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<document id="6C302246B88A9260667B6ADCC8B74423" ID-CLB-Dataset="299063" ID-DOI="10.11646/zootaxa.5474.2.6" ID-GBIF-Dataset="ccf79856-d670-4d44-affa-697cc6311fb9" ID-ISSN="1175-5326" ID-Zenodo-Dep="12567475" ID-ZooBank="E7DFCA2B-24F3-4F9B-B051-3C0C9ADE3230" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="guilherme" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1719479159966" checkinUser="plazi" docAuthor="Gualdrón-Díaz, Juliette C. &amp; Chani-Posse, Mariana R." docDate="2024" docId="F34AD63080082C75E5C5FE12FDD55209" docLanguage="en" docName="zootaxa.5474.2.6.pdf" docOrigin="Zootaxa 5474 (2)" docSource="http://dx.doi.org/10.11646/zootaxa.5474.2.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tetradonia vonbeerensis Gualdron-Diaz &amp; Chani-Posse 2024, sp. n." docType="treatment" docVersion="5" lastPageNumber="186" masterDocId="0F73AE48800A2C70E552FF80FFE95520" masterDocTitle="First record of the genus Tetradonia Wasmann, 1894 (Coleoptera: Staphylinidae: Aleocharinae) in Colombia, with description of two new species" masterLastPageNumber="191" masterPageNumber="181" pageNumber="183" updateTime="1719585302427" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="7D5E52E8E442E98CB57156D621F4FC32">First record of the genus Tetradonia Wasmann, 1894 (Coleoptera: Staphylinidae: Aleocharinae) in Colombia, with description of two new species</mods:title>
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<mods:name id="835C9CBA9C0265AA4C8E693EEC04BEDB" type="personal">
<mods:role id="61936EDFB55E230B0D1EB94B4E369CEB">
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<mods:namePart id="17C442D10FBF83DF16C069F9F41679A2">Gualdrón-Díaz, Juliette C.</mods:namePart>
<mods:affiliation id="6A865BCBE660138A70E5E3B0BA3C668A">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Aridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. &amp; Programa de Doctorado en Ciencias Biológicas, Universidad Nacional de Córdoba, Córdoba, Argentina.</mods:affiliation>
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<mods:namePart id="12B65B0C978C288C020C6B10D46F53C3">Chani-Posse, Mariana R.</mods:namePart>
<mods:affiliation id="011445DE7CCA0765CC5B849394D40375">Laboratorio de Entomología, Instituto Argentino de Investigaciones de las Zonas Aridas (IADIZA, CCT CONICET, Mendoza), Casilla de Correo 507, 5500 Mendoza, Argentina. &amp; Facultad de Ciencias Exactas y Naturales, Universidad Nacional de Cuyo, Mendoza, Argentina.</mods:affiliation>
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<mods:title id="A656596EAFD92C7BAACFB44C90BDA7F8">Zootaxa</mods:title>
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<mods:date id="E98ED32DEA6D6F4EA657EA471A98CD3A">2024</mods:date>
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<mods:number id="F232C6C9E23FC4A8E1DECE05F1D7D572">2024-06-26</mods:number>
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<mods:identifier id="C24FBDF5C9E8D8815D7EACCB0335F18E" type="DOI">10.11646/zootaxa.5474.2.6</mods:identifier>
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<treatment id="F34AD63080082C75E5C5FE12FDD55209" ID-DOI="http://doi.org/10.5281/zenodo.12567479" ID-GBIF-Taxon="233851334" ID-Zenodo-Dep="12567479" LSID="urn:lsid:plazi:treatment:F34AD63080082C75E5C5FE12FDD55209" httpUri="http://treatment.plazi.org/id/F34AD63080082C75E5C5FE12FDD55209" lastPageId="5" lastPageNumber="186" pageId="2" pageNumber="183">
<subSubSection id="33F934AD80082C72E5C5FE12FC67548D" box="[151,910,402,429]" pageId="2" pageNumber="183" type="nomenclature">
<paragraph id="7B5C672680082C72E5C5FE12FC67548D" blockId="2.[151,910,402,465]" box="[151,910,402,429]" pageId="2" pageNumber="183">
<heading id="2014D04A80082C72E5C5FE12FC67548D" bold="true" box="[151,910,402,429]" fontSize="11" level="1" pageId="2" pageNumber="183" reason="1">
<emphasis id="4997BB3480082C72E5C5FE12FC67548D" bold="true" box="[151,910,402,429]" pageId="2" pageNumber="183">
<taxonomicName id="BCE31CA580082C72E5C5FE12FCAD548C" authority="Gualdron-Diaz &amp; Chani-Posse" authorityName="Gualdron-Diaz &amp; Chani-Posse" authorityYear="2024" box="[151,836,402,429]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="vonbeerensis" status="sp. nov.">
<emphasis id="4997BB3480082C72E5C5FE12FE54548D" bold="true" box="[151,445,402,429]" italics="true" pageId="2" pageNumber="183">Tetradonia vonbeerensis</emphasis>
Gualdrón-Diaz &amp; Chani-Posse
</taxonomicName>
<taxonomicNameLabel id="52A4064F80082C72E61EFE13FC67548D" box="[844,910,403,429]" pageId="2" pageNumber="183" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="33F934AD80082C72E5C5FE37FEE454F1" box="[151,269,439,465]" pageId="2" pageNumber="183" type="description">
<paragraph id="7B5C672680082C72E5C5FE37FEE454F1" blockId="2.[151,910,402,465]" box="[151,269,439,465]" pageId="2" pageNumber="183">
