treatments-xml/data/09/4E/48/094E480B5C2DFFD6FF3E4EB66BE0D70B.xml

<document id="A2DF44E0EEF82C8AA4E54BD072EE21AB" ID-DOI="10.11646/zootaxa.3691.3.1" ID-GBIF-Dataset="ad034b98-4029-470c-9a6e-4b5d168b46e4" ID-ISSN="1175-5326" ID-Zenodo-Dep="248009" ID-ZooBank="88298B5D-2941-4A52-8C47-0330168EA240" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460650775633" checkinUser="plazi" docAuthor="Walter, David Evans" docDate="2013" docId="094E480B5C2DFFD6FF3E4EB66BE0D70B" docLanguage="en" docName="zt03691p323.pdf" docOrigin="Zootaxa 3691 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Reginacharlottia Walter, 2013, gen. nov." docType="treatment" docVersion="9" lastPageNumber="315" masterDocId="F57730735C2EFFD8FFA94C676921D33F" masterDocTitle="A new genus and family of sejine mites (Acari, Parasitiformes, Mesostigmata, Sejoidea) based on new species from Lord Howe Island and Brazil, and a redescription of Sejus americanus (Banks, 1