743 lines
69 KiB
XML
743 lines
69 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zse.97.59559" ID-GBIF-Dataset="a466a75c-706c-4cbb-b591-99df3300f175" ID-GBIF-Taxon="177318570" ID-Pensoft-Pub="1860-0743-1-121" ID-Pensoft-UUID="D6B72C47EAF45F7A94A24A00BAFAF95B" ID-ZooBank="94AF3450CEAF4F0C8149C3F9BA9E405A" ModsDocID="1860-0743-1-121" checkinTime="1612984351702" checkinUser="pensoft" docAuthor="Molina-Acevedo, Isabel Cristina & Idris, Izwandy" docDate="2021" docId="D4D842D0F6E65F99BBBE65DD8E236CF1" docLanguage="en" docName="ZoosystEvol 97(1): 121-139" docOrigin="Zoosystematics and Evolution 97 (1)" docPubDate="2021-02-10" docSource="http://dx.doi.org/10.3897/zse.97.59559" docTitle="Marphysa fijiensis Molina-Acevedo & Idris, 2021, nom. nov." docType="treatment" docVersion="3" id="D6B72C47EAF45F7A94A24A00BAFAF95B" lastPageNumber="121" masterDocId="D6B72C47EAF45F7A94A24A00BAFAF95B" masterDocTitle="Unravelling the convoluted nomenclature of Marphysa simplex (Annelida, Eunicidae) with the proposal of a new name and the re-description of species" masterLastPageNumber="139" masterPageNumber="121" pageId="0" pageNumber="121" updateTime="1643426773792" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Unravelling the convoluted nomenclature of Marphysa simplex (Annelida, Eunicidae) with the proposal of a new name and the re-description of species</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Molina-Acevedo, Isabel Cristina</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-5487-895X</mods:nameIdentifier>
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<mods:affiliation>Estructura y Funcion del Bentos, Depto. Sistematica y Ecologia Acuatica, El Colegio de la Frontera Sur, Chetumal, Quintana Roo, Mexico & South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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<mods:nameIdentifier type="email">imolina@ecosur.edu.mx</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Idris, Izwandy</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-1516-8175</mods:nameIdentifier>
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<mods:affiliation>South China Sea Repository and Reference Centre, Institute of Oceanography and Environment, Universiti Malaysia Terengganu, 21030 Kuala Nerus, Terengganu, Malaysia</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2021</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2021-02-10</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>97</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>121</mods:start>
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<mods:end>139</mods:end>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.97.59559</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.97.59559</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-121</mods:identifier>
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<mods:identifier type="ZooBank">94AF3450CEAF4F0C8149C3F9BA9E405A</mods:identifier>
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<mods:identifier type="Pensoft-UUID">D6B72C47EAF45F7A94A24A00BAFAF95B</mods:identifier>
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<treatment ID-GBIF-Taxon="177318570" LSID="urn:lsid:plazi:treatment:D4D842D0F6E65F99BBBE65DD8E236CF1" httpUri="http://treatment.plazi.org/id/D4D842D0F6E65F99BBBE65DD8E236CF1" lastPageNumber="121" pageId="0" pageNumber="121">
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<subSubSection pageId="0" pageNumber="121" type="nomenclature">
|
||
<paragraph pageId="0" pageNumber="121">
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||
<taxonomicName LSID="D4D842D0-F6E6-5F99-BBBE-65DD8E236CF1" authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis" status="nom. nov.">Marphysa fijiensis</taxonomicName>
|
||
<taxonomicNameLabel pageId="0" pageNumber="121">nom. nov.</taxonomicNameLabel>
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">Figures 1</figureCitation>
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">, 2</figureCitation>
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of branchial filaments throughout the body. A. Holotype of Marphysa fijiensis nom. nov. (AMNH 1920 - 1530) with L 10: 8.2 mm, TL: 93 mm and 198 chaetigers; B. Paralectotype of Marphysa mossambica (Peters, 1854) (ZMB 47) with L 10: 10.3 mm, TL: 290 mm and 429 chaetigers." figureDoi="10.3897/zse.97.59559.figure3" httpUri="https://binary.pensoft.net/fig/507567" pageId="0" pageNumber="121">, 3</figureCitation>