<figureCitation id="E3D87BA380082C72E5C5FE37FEE454F1" box="[151,269,439,465]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–7" captionStart-2="FIGURES 8–17" captionStartId-0="2.[151,264,1826,1850]" captionStartId-1="3.[151,264,1380,1404]" captionStartId-2="4.[151,264,1831,1855]" captionTargetBox-0="[151,1436,666,1800]" captionTargetBox-1="[160,1431,237,1349]" captionTargetBox-2="[164,1416,187,1801]" captionTargetId-0="figure-179@2.[151,1436,666,1800]" captionTargetId-1="figure-55@3.[151,1436,231,1356]" captionTargetId-2="figure-25@4.[158,1430,181,1807]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURES 1–2. Tetradonia vonbeerensis sp. n., male holotype [MPUJ_ENT ©], (1) habitus dorsal; (2) habitus lateral." captionText-1="FIGURES 3–7. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (3) antenna; (4) labrum, dorsal view; (5) labium and mentum, ventral view; (6) right and left mandibles respectively, ventral view; (7) maxilla, dorsal view." captionText-2="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi-0="http://doi.org/10.5281/zenodo.12567477" figureDoi-1="http://doi.org/10.5281/zenodo.12567483" figureDoi-2="http://doi.org/10.5281/zenodo.12567487" httpUri-0="https://zenodo.org/record/12567477/files/figure.png" httpUri-1="https://zenodo.org/record/12567483/files/figure.png" httpUri-2="https://zenodo.org/record/12567487/files/figure.png" pageId="2" pageNumber="183">Figs. 1–17</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="33F934AD80082C72E5C5FE7EFD0357A5" pageId="2" pageNumber="183" type="materials_examined">
<paragraph id="7B5C672680082C72E5C5FE7EFD0357A5" blockId="2.[151,1437,510,645]" pageId="2" pageNumber="183">
<emphasis id="4997BB3480082C72E5C5FE7EFDC25738" bold="true" box="[151,555,510,537]" pageId="2" pageNumber="183">Material studied. Type material.</emphasis>
<materialsCitation id="CB8B6D7B80082C72E765FE7FFBCC5741" ID-GBIF-Occurrence="4899950305" collectingDate="2013-04-08" collectingDateMax="2013-04-12" collectingDateMin="2013-04-08" collectionCode="UV, MPUJ" collectorName="Duque N" country="Colombia" county="San Antonio del Tequendama" elevation="2221" latitude="4.6148667" location="Trampa" longLatPrecision="1" longitude="-74.31129" municipality="Reserva Chicaque" pageId="2" pageNumber="183" specimenCount="1" specimenCount-male="1" stateProvince="Cundinamarca" typeStatus="holotype">
<typeStatus id="A458D98480082C72E765FE7FFD765739" box="[567,671,511,537]" pageId="2" pageNumber="183" type="holotype">Holotype</typeStatus>
<specimenCount id="6DE5ACAF80082C72E7FFFE7FFD2D5739" box="[685,708,511,537]" count="1" pageId="2" pageNumber="183" type="male">♂</specimenCount>
,
<emphasis id="4997BB3480082C72E784FE7EFC9C5738" bold="true" box="[726,885,510,536]" pageId="2" pageNumber="183">
<collectingCountry id="03F427B680082C72E784FE7EFC9C5738" box="[726,885,510,536]" name="Colombia" pageId="2" pageNumber="183">COLOMBIA</collectingCountry>
</emphasis>
, with one labels: [white label] “COLOMBIA,
<collectingRegion id="B927A9C480082C72E5C5FDA3FED3571D" box="[151,314,547,573]" country="Colombia" name="Cundinamarca" pageId="2" pageNumber="183">Cundinamarca</collectingRegion>
/
<collectingCounty id="923D1FAA80082C72E409FDA3FD5C571D" box="[347,693,547,573]" pageId="2" pageNumber="183">San Antonio del Tequendama</collectingCounty>
/
<collectingMunicipality id="9B38FD5C80082C72E784FDA3FC4F571D" box="[726,934,547,573]" pageId="2" pageNumber="183">Reserva Chicaque</collectingMunicipality>
refugio /
<location id="7E3C31FD80082C72E171FDA3FB90571D" LSID="urn:lsid:plazi:treatment:F34AD63080082C75E5C5FE12FDD55209:7E3C31FD80082C72E171FDA3FB90571D" box="[1059,1145,547,573]" country="Colombia" county="San Antonio del Tequendama" latitude="4.6148667" longLatPrecision="1" longitude="-74.31129" municipality="Reserva Chicaque" name="Trampa" pageId="2" pageNumber="183" stateProvince="Cundinamarca">Trampa</location>
de luz
<collectionCode id="1DF2FFE380082C72E18EFDA3FAED571D" box="[1244,1284,547,573]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="2" pageNumber="183">UV</collectionCode>
,
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<geoCoordinate id="1ED701E180082C72E5C5FDC7FECC5741" box="[151,293,583,609]" degrees="74" direction="west" minutes="18.677" orientation="longitude" pageId="2" pageNumber="183" precision="1" value="-74.31129">74°18.677W</geoCoordinate>
/
<quantity id="BC1BCAC380082C72E466FDC7FE6D5741" box="[308,388,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.221" pageId="2" pageNumber="183" unit="m" value="2221.0">
<elevation id="F0CE801580082C72E466FDC7FE6D5741" box="[308,388,583,609]" metricMagnitude="3" metricUnit="m" metricValue="2.221" pageId="2" pageNumber="183" unit="m" value="2221.0">2221m</elevation>
</quantity>
,
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<collectingDate id="1F19B80E80082C72E4DCFDC7FDF75741" box="[398,542,583,609]" pageId="2" pageNumber="183" value="2013-04-08" valueMax="2013-04-12" valueMin="2013-04-08">8-12.iv.2013</collectingDate>
</date>
,
<collectorName id="D61602F080082C72E77BFDC7FD645741" box="[553,653,583,609]" pageId="2" pageNumber="183">Duque N</collectorName>
leg.” [white, printed] (
<collectionCode id="1DF2FFE380082C72E6D8FDC7FC3C5741" box="[906,981,583,609]" pageId="2" pageNumber="183">MPUJ</collectionCode>
_ENT)
</materialsCitation>
.
