorferi fruhtorferi (81, 82—Vietnam, Montes Mauson; 83— Vietnam, Na Hang; 84—Vietnam, Pu Hoat N.P.; 85—Vietnam, Tam Dao; 86—Laos, Ban Nape—Kaev Nua Pass; 87—Laos, Ban Nape); 88, 89—T. fruhstorferi australis ssp. n. (88—Vietnam, Kon Ka Kinh N.P.; 89—Vietnam, Pak Khe River); 90– 93—T. pseudovitalisi pseudovitalisi ssp. n. (90, 93—Laos, Ban Song Cha; 91—Laos, Ban Bun Neua; 92—Laos, Oudom Xai); 94, 95—T. pseudovitalisi tibetana ssp. n. (China, Xia-Cha-Yu); 96—T. ziyardamensis sp. n. (Myanmar, Zi Yar Dam); 97, 98—T. motoensis (China, Hanmi); 81, 83, 84, 87, 88, 90, 91, 94, 96, 97—males; 82, 85, 86, 89, 92, 93, 95, 98—females; 81, 82—paralectotypes; 88, 90, 94, 96—holotypes; 89, 91–93, 95—paratype." figureDoi="http://doi.org/10.5281/zenodo.7758894" httpUri="https://zenodo.org/record/7758894/files/figure.png" pageId="30" pageNumber="431">Fig. 95</figureCitation>
).......................................
<taxonomicName authorityName="Matalin et Wiesner" authorityYear="2023" box="[1125,1367,973,995]" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="431" phylum="Arthropoda" rank="subSpecies" species="pseudovitalisi" subSpecies="tibetana">
<emphasis box="[1125,1367,973,995]" italics="true" pageId="30" pageNumber="431">T. pseudovitalisi tibetana</emphasis>
</taxonomicName>
<emphasis bold="true" box="[1373,1436,973,994]" pageId="30" pageNumber="431">ssp. n.</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>
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