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Limbate chaetae, chaetiger 8; B. Limbate chaetae, chaetiger 188; C. Thick isodont narrow, with short and slender teeth, chaetiger 3; D. Thick isodont wide with short and slender teeth, chaetiger 152; E. Thick isodont wide with short and slender teeth, chaetiger 188; F. Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40 μm (A, B, F, G); 12.5 μm (C-E)." figureDoi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121">, 4</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological features of Marphysa group A sensu Fauchald (1970). Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; COp: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p / a: present / absent; AR: anterior region; MR: median region; PR: posterior region; SH: subacicular hook. INSS: Isodont narrow with short and slender teeth; INLS: Isodont narrow with long and slender teeth; IWSS: Isodont wide with short and slender teeth; IWLS: Isodont wide with long and slender teeth; AWLT: Anodont wide with long and thick teeth." pageId="0" pageNumber="121">Table 1</figureCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Marphysa simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex" status="nom. nov.">Marphysa simplex</taxonomicName>
|
||
Treadwell, 1922: 151-152, pl. 5, figs 8-12, text-figs 39 (
|
||
<emphasis italics="true" pageId="0" pageNumber="121">non</emphasis>
|
||
Crossland, 1903); -
|
||
<bibRefCitation author="Hartman, O" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="121" pagination="239 - 310" refId="B23" refString="Hartman, O, 1956. Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology. Bulletin of the American Museum of Natural History 109 (2): 239 - 310" title="Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology." volume="109" year="1956">Hartman 1956</bibRefCitation>
|
||
: 254, 262, 268, 286.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="121">Material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Holotype.</emphasis>
|
||
Fiji · Suva Harbour; Apr-Jun, 1920; AMNH 1920-1530.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="materials_examined">
|
||
<paragraph pageId="0" pageNumber="121">Comparative material examined.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName class="Polychaeta" family="Eunicidae" genus="Eunice" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eunice mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Eunice mossambica</emphasis>
|
||
</taxonomicName>
|
||
Peters, 1854
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Mozambique · one specimen and vial with six parapodia;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">lectotype</emphasis>
|
||
ZMB 4005 · 3 adult specimens, same data as for the lectotype;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">paralectotypes</emphasis>
|
||
ZMB 47 · one specimen; same data as for the lectotype; ZMB F2046 · seven adult specimens; same data as for the lectotype; ZMB 4005.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Idris, Hutchings & Arshad" authorityYear="2014" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa moribidii" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="moribidii">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa moribidii</emphasis>
|
||
</taxonomicName>
|
||
Idris, Hutchings & Arshad, 2014
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Malaysia · two adult specimens; Pantai Kelanang, Morib, Selangor;
|
||
<geoCoordinate degrees="2" direction="north" minutes="45" orientation="latitude" precision="1" seconds="39.85" value="2.7610695">2°45'39.85"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="101" direction="east" minutes="26" orientation="longitude" precision="15" seconds="08" value="101.435555">101°26'08"E</geoCoordinate>
|
||
; in mangrove vegetation; 19 Jul 2011; I. Idris leg.;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">paratype</emphasis>
|
||
AM W.38690.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authority="Kinberg, 1865" authorityName="Kinberg" authorityYear="1865" class="Polychaeta" family="Eunicidae" genus="Nauphanta" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Nauphanta novaehollandiae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="novaehollandiae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Nauphanta novaehollandiae</emphasis>
|
||
</taxonomicName>
|
||
Kinberg, 1865
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Australia · one specimen divided into four vials, one of them with maxillary apparatus; Sydney Port Jackson;
|
||
<geoCoordinate degrees="33" direction="south" minutes="54" orientation="latitude" precision="925" value="-33.9">33°54'S</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="151" direction="east" minutes="11" orientation="longitude" precision="925" value="151.18333">151°11'E</geoCoordinate>
|
||
; Eugenie Epx. 1851-53;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">holotype</emphasis>
|
||
SMNH-type-432.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="etymology">
|
||
<paragraph pageId="0" pageNumber="121">Etymology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">The new name denotes the geographic region where the specimen was collected.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="description">
|
||
<paragraph pageId="0" pageNumber="121">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Holotype complete (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1A-F</figureCitation>
|
||