<materialsCitation id="CB8B6D7B80082C72E17CFDC7FD0F57A5" ID-GBIF-Occurrence="4899950302" collectingDate="2013-04-08" collectingDateMax="2013-04-12" collectingDateMin="2013-04-08" collectionCode="UV, MPUJ" collectorName="Duque N" country="Colombia" county="San Antonio del Tequendama" elevation="2221" latitude="4.6148667" location="Trampa" longLatPrecision="1" longitude="-74.31129" municipality="Reserva Chicaque" pageId="2" pageNumber="183" specimenCount="6" specimenCount-male="6" stateProvince="Cundinamarca" typeStatus="paratype">
<typeStatus id="A458D98480082C72E17CFDC7FB775741" box="[1070,1182,583,609]" pageId="2" pageNumber="183" type="paratype">Paratypes</typeStatus>
,
<specimenCount id="6DE5ACAF80082C72E1FAFDC7FB385741" box="[1192,1233,583,609]" count="6" pageId="2" pageNumber="183" type="male">6 ♂</specimenCount>
(
<quantity id="BC1BCAC380082C72E1B2FDC7FAE25741" box="[1248,1291,583,609]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="2" pageNumber="183" unit="in" value="3.0">3 in</quantity>
alcohol 96% and 2 dissected on slides): same data as for holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="2F9C37AE80082C72E5C5F8A2FAA0521A" ID-DOI="http://doi.org/10.5281/zenodo.12567477" ID-Zenodo-Dep="12567477" box="[151,1353,1826,1853]" httpUri="https://zenodo.org/record/12567477/files/figure.png" pageId="2" pageNumber="183" startId="2.[151,264,1826,1850]" targetBox="[151,1436,666,1800]" targetPageId="2" targetType="figure">
<paragraph id="7B5C672680082C72E5C5F8A2FAA0521A" blockId="2.[151,1353,1826,1853]" box="[151,1353,1826,1853]" pageId="2" pageNumber="183">
<emphasis id="4997BB3480082C72E5C5F8A2FED3521A" bold="true" box="[151,314,1826,1850]" pageId="2" pageNumber="183">FIGURES 1–2.</emphasis>
<taxonomicName id="BCE31CA580082C72E413F8A2FDD1521A" authorityName="Gualdrón-Díaz &amp; Chani-Posse" authorityYear="2024" box="[321,568,1826,1850]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="vonbeerensis" status="sp. nov.">
<emphasis id="4997BB3480082C72E413F8A2FDD1521A" box="[321,568,1826,1850]" italics="true" pageId="2" pageNumber="183">Tetradonia vonbeerensis</emphasis>
</taxonomicName>
<emphasis id="4997BB3480082C72E76DF8A2FD91521A" bold="true" box="[575,632,1826,1850]" pageId="2" pageNumber="183">
<taxonomicNameLabel id="52A4064F80082C72E76DF8A2FD91521A" box="[575,632,1826,1850]" pageId="2" pageNumber="183" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male holotype [MPUJ_ENT ©], (1) habitus dorsal; (2) habitus lateral.
</paragraph>
</caption>
<subSubSection id="33F934AD80082C73E595F80CFBAE55F5" lastPageId="3" lastPageNumber="184" pageId="2" pageNumber="183" type="diagnosis">
<paragraph id="7B5C672680082C73E595F80CFBAE55F5" blockId="2.[151,1437,1932,2031]" lastBlockId="3.[151,1436,151,213]" lastPageId="3" lastPageNumber="184" pageId="2" pageNumber="183">
<emphasis id="4997BB3480082C72E595F80CFEAB5286" bold="true" box="[199,322,1932,1958]" pageId="2" pageNumber="183">Diagnosis.</emphasis>
<taxonomicName id="BCE31CA580082C72E415F80DFCA65286" authority="Gualdron-Diaz &amp; Chani-Posse" authorityName="Gualdron-Diaz &amp; Chani-Posse" authorityYear="2024" box="[327,847,1932,1958]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="species" species="vonbeerensis" status="sp. n.">
<emphasis id="4997BB3480082C72E415F80DFE1A5286" box="[327,499,1932,1958]" italics="true" pageId="2" pageNumber="183">T. vonbeerensis</emphasis>
Gualdrón-Diaz &amp; Chani-Posse
</taxonomicName>
<emphasis id="4997BB3480082C72E607F80CFC7A5286" bold="true" box="[853,915,1932,1958]" pageId="2" pageNumber="183">
<taxonomicNameLabel id="52A4064F80082C72E607F80CFC7A5286" box="[853,915,1932,1958]" pageId="2" pageNumber="183" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
differs from the other species of
<taxonomicName id="BCE31CA580082C72E052F80CFA935286" authorityName="Wasmann" authorityYear="1894" box="[1280,1402,1932,1958]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="183" phylum="Arthropoda" rank="genus">
<emphasis id="4997BB3480082C72E052F80CFA935286" box="[1280,1402,1932,1958]" italics="true" pageId="2" pageNumber="183">Tetradonia</emphasis>
</taxonomicName>
by the following combination of characters: pronotum with a carina near to posterior margin and four long macrosetae along lateral margin (the first two setae long and the others short); elytral surface weakly punctate with a thin ridge along the lateral margin; sternite III with a very short extension from posterior angle and tergite III with two macrosetae, tergite VII with 4+4 macrosetae in a small, raised area postero-laterally.