), ventrally dissected (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1C</figureCitation>
|
||
), with 198 chaetigers, L10 = 8.2 mm, W10 = 2.5 mm, TL = 93 mm. Anterior region with dorsum convex, flat ventre, body depressed from chaetiger 6 (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1D</figureCitation>
|
||
), widest at chaetiger 17, tapering after chaetiger 41.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121" start="Figure 1" startId="F1">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 1.</emphasis>
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov., holotype (AMNH 1920-1530).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, B.</emphasis>
|
||
Anterior end, dorsal view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Anterior end, ventral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D.</emphasis>
|
||
Anterior end, lateral view;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E.</emphasis>
|
||
Median region, lateral and ventral views;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F.</emphasis>
|
||
Pygidium, lateral view. Scale bars: 0.35 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A</emphasis>
|
||
); 2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B-D</emphasis>
|
||
); 0.5 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E, F</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Prostomium bilobed, 1.3 mm long, 1.6 mm wide; lobes frontally rounded; median sulcus shallow and deep ventrally (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1A, B</figureCitation>
|
||
). Prostomial appendages in semicircle, median antenna isolated by a gap. Palps reaching second peristomial ring; lateral antennae reaching first chaetiger; median antenna reaching second chaetiger. Palpophores and ceratophores ring-shaped, short, slender; palpostyles and ceratostyles tapering, thick. Eyes absent.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Peristomium (1.7 mm long, 2.3 mm wide) larger than prostomium, first ring twice as long as second ring, separation between rings distinct on all sides (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1A-C</figureCitation>
|
||
). Ventral lip dissected, with several shallow wrinkles (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">Maxillary apparatus lost, according to Treadwell with MF = 1+1, 5+5-6, 9+0 2+8, 1+1.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Branchiae pectinate with up to five long filaments, from chaetigers 22 to 184L-195R (Figs
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">2D, E</figureCitation>
|
||
). Six first branchiae with one filament; reaching maximum five filaments in chaetigers 79L-178L; last 12 branchiae with one filament (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of branchial filaments throughout the body. A. Holotype of Marphysa fijiensis nom. nov. (AMNH 1920 - 1530) with L 10: 8.2 mm, TL: 93 mm and 198 chaetigers; B. Paralectotype of Marphysa mossambica (Peters, 1854) (ZMB 47) with L 10: 10.3 mm, TL: 290 mm and 429 chaetigers." figureDoi="10.3897/zse.97.59559.figure3" httpUri="https://binary.pensoft.net/fig/507567" pageId="0" pageNumber="121">3A</figureCitation>
|
||
). Branchial filaments longer than dorsal cirri, except in first six and last 10 branchiae.
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121" start="Figure 2" startId="F2">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 2.</emphasis>
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov., holotype (AMNH 1920-1530).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A.</emphasis>
|
||
Chaetiger 3;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B.</emphasis>
|
||
Chaetiger 8;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Chaetiger 14;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D.</emphasis>
|
||
Chaetiger 123;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E.</emphasis>
|
||
Chaetiger 188.
|
||
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1854" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa mossambica</emphasis>
|
||
</taxonomicName>
|
||
(Peters, 1854), lectotype (ZMB 4005a).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F.</emphasis>
|
||
Chaetiger 3;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">G.</emphasis>
|
||
Chaetiger 8;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">H.</emphasis>
|
||
Chaetiger 15;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">I.</emphasis>
|
||
Chaetiger 293;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">J.</emphasis>
|
||
Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A-E</emphasis>
|
||
); 0.2 mm (
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F-J</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<caption doi="10.3897/zse.97.59559.figure3" httpUri="https://binary.pensoft.net/fig/507567" pageId="0" pageNumber="121" start="Figure 3" startId="F3">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 3.</emphasis>
|
||
Distribution of branchial filaments throughout the body.
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A.</emphasis>
|
||
Holotype of
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. (AMNH 1920-1530) with L10: 8.2 mm, TL: 93 mm and 198 chaetigers;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B.</emphasis>
|
||
Paralectotype of
|
||
<taxonomicName baseAuthorityName="Peters" baseAuthorityYear="1854" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa mossambica</emphasis>
|
||
</taxonomicName>
|
||
(Peters, 1854) (ZMB 47) with L10: 10.3 mm, TL: 290 mm and 429 chaetigers.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
First two parapodia smaller; best developed in chaetigers 4-21, following parapodia gradually decreasing in size. Dorsal cirri conical in all chaetigers; longer than ventral cirri in anterior and posterior chaetigers, of similar size in median chaetigers; best developed in chaetigers 3-30, gradually decreasing posteriorly (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">2A-E</figureCitation>
|
||
). Prechaetal lobes short, as transverse folds in first three chaetigers, following lobes with upper edge longer than lower; transverse folds in most posterior chaetigers (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">2A-E</figureCitation>
|
||
). Chaetal lobes rounded in most chaetigers, shorter than postchaetal lobe, with aciculae emerging dorsal to mid-line; from chaetiger 33, longer than other lobes, aciculae emerging in mid-line (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">2A-E</figureCitation>
|
||
). Postchaetal lobes slightly developed in first 55 chaetigers; ovoid in first two chaetigers, rounded in the following ones, progressively smaller from chaetiger 14; from chaetiger 56 inconspicuous (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">2A-E</figureCitation>
|
||
). Ventral cirri digitiform in first 13 chaetigers; in chaetigers 14 to 126 with rounded, poorly developed swollen base and digitiform tip; conical from chaetiger 127, gradually decreasing in size posteriorly (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">2A-E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Aciculae blunt, basally reddish and translucent distally; colourless in posterior chaetigers (Fig.
|
||
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Chaetiger 3; B. Chaetiger 8; C. Chaetiger 14; D. Chaetiger 123; E. Chaetiger 188. Marphysa mossambica (Peters, 1854), lectotype (ZMB 4005 a). F. Chaetiger 3; G. Chaetiger 8; H. Chaetiger 15; I. Chaetiger 293; J. Chaetiger 398. All chaetigers in anterior view. The colours in drawings indicate the prechaetal (light brown), chaetal (light yellow) and postchaetal (light green) lobes. Scale bars: 0.1 mm (A-E); 0.2 mm (F-J)." figureDoi="10.3897/zse.97.59559.figure2" httpUri="https://binary.pensoft.net/fig/507566" pageId="0" pageNumber="121">2A-E</figureCitation>
|
||
). First two chaetigers with two aciculae; in chaetigers 3-10 with three or four aciculae; in chaetigers 11-24 with four aciculae; in chaetigers 25-43 with three aciculae; in chaetigers 44-85 with two aciculae; from 86 with only one acicula. In median-posterior region, aciculae twice as wide as subacicular hook (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Limbate chaetae, chaetiger 8; B. Limbate chaetae, chaetiger 188; C. Thick isodont narrow, with short and slender teeth, chaetiger 3; D. Thick isodont wide with short and slender teeth, chaetiger 152; E. Thick isodont wide with short and slender teeth, chaetiger 188; F. Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40 μm (A, B, F, G); 12.5 μm (C-E)." figureDoi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121">4F, G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption doi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121" start="Figure 4" startId="F4">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Figure 4.</emphasis>
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov., holotype (AMNH 1920-1530).
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A.</emphasis>
|
||
Limbate chaetae, chaetiger 8;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">B.</emphasis>
|
||
Limbate chaetae, chaetiger 188;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C.</emphasis>
|
||
Thick isodont narrow, with short and slender teeth, chaetiger 3;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">D.</emphasis>
|
||
Thick isodont wide with short and slender teeth, chaetiger 152;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">E.</emphasis>
|
||
Thick isodont wide with short and slender teeth, chaetiger 188;
|
||
<emphasis bold="true" pageId="0" pageNumber="121">F.</emphasis>
|
||
Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="121">A, B, F, G</emphasis>
|
||
); 12.5
|
||
<normalizedToken originalValue="μm">μm</normalizedToken>
|
||
(
|
||
<emphasis bold="true" pageId="0" pageNumber="121">C-E</emphasis>
|
||