</paragraph>
</subSubSection>
<caption id="2F9C37AE80092C73E5C5FAE4FAF75080" ID-DOI="http://doi.org/10.5281/zenodo.12567483" ID-Zenodo-Dep="12567483" httpUri="https://zenodo.org/record/12567483/files/figure.png" pageId="3" pageNumber="184" startId="3.[151,264,1380,1404]" targetBox="[160,1431,237,1349]" targetPageId="3" targetType="figure">
<paragraph id="7B5C672680092C73E5C5FAE4FAF75080" blockId="3.[151,1436,1380,1440]" pageId="3" pageNumber="184">
<emphasis id="4997BB3480092C73E5C5FAE4FED7505C" bold="true" box="[151,318,1380,1404]" pageId="3" pageNumber="184">FIGURES 3–7.</emphasis>
<taxonomicName id="BCE31CA580092C73E415FAE4FDAB505C" authorityName="Gualdrón-Díaz &amp; Chani-Posse" authorityYear="2024" box="[327,578,1380,1404]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="184" phylum="Arthropoda" rank="species" species="vonbeerensis" status="sp. nov.">
<emphasis id="4997BB3480092C73E415FAE4FDAB505C" box="[327,578,1380,1404]" italics="true" pageId="3" pageNumber="184">Tetradonia vonbeerensis</emphasis>
</taxonomicName>
<emphasis id="4997BB3480092C73E71EFAE4FD61505C" bold="true" box="[588,648,1380,1404]" pageId="3" pageNumber="184">
<taxonomicNameLabel id="52A4064F80092C73E71EFAE4FD61505C" box="[588,648,1380,1404]" pageId="3" pageNumber="184" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male paratype [MPUJ_ENT ©], (3) antenna; (4) labrum, dorsal view; (5) labium and mentum, ventral view; (6) right and left mandibles respectively, ventral view; (7) maxilla, dorsal view.
</paragraph>
</caption>
<subSubSection id="33F934AD80092C75E5C5FA67FE7F5075" lastPageId="5" lastPageNumber="186" pageId="3" pageNumber="184" type="description">
<paragraph id="7B5C672680092C73E5C5FA67FEC05321" blockId="3.[151,1437,1511,1970]" box="[151,297,1511,1537]" pageId="3" pageNumber="184">
<emphasis id="4997BB3480092C73E5C5FA67FEC05321" bold="true" box="[151,297,1511,1537]" pageId="3" pageNumber="184">Description.</emphasis>
</paragraph>
<paragraph id="7B5C672680092C73E595F98CFDEF536A" blockId="3.[151,1437,1511,1970]" pageId="3" pageNumber="184">Measurements: LPL: 1.43; LEL: 2.42; HW: 0.80; HL: 0.62; AL: 2.81; PW: 0.79; PL: 0.72; EL: 0.46; TL: 0.03; NW: 0.31; EtW: 0.74; EtL: 0.99.</paragraph>
<paragraph id="7B5C672680092C73E595F9D3FC5E5395" blockId="3.[151,1437,1511,1970]" pageId="3" pageNumber="184">
Body length,
<quantity id="BC1BCAC380092C73E432F9D3FDED534E" box="[352,516,1619,1646]" metricMagnitude="-3" metricUnit="m" metricValue="4.89" metricValueMax="5.25" metricValueMin="4.53" pageId="3" pageNumber="184" unit="mm" value="4.89" valueMax="5.25" valueMin="4.53">4.53–5.25 mm</quantity>
(
<figureCitation id="E3D87BA380092C73E741F9D3FD69534E" box="[531,640,1619,1646]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1826,1850]" captionTargetBox="[151,1436,666,1800]" captionTargetId="figure-179@2.[151,1436,666,1800]" captionTargetPageId="2" captionText="FIGURES 1–2. Tetradonia vonbeerensis sp. n., male holotype [MPUJ_ENT ©], (1) habitus dorsal; (2) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.12567477" httpUri="https://zenodo.org/record/12567477/files/figure.png" pageId="3" pageNumber="184">Figs. 1–2</figureCitation>
). Yellowish-brown but head dark brown; pronotum with carina near to posterior margin; abdomen with long erect dark brown setae and short recumbent light brown setae; head, pronotum and elytra with short recumbent light brown setae; whole body shining.