).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Limbate chaetae in supra- and subacicular positions. Limbate supracicular chaetae reduced in number around chaetiger 16, chaetae of two lengths in same chaetiger, with longer blades in dorsal position and with short blades in ventral position. Limbate subacicular of two lengths, with short blades in dorsal position and with longer blades in ventral position (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Limbate chaetae, chaetiger 8; B. Limbate chaetae, chaetiger 188; C. Thick isodont narrow, with short and slender teeth, chaetiger 3; D. Thick isodont wide with short and slender teeth, chaetiger 152; E. Thick isodont wide with short and slender teeth, chaetiger 188; F. Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40 μm (A, B, F, G); 12.5 μm (C-E)." figureDoi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121">4A, B</figureCitation>
|
||
). Two types of pectinate chaetae; in all chaetigers, 2-3 thick isodont narrow chaetae, with up to 16-18 teeth short and slender teeth (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Limbate chaetae, chaetiger 8; B. Limbate chaetae, chaetiger 188; C. Thick isodont narrow, with short and slender teeth, chaetiger 3; D. Thick isodont wide with short and slender teeth, chaetiger 152; E. Thick isodont wide with short and slender teeth, chaetiger 188; F. Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40 μm (A, B, F, G); 12.5 μm (C-E)." figureDoi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121">4C</figureCitation>
|
||
); in median-posterior chaetigers, 4-5 thick isodont wide chaetae, with up to 38-42 teeth short and slender teeth (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Limbate chaetae, chaetiger 8; B. Limbate chaetae, chaetiger 188; C. Thick isodont narrow, with short and slender teeth, chaetiger 3; D. Thick isodont wide with short and slender teeth, chaetiger 152; E. Thick isodont wide with short and slender teeth, chaetiger 188; F. Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40 μm (A, B, F, G); 12.5 μm (C-E)." figureDoi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121">4D, E</figureCitation>
|
||
); anodont pectinate chaetae not observed. Compound chaetae absent. Subacicular hooks starting from chaetigers 38R-39L, one per chaetiger, with discontinuous distribution, in last 25 chaetigers, the hooks are absent; unidentate in median region with one hood (possibly with second hood broken) (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Limbate chaetae, chaetiger 8; B. Limbate chaetae, chaetiger 188; C. Thick isodont narrow, with short and slender teeth, chaetiger 3; D. Thick isodont wide with short and slender teeth, chaetiger 152; E. Thick isodont wide with short and slender teeth, chaetiger 188; F. Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40 μm (A, B, F, G); 12.5 μm (C-E)." figureDoi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121">4F</figureCitation>
|
||
); bidentate in median-posterior region, translucent, with blunt teeth, distal tooth directed upwards, proximal tooth larger, directed laterally (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A. Limbate chaetae, chaetiger 8; B. Limbate chaetae, chaetiger 188; C. Thick isodont narrow, with short and slender teeth, chaetiger 3; D. Thick isodont wide with short and slender teeth, chaetiger 152; E. Thick isodont wide with short and slender teeth, chaetiger 188; F. Unidentate subacicular hook, chaetiger 123; Bidentate subacicular hook, chaetiger 152. Arrows in F: upper one indicates the acicula; lower one indicates the hood of the subacicular hook. Scale bars: 40 μm (A, B, F, G); 12.5 μm (C-E)." figureDoi="10.3897/zse.97.59559.figure4" httpUri="https://binary.pensoft.net/fig/507569" pageId="0" pageNumber="121">4G</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Pygidium with two pairs of anal cirri broken (Fig.
|
||
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Marphysa fijiensis nom. nov., holotype (AMNH 1920 - 1530). A, B. Anterior end, dorsal view; C. Anterior end, ventral view; D. Anterior end, lateral view; E. Median region, lateral and ventral views; F. Pygidium, lateral view. Scale bars: 0.35 mm (A); 2 mm (B-D); 0.5 mm (E, F)." figureDoi="10.3897/zse.97.59559.figure1" httpUri="https://binary.pensoft.net/fig/507565" pageId="0" pageNumber="121">1F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="distribution">
|
||
<paragraph pageId="0" pageNumber="121">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">Known only from the type locality.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="habitat">
|
||
<paragraph pageId="0" pageNumber="121">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
Uncertain. Possibly coral reefs or mudflats (
|
||
<bibRefCitation author="Treadwell, AL" journalOrPublisher="Carnegie Institute of Washington Publication" pageId="0" pageNumber="121" pagination="127 - 170" refId="B51" refString="Treadwell, AL, 1922. Leodicidae from Fiji and Samoa. Carnegie Institute of Washington Publication 312: 127 - 170" title="Leodicidae from Fiji and Samoa." volume="312" year="1922">Treadwell 1922</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="121" type="remarks">
|
||
<paragraph pageId="0" pageNumber="121">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
The first comment on the synonymy of
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. (as
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. simplex" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="simplex">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. simplex</emphasis>
|
||
</taxonomicName>
|
||
Treadwell, 1922) and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. mossambica</emphasis>
|
||
</taxonomicName>
|
||
was made by
|
||
<bibRefCitation author="Hartman, O" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="121" pagination="239 - 310" refId="B23" refString="Hartman, O, 1956. Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology. Bulletin of the American Museum of Natural History 109 (2): 239 - 310" title="Polychaetous annelids erected by Treadwell, 1891 to 1948, together with a brief chronology." volume="109" year="1956">Hartman (1956)</bibRefCitation>
|
||
. Subsequently,
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.2352.1.2" author="Glasby, CJ" journalOrPublisher="Zootaxa" pageId="0" pageNumber="121" pagination="29 - 45" refId="B19" refString="Glasby, CJ, Hutchings, PA, 2010. A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865. Zootaxa 2352: 29 - 45, DOI: https://doi.org/10.11646/zootaxa.2352.1.2" title="A new species of Marphysa Quatrefages, 1865 (Polychaeta: Eunicida: Eunicidae) from northern Australia and a review of similar taxa from the Indo-west Pacific, including the genus Nauphanta Kinberg, 1865." url="https://doi.org/10.11646/zootaxa.2352.1.2" volume="2352" year="2010">Glasby and Hutchings (2010)</bibRefCitation>
|
||
supported this idea, stating that the morphology described by Treadwell was very similar to the smaller-sized specimens of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. mossambica</emphasis>
|
||
</taxonomicName>
|
||
. However, after a detailed morphological comparison of both
|
||
<normalizedToken originalValue="species’">species'</normalizedToken>
|
||
type material, we found marked differences.
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. lacks eyes, whereas they are present in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. mossambica</emphasis>
|
||
</taxonomicName>
|
||
. Additionally,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. (L10: 8.2 mm) has a maximum number of five branchial filaments in the median region (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of branchial filaments throughout the body. A. Holotype of Marphysa fijiensis nom. nov. (AMNH 1920 - 1530) with L 10: 8.2 mm, TL: 93 mm and 198 chaetigers; B. Paralectotype of Marphysa mossambica (Peters, 1854) (ZMB 47) with L 10: 10.3 mm, TL: 290 mm and 429 chaetigers." figureDoi="10.3897/zse.97.59559.figure3" httpUri="https://binary.pensoft.net/fig/507567" pageId="0" pageNumber="121">3</figureCitation>
|
||
), whereas
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. mossambica</emphasis>
|
||
</taxonomicName>
|
||
(L10:8.5-11.5 mm), has a maximum number of eight branchial filaments in the mid-posterior region (Fig.
|
||
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Distribution of branchial filaments throughout the body. A. Holotype of Marphysa fijiensis nom. nov. (AMNH 1920 - 1530) with L 10: 8.2 mm, TL: 93 mm and 198 chaetigers; B. Paralectotype of Marphysa mossambica (Peters, 1854) (ZMB 47) with L 10: 10.3 mm, TL: 290 mm and 429 chaetigers." figureDoi="10.3897/zse.97.59559.figure3" httpUri="https://binary.pensoft.net/fig/507567" pageId="0" pageNumber="121">3</figureCitation>
|
||
). In
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov., the chaetal lobe is rounded in the anterior region and the postchaetal lobe is oval in the first three chaetigers. In contrast, in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. mossambica</emphasis>
|
||
</taxonomicName>
|
||
, the chaetal lobe is rectangular in the anterior region and the postchaetal lobe is digitiform in the first three chaetigers. Likewise,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. has the subacicular hook present from chaetiger 25, contrasting to chaetigers 35-65 in
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. mossambica</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<taxonomicName authorityName="Molina-Acevedo & Idris" authorityYear="2021" class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. resembles
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. moribidii" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="moribidii">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. moribidii</emphasis>
|
||
</taxonomicName>
|
||
Idris, Hutchings & Arshad, 2014 and
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. novaehollandiae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="novaehollandiae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. novaehollandiae</emphasis>
|
||
</taxonomicName>
|
||
(Kinberg, 1865) in lacking compound chaetae. However,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. lacks the peduncle in prostomial appendages, with swollen base in ventral cirri starting from chaetiger 14 and the acicula is twice as wide as the subacicular hook in the median-posterior region. In contrast,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. moribidii" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="moribidii">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. moribidii</emphasis>