</paragraph>
<paragraph id="7B5C672680092C73E595F93FFE9C5292" blockId="3.[151,1437,1511,1970]" pageId="3" pageNumber="184">
Head capsule transverse (HW/HL = 1.29), as wide as pronotum at widest point (HW/PW = 1.01), surface smooth and glossy, covered with long thin recumbent light brown setae; eyes extremely large, occupying almost entire sides of head, notably much longer than temples (EL/TL=15.33) seen from above (
<figureCitation id="E3D87BA380092C73E15DF887FBBB5202" box="[1039,1106,1799,1826]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1826,1850]" captionTargetBox="[151,1436,666,1800]" captionTargetId="figure-179@2.[151,1436,666,1800]" captionTargetPageId="2" captionText="FIGURES 1–2. Tetradonia vonbeerensis sp. n., male holotype [MPUJ_ENT ©], (1) habitus dorsal; (2) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.12567477" httpUri="https://zenodo.org/record/12567477/files/figure.png" pageId="3" pageNumber="184">Fig. 1</figureCitation>
); antennae long, longer than head, pronotum and elytra combined (AL=
<quantity id="BC1BCAC380092C73E720F8ACFD395266" box="[626,720,1836,1862]" metricMagnitude="-3" metricUnit="m" metricValue="2.81" pageId="3" pageNumber="184" unit="mm" value="2.81">2.81mm</quantity>
) (
<figureCitation id="E3D87BA380092C73E7B4F8ABFCC05266" box="[742,809,1835,1862]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1380,1404]" captionTargetBox="[160,1431,237,1349]" captionTargetId="figure-55@3.[151,1436,231,1356]" captionTargetPageId="3" captionText="FIGURES 3–7. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (3) antenna; (4) labrum, dorsal view; (5) labium and mentum, ventral view; (6) right and left mandibles respectively, ventral view; (7) maxilla, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12567483" httpUri="https://zenodo.org/record/12567483/files/figure.png" pageId="3" pageNumber="184">Fig. 3</figureCitation>
); all antennomeres longer than wide; antennomere 1 and 11 almost the same size; antennomeres 2–11 slightly wider at apex; scape widened medially; pedicel with apical area translucent; antennomere 10 short (
<quantity id="BC1BCAC380092C73E70CF8F3FD2052AE" box="[606,713,1907,1934]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="184" unit="mm" value="0.16">0.16 mm</quantity>
), 2.37 x shorter than scape; last antennomere narrowed apically into a pointed apex.
</paragraph>
<caption id="2F9C37AE800E2C74E5C5F8A7FB7A52D3" ID-DOI="http://doi.org/10.5281/zenodo.12567487" ID-Zenodo-Dep="12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="4" pageNumber="185" startId="4.[151,264,1831,1855]" targetBox="[164,1416,187,1801]" targetPageId="4" targetType="figure">
<paragraph id="7B5C6726800E2C74E5C5F8A7FB7A52D3" blockId="4.[151,1437,1831,2035]" pageId="4" pageNumber="185">
<emphasis id="4997BB34800E2C74E5C5F8A7FEA1521F" bold="true" box="[151,328,1831,1855]" pageId="4" pageNumber="185">FIGURES 8–17.</emphasis>
<taxonomicName id="BCE31CA5800E2C74E41DF8A7FDA1521F" authorityName="Gualdrón-Díaz &amp; Chani-Posse" authorityYear="2024" box="[335,584,1831,1855]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="185" phylum="Arthropoda" rank="species" species="vonbeerensis" status="sp. nov.">
<emphasis id="4997BB34800E2C74E41DF8A7FDA1521F" box="[335,584,1831,1855]" italics="true" pageId="4" pageNumber="185">Tetradonia vonbeerensis</emphasis>
</taxonomicName>
<emphasis id="4997BB34800E2C74E71DF8A7FD60521F" bold="true" box="[591,649,1831,1855]" pageId="4" pageNumber="185">
<taxonomicNameLabel id="52A4064F800E2C74E71DF8A7FD60521F" box="[591,649,1831,1855]" pageId="4" pageNumber="185" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, male paratype [MPUJ_ENT ©], (8) abdominal sternites III
<emphasis id="4997BB34800E2C74E1A2F8A7FB14521F" bold="true" box="[1264,1277,1831,1855]" pageId="4" pageNumber="185">–</emphasis>
V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below.
</paragraph>
</caption>
<paragraph id="7B5C6726800F2C75E595FF17FCA757A5" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
Labrum transverse, 1.56 x as wide as long, with 18 setae on each side of midline, most of them on anterior half, lateral margins convex, anterior margin concave, and posterior margin nearly straight with two long apodemes (
<figureCitation id="E3D87BA3800F2C75E03DFF3BFF4155D9" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1380,1404]" captionTargetBox="[160,1431,237,1349]" captionTargetId="figure-55@3.[151,1436,231,1356]" captionTargetPageId="3" captionText="FIGURES 3–7. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (3) antenna; (4) labrum, dorsal view; (5) labium and mentum, ventral view; (6) right and left mandibles respectively, ventral view; (7) maxilla, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12567483" httpUri="https://zenodo.org/record/12567483/files/figure.png" pageId="5" pageNumber="186">Fig. 4</figureCitation>
); apico-medial margin of epipharynx not modified into setose or spinose process; basal region of epipharynx with four pores, medial region of epipharynx with approximately 28 pseudopores in an irregular array; lateral pores row extended to anterior 1/2–1/3 of epipharynx; labium with ligula bifid and short, apically rounded lobes; prementum with discal seta widely separated, medial pseudopore field with a few pseudopores near base, lateral pseudopore fields with two real pores and one setal pore; labial palpi 3-articled, first article long, subequal in length to 3, article 2 shorter in length, the first two articles bear several setae and some pores; mentum sub-trapezoidal, with 2 pseudopores, and several sensory pores primarily on each side of midline (
<figureCitation id="E3D87BA3800F2C75E6B7FE37FBC254F1" box="[997,1067,438,465]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1380,1404]" captionTargetBox="[160,1431,237,1349]" captionTargetId="figure-55@3.[151,1436,231,1356]" captionTargetPageId="3" captionText="FIGURES 3–7. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (3) antenna; (4) labrum, dorsal view; (5) labium and mentum, ventral view; (6) right and left mandibles respectively, ventral view; (7) maxilla, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12567483" httpUri="https://zenodo.org/record/12567483/files/figure.png" pageId="5" pageNumber="186">Fig. 5</figureCitation>
); mandibles nearly symmetrical, but right mandible with one large tooth, left mandible lacking teeth (
<figureCitation id="E3D87BA3800F2C75E6E9FE5BFBEA54D5" box="[955,1027,475,501]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1380,1404]" captionTargetBox="[160,1431,237,1349]" captionTargetId="figure-55@3.[151,1436,231,1356]" captionTargetPageId="3" captionText="FIGURES 3–7. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (3) antenna; (4) labrum, dorsal view; (5) labium and mentum, ventral view; (6) right and left mandibles respectively, ventral view; (7) maxilla, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12567483" httpUri="https://zenodo.org/record/12567483/files/figure.png" pageId="5" pageNumber="186">Fig. 6</figureCitation>
). Maxillae with galea elongate and densely setose at apex, lacinia elongate, longer and wider than galea, with some short setae near base, numerous long setae on middle third and densenly setose toward at apex; maxillary palpi with palpomere I tranverse and very short, palpomere II a bit longer than III, with few long setae, palpomere III a bit wider than II, with several long setae, apical palpomere much narrower, lacking setae (
<figureCitation id="E3D87BA3800F2C75E7AEFDEBFCA957A5" box="[764,832,619,645]" captionStart="FIGURES 3–7" captionStartId="3.[151,264,1380,1404]" captionTargetBox="[160,1431,237,1349]" captionTargetId="figure-55@3.[151,1436,231,1356]" captionTargetPageId="3" captionText="FIGURES 3–7. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (3) antenna; (4) labrum, dorsal view; (5) labium and mentum, ventral view; (6) right and left mandibles respectively, ventral view; (7) maxilla, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.12567483" httpUri="https://zenodo.org/record/12567483/files/figure.png" pageId="5" pageNumber="186">Fig. 7</figureCitation>
).
</paragraph>
<paragraph id="7B5C6726800F2C75E595FD0FFC1D56CD" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
Pronotum almost slightly wider than long (WP/LP= 1.09), distinctly margined; dorsal surface smoothly convex, with carina near to posterior margin (
<figureCitation id="E3D87BA3800F2C75E702FD33FD7F57ED" box="[592,662,691,717]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1826,1850]" captionTargetBox="[151,1436,666,1800]" captionTargetId="figure-179@2.[151,1436,666,1800]" captionTargetPageId="2" captionText="FIGURES 1–2. Tetradonia vonbeerensis sp. n., male holotype [MPUJ_ENT ©], (1) habitus dorsal; (2) habitus lateral." figureDoi="http://doi.org/10.5281/zenodo.12567477" httpUri="https://zenodo.org/record/12567477/files/figure.png" pageId="5" pageNumber="186">Fig. 1</figureCitation>
), covered with short recumbent light brown setae; with two short macrosetae along anterior margin and four long macrosetae along lateral margin (the first two setae long and the others short); anterior margin straight to slightly sinuous on midline, angles rounded; lateral margins nearly straight, slightly constricted in posterior third; posterior margin slightly convex; hypomera visible in lateral view; elytra longer than pronotum (EL/PL= 1.37), weakly punctate, shining, covered with short recumbent light brown setae and without macrosetae (
<figureCitation id="E3D87BA3800F2C75E4FFFCE7FE1656A1" box="[429,511,871,897]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 10</figureCitation>
), lateral margins broadly convex, with thin ridge along the lateral margins in dorsal view, and with anteromesial border densely scaled; hind wings fully developed, wing flabellum with 8 rays attached to short broad lobe (
<figureCitation id="E3D87BA3800F2C75E42CFC2FFE2A56E9" box="[382,451,943,969]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 9</figureCitation>
). Mesoventrite with intercoxal process extremely short and broad; metaventral process longer than mesoventral process, ending in a V shape with the apex rounded.
</paragraph>
<paragraph id="7B5C6726800F2C75E595FC77FC8051C9" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
Abdomen with tergite III–VI with two macrosetae; sternite III with very short extension from posterior angle (
<figureCitation id="E3D87BA3800F2C75E5CDFB9BFF1C5115" box="[159,245,1051,1077]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 8a</figureCitation>
); sternite IV with pair of gland reservoir on anterior margin (
<figureCitation id="E3D87BA3800F2C75E698FB9BFBF85115" box="[970,1041,1051,1077]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 8</figureCitation>
); tergite VII (
<figureCitation id="E3D87BA3800F2C75E1E5FB9BFAE35115" box="[1207,1290,1051,1077]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 11</figureCitation>
) with 4 + 4 macrosetae on postero-laterally raised area, 8+8 macrosetae along apical margin, and with dense apical comb consisting of
<specimenCount id="6DE5ACAF800F2C75E463FBE3FE77515D" box="[305,414,1123,1149]" count="2" pageId="5" pageNumber="186" type="generic" typeStatus="types">two types</specimenCount>
of teeth (
<figureCitation id="E3D87BA3800F2C75E75FFBE3FD84515D" box="[525,621,1123,1149]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 11a</figureCitation>
); tergite VIII (
<figureCitation id="E3D87BA3800F2C75E641FBE3FC8F515D" box="[787,870,1123,1149]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 12</figureCitation>
) with four distinct processes on posterior margin, two on each side of longitudinal midline, 4+4 long macrosetae in raised area and several short macrosetae on lateral margin; sternite VIII as in
<figureCitation id="E3D87BA3800F2C75E492FB2BFDFB51E5" box="[448,530,1195,1221]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 13</figureCitation>
; tergite and sternite IX (
<figureCitation id="E3D87BA3800F2C75E670FB2BFC9A51E5" box="[802,883,1195,1221]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 14</figureCitation>
) with 4 + 4 macrosetae on lateral lobes and 4 + 4 macrosetae and row of yellow fine setae at base on median lobe.
</paragraph>
<paragraph id="7B5C6726800F2C75E595FB73FB605011" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
Male. Aedeagus elongated and sclerotized (
<figureCitation id="E3D87BA3800F2C75E7E1FB73FCE1502D" box="[691,776,1266,1293]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 15</figureCitation>
); parameres, as in
<figureCitation id="E3D87BA3800F2C75E68FFB73FBD8502D" box="[989,1073,1267,1293]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 16</figureCitation>
; median lobe with bulbous base and apical portion gradually narrowed toward apex, with athetine bridge present (
<figureCitation id="E3D87BA3800F2C75E17BFA97FB955011" box="[1065,1148,1303,1329]" captionStart="FIGURES 8–17" captionStartId="4.[151,264,1831,1855]" captionTargetBox="[164,1416,187,1801]" captionTargetId="figure-25@4.[158,1430,181,1807]" captionTargetPageId="4" captionText="FIGURES 8–17. Tetradonia vonbeerensis sp. n., male paratype [MPUJ_ENT ©], (8) abdominal sternites III–V, ventral view, white head rows showing sternite IV with a pair of gland reservoir on the anterior margin; (8a) paratergite and sternite III, lateral view, white head row showing sternite III with a short extension from posterior angle; (9) wing flabellum, numbers in the photo showing the number of rays; (10) elytron, dorsal view; (11) tergite VII with a dense comb on the posterior margin (11a); (12) tergite VIII; (13) sternite VIII; (14) tergite and sternite IX; (15) aedeagus, lateral view; (16) parameres, lateral view above and dorsal view below (17) median lobe, ventral view above, dorsal view middle and lateral view below." figureDoi="http://doi.org/10.5281/zenodo.12567487" httpUri="https://zenodo.org/record/12567487/files/figure.png" pageId="5" pageNumber="186">Fig. 17</figureCitation>
).
</paragraph>
<paragraph id="7B5C6726800F2C75E595FABBFE7F5075" blockId="5.[151,1437,150,1833]" box="[199,406,1339,1365]" pageId="5" pageNumber="186">Female. Unknown</paragraph>
</subSubSection>
<subSubSection id="33F934AD800F2C75E595FADFFE325355" pageId="5" pageNumber="186" type="discussion">
<paragraph id="7B5C6726800F2C75E595FADFFF31530D" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
<emphasis id="4997BB34800F2C75E595FADFFED25059" bold="true" box="[199,315,1375,1401]" pageId="5" pageNumber="186">Remarks.</emphasis>
<taxonomicName id="BCE31CA5800F2C75E413FADFFE045059" authorityName="Gualdrón-Díaz &amp; Chani-Posse" authorityYear="2024" box="[321,493,1375,1401]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="vonbeerensis">
<emphasis id="4997BB34800F2C75E413FADFFE045059" box="[321,493,1375,1401]" italics="true" pageId="5" pageNumber="186">T. vonbeerensis</emphasis>
</taxonomicName>
is similar to
<taxonomicName id="BCE31CA5800F2C75E729FADFFC265059" authority="Borgmeier, 1959" authorityName="Borgmeier" authorityYear="1959" box="[635,975,1374,1401]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="4997BB34800F2C75E729FADFFCE45059" box="[635,781,1375,1401]" italics="true" pageId="5" pageNumber="186">T. alvarengai</emphasis>
Borgmeier, 1959
</taxonomicName>
sharing the presence of a thin ridge along the lateral margin of the elytra and eyes extremely large, occupying almost the entire sides of head. However,
<taxonomicName id="BCE31CA5800F2C75E0DAFA03FEC150E1" authorityName="Gualdrón-Díaz &amp; Chani-Posse" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="vonbeerensis">
<emphasis id="4997BB34800F2C75E0DAFA03FEC150E1" italics="true" pageId="5" pageNumber="186">T. vonbeerensis</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="BCE31CA5800F2C75E700FA27FD0D50E1" authorityName="Borgmeier" authorityYear="1959" box="[594,740,1447,1473]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="4997BB34800F2C75E700FA27FD0D50E1" box="[594,740,1447,1473]" italics="true" pageId="5" pageNumber="186">T. alvarengai</emphasis>
</taxonomicName>
by the number of macrosetae on the tergite III (
<taxonomicName id="BCE31CA5800F2C75E1A3FA27FA7550E1" authorityName="Gualdrón-Díaz &amp; Chani-Posse" authorityYear="2024" box="[1265,1436,1447,1473]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="vonbeerensis">
<emphasis id="4997BB34800F2C75E1A3FA27FA7550E1" box="[1265,1436,1447,1473]" italics="true" pageId="5" pageNumber="186">T. vonbeerensis</emphasis>
</taxonomicName>
has 2 macrosetae and
<taxonomicName id="BCE31CA5800F2C75E4C2FA4BFDCD50C5" authorityName="Borgmeier" authorityYear="1959" box="[400,548,1483,1509]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="4997BB34800F2C75E4C2FA4BFDCD50C5" box="[400,548,1483,1509]" italics="true" pageId="5" pageNumber="186">T. alvarengai</emphasis>
</taxonomicName>
has 4) and tergite VII (4+4 against 3+3 respectively) as well as by having the posterior margin of outer paratergite III not extended, opposite to
<taxonomicName id="BCE31CA5800F2C75E62BFA6FFBE45329" authorityName="Borgmeier" authorityYear="1959" box="[889,1037,1519,1545]" class="Insecta" family="Staphylinidae" genus="Tetradonia" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="186" phylum="Arthropoda" rank="species" species="alvarengai">
<emphasis id="4997BB34800F2C75E62BFA6FFBE45329" box="[889,1037,1519,1545]" italics="true" pageId="5" pageNumber="186">T. alvarengai</emphasis>
</taxonomicName>
where it is extended to form a long point.
</paragraph>
<paragraph id="7B5C6726800F2C75E595F9B6FE325355" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
<emphasis id="4997BB34800F2C75E595F9B6FDEA5370" bold="true" box="[199,515,1590,1616]" pageId="5" pageNumber="186">Geographical distribution.</emphasis>
<materialsCitation id="CB8B6D7B800F2C75E75BF9B7FE3E5355" ID-GBIF-Occurrence="4899950303" country="Colombia" location="San Antonio del Tequendama" pageId="5" pageNumber="186" specimenCount="1" stateProvince="Cundinamarca">
The new species is only known from its
<typeStatus id="A458D984800F2C75E690F9B7FC1B5371" box="[962,1010,1591,1617]" pageId="5" pageNumber="186">type</typeStatus>
locality,
<location id="7E3C31FD800F2C75E105F9B7FA745371" LSID="urn:lsid:plazi:treatment:F34AD63080082C75E5C5FE12FDD55209:7E3C31FD800F2C75E105F9B7FA745371" box="[1111,1437,1591,1617]" country="Colombia" name="San Antonio del Tequendama" pageId="5" pageNumber="186" stateProvince="Cundinamarca">San Antonio del Tequendama</location>
in
<collectingRegion id="B927A9C4800F2C75E5E6F9DBFEB35355" box="[180,346,1627,1653]" country="Colombia" name="Cundinamarca" pageId="5" pageNumber="186">Cundinamarca</collectingRegion>
,
<collectingCountry id="03F427B6800F2C75E437F9DBFE3E5355" box="[357,471,1627,1653]" name="Colombia" pageId="5" pageNumber="186">Colombia</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="33F934AD800F2C75E595F9FFFDD55209" pageId="5" pageNumber="186" type="etymology">
<paragraph id="7B5C6726800F2C75E595F9FFFC70539D" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
<emphasis id="4997BB34800F2C75E595F9FFFEA753B9" bold="true" box="[199,334,1663,1689]" pageId="5" pageNumber="186">Etymology.</emphasis>
The specific epithet honors Dr. Christoph von Beeren, a distinguished staphylinologist, who has contributed to the study of various aspects of myrmecophile biology.
</paragraph>
<paragraph id="7B5C6726800F2C75E595F947FDD55209" blockId="5.[151,1437,150,1833]" pageId="5" pageNumber="186">
<emphasis id="4997BB34800F2C75E595F947FDF953C0" bold="true" box="[199,528,1734,1761]" pageId="5" pageNumber="186">Natural history and habits.</emphasis>
<materialsCitation id="CB8B6D7B800F2C75E74BF947FDD15209" ID-GBIF-Occurrence="4899950304" collectionCode="UV" county="The" elevation="2221" location="San Antonio del Tequendama" municipality="Reserva Chicaque" pageId="5" pageNumber="186" specimenCount="1" stateProvince="Bogota">
The specimens were collected using an
<collectionCode id="1DF2FFE3800F2C75E6B2F947FBE153C1" box="[992,1032,1735,1761]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="5" pageNumber="186">UV</collectionCode>
light trap at
<quantity id="BC1BCAC3800F2C75E1CCF947FB1553C1" box="[1182,1276,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="2.221" pageId="5" pageNumber="186" unit="m" value="2221.0">
<elevation id="F0CE8015800F2C75E1CCF947FB1553C1" box="[1182,1276,1735,1761]" metricMagnitude="3" metricUnit="m" metricValue="2.221" pageId="5" pageNumber="186" unit="m" value="2221.0">2221 m</elevation>
</quantity>
.a.s.l.
<collectingCounty id="923D1FAA800F2C75E06AF947FA8D53C1" box="[1336,1380,1735,1761]" pageId="5" pageNumber="186">The</collectingCounty>
<typeStatus id="A458D984800F2C75E03EF947FA7553C1" box="[1388,1436,1735,1761]" pageId="5" pageNumber="186">type</typeStatus>
locality,
<collectingMunicipality id="9B38FD5C800F2C75E5A4F96BFE2A5225" box="[246,451,1771,1797]" pageId="5" pageNumber="186">Reserva Chicaque</collectingMunicipality>
, is located in the municipality of
<location id="7E3C31FD800F2C75E662F96BFB935225" LSID="urn:lsid:plazi:treatment:F34AD63080082C75E5C5FE12FDD55209:7E3C31FD800F2C75E662F96BFB935225" box="[816,1146,1771,1797]" county="The" municipality="Reserva Chicaque" name="San Antonio del Tequendama" pageId="5" pageNumber="186" stateProvince="Bogota">San Antonio del Tequendama</location>
, southwest of
<collectingRegion id="B927A9C4800F2C75E044F96BFA835225" box="[1302,1386,1771,1797]" country="Colombia" name="Bogota" pageId="5" pageNumber="186">Bogotá</collectingRegion>
, and is surrounded by high
<location id="7E3C31FD800F2C75E4C2F88FFDD15209" LSID="urn:lsid:plazi:treatment:F34AD63080082C75E5C5FE12FDD55209:7E3C31FD800F2C75E4C2F88FFDD15209" box="[400,568,1807,1833]" county="The" municipality="Reserva Chicaque" name="Andean Forest" pageId="5" pageNumber="186" stateProvince="Bogota">Andean Forest</location>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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