|
||
</taxonomicName>
|
||
(L10: 12.2-20 mm) has a peduncle at the base of the palpo- and ceratostyles, bears ventral cirri with a swollen base starting from chaetiger 6 and has a subacicular hook similar in width to the acicula throughout the body. Furthermore,
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. has the prechaetal lobe as a transverse fold throughout the body, the chaetal lobe rounded throughout the body, the ventral cirri with a swollen base starting from chaetiger 14 and the subacicular hook starting from chaetiger 25; while
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. novaehollandiae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="novaehollandiae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. novaehollandiae</emphasis>
|
||
</taxonomicName>
|
||
(L10: 6.6-9.6 mm) has the prechaetal lobe with dorsal edge longer than the ventral side in the first chaetigers, the chaetal lobe rectangular in the anterior region, the ventral cirri with a swollen base starting from chaetiger 8 and the subacicular hook starting from chaetigers 39-42. The comparison of
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov. with related species is provided in Table
|
||
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Morphological features of Marphysa group A sensu Fauchald (1970). Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; COp: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p / a: present / absent; AR: anterior region; MR: median region; PR: posterior region; SH: subacicular hook. INSS: Isodont narrow with short and slender teeth; INLS: Isodont narrow with long and slender teeth; IWSS: Isodont wide with short and slender teeth; IWLS: Isodont wide with long and slender teeth; AWLT: Anodont wide with long and thick teeth." httpUri="http://table.plazi.org/id/AFFEBF2D11FC3EF1FF80D8CEB16CFB15" pageId="0" pageNumber="121" tableUuid="AFFEBF2D11FC3EF1FF80D8CEB16CFB15">1</tableCitation>
|
||
.
|
||
</paragraph>
|
||
<caption ID-Table-UUID="AFFEBF2D11FC3EF1FF80D8CEB16CFB15" httpUri="http://table.plazi.org/id/AFFEBF2D11FC3EF1FF80D8CEB16CFB15" pageId="0" pageNumber="121" start="Table 1" startId="T1">
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<emphasis bold="true" pageId="0" pageNumber="121">Table 1.</emphasis>
|
||
Morphological features of
|
||
<taxonomicName class="Polychaeta" family="Eunicidae" genus="Marphysa" kingdom="Animalia" lsidName="Marphysa" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">Marphysa</emphasis>
|
||
</taxonomicName>
|
||
group A sensu
|
||
<bibRefCitation author="Fauchald, K" journalOrPublisher="Allan Hancock Monographs in Marine Biology" pageId="0" pageNumber="121" pagination="1 - 335" refId="B12" refString="Fauchald, K, 1970. Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from Western Mexico. Allan Hancock Monographs in Marine Biology 5: 1 - 335" title="Polychaetous annelids of the families Eunicidae, Lumbrineridae, Iphitimidae, Arabellidae, Lysaretidae and Dorvilleidae from Western Mexico." volume="5" year="1970">Fauchald (1970)</bibRefCitation>
|
||
. Abbreviations: MF: Maxillary formula, roman numerals refer to number of maxilla; MxC: maxillary carriers; CIS: closing system; COp: cavity opening; PR-I: first peristomial ring; PR-II: second peristomial ring; Chaet: chaetiger; p/a: present/absent; AR: anterior region; MR: median region; PR: posterior region; SH: subacicular hook. INSS: Isodont narrow with short and slender teeth; INLS: Isodont narrow with long and slender teeth; IWSS: Isodont wide with short and slender teeth; IWLS: Isodont wide with long and slender teeth; AWLT: Anodont wide with long and thick teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph pageId="0" pageNumber="121">
|
||
<table pageId="0" pageNumber="121">
|
||
<tr pageId="0" pageNumber="121">
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">Morphological feature</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. moribidii" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="moribidii">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. moribidii</emphasis>
|
||
</taxonomicName>
|
||
Idris et al., 2014
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. mossambica" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="mossambica">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. mossambica</emphasis>
|
||
</taxonomicName>
|
||
(Peters, 1854)
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. novaehollandiae" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="novaehollandiae">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. novaehollandiae</emphasis>
|
||
</taxonomicName>
|
||
(Kinberg, 1865)
|
||
</th>
|
||
<th colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
<taxonomicName class="Polychaeta" genus="M." higherTaxonomySource="GBIF,CoL" kingdom="Animalia" lsidName="M. fijiensis" order="Eunicida" pageId="0" pageNumber="121" phylum="Annelida" rank="species" species="fijiensis">
|
||
<emphasis italics="true" pageId="0" pageNumber="121">M. fijiensis</emphasis>
|
||
</taxonomicName>
|
||
nom. nov.
|
||
</th>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Source of information</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">paratypes AM W. 38690; additional material</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">lectotype ZMB 4005a; paralectotypes ZMB 47; ZMB F2046, ZMB 4005</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">holotype SMNH-type-432; AM W.33021</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">holotype AMNH 1920-1530</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Size (mm): L10, W10</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">12.2-20, 6.3-8.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">8-11.5, 3.6-8.1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">6.6-9.6, 4-4.2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">8.2, 2.5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Prostomium: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bilobed</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Palps: reaching</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II or Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Lateral antennae: reaching</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">PR-II or Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">middle Chaet 1 or 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">middle Chaet 1</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Median antennae: reaching</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 1 or 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 2 or 3</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">middle Chaet 2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 2</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Peduncle in prostomial appendages</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Eyes</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">absent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MF: MII, MIII, MIV</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5-6+4-6, 7-8, 6+8-10</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5+5-6, 6-7, 3-4+8-9</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">4-5+6, 7, 5+8-9</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5+5-6, 9, 2+8</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MI vs. MxC: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2.4-2.8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2-3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
2.4-3.2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than MxC
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">?</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MI vs. CIS: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4.3-5.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
5-7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4.4-8
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than CIS
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">?</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MII vs. COp: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4.3-4.7
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
3.2-4
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
4.5-5.3
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
longer than COp
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">?</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Branchiae: shaped</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">pectinate</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Branchiae: start chaetiger; last chaetiger before pygidium</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">27-39; 15-37</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">23-48; 29-126</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">21-25; 15</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">22; 3</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Branchial filaments: numbers; length of the filaments</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">7-10; long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">7-8; long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">6-7; long</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">5; long</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Dorsal cirri: shaped</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical with wide base</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Prechaetal lobe: shaped</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: upper edge longer than lower, MR, PR: transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR, MR: upper edge longer than lower, PR: transverse fold</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR, MR: upper edge longer than lower, PR: transverse fold</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaetal lobe: shaped</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR: rectangular MR, PR: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">AR, MR: rectangular, PR: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">rounded</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Developed postchaetal lobe: end chaetiger</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">50-96</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">27-70</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">32-38</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">55</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Postchaetal lobe: shape in body regions</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 4: digitiform short, Chaet 4-10, 10: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 4: digitiform short, Chaet 4-10, 10: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 4: ovoid, Chaet 4-10, 10: rounded</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Chaet 4: ovoid, Chaet 4-10, 10: rounded</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Ventral cirri in first chaetigers: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">digitiform</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Ventral cirri with swollen base: start chaetiger; last chaetiger before pygidium</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">6; 62-96</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">7-9; 96-208</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">8; 41</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">14; 72</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Ventral cirri in most posterior chaetigers: shape</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">conical</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Aciculae: shape; colour</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">blunt, dark</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular limbate chaetae: (p/a); distribution</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present; all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present; all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present; all chaet</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">present; all chaet</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Pectinate chaetae: type in AR; MR, PR</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; IWSS, IWLS, AWLT</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; IWSS, IWLS, AWLT</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INLS; IWSS, IWLS,?</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">INSS; IWSS</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Pectinate chaetae: number per type</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">1-2; 3-4, 1-2, 1-2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">1-2; 2-3, 2-3, 1-2</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">1-2; 1-2, 3-4;?</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">2-3; 4-5</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Pectinate chaetae teeth: number per type</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">18; 52, 26, 7</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">18-19; 56, 27, 9-10</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">25; 50-51, 35;?</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">16-18; 38-42</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular hook: start chaetiger</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">56-65</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">35-65</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">39-42</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">38</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular hook: shape; colour</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, translucent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, translucent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">bidentate, translucent</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">MR: unidentate, PR: bidentate, translucent</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Width acicula vs. SH in MR-PR: proportion</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">similar width</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
Acicula 2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wider than SH
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
Acicula 2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wider than SH
|
||
</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">
|
||
Acicula 2
|
||
<normalizedToken originalValue="×">x</normalizedToken>
|
||
wider than SH
|
||
</td>
|
||
</tr>
|
||
<tr pageId="0" pageNumber="121">
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">Subacicular hook: distribution</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
<td colspan="1" pageId="0" pageNumber="121" rowspan="1">discontinuous</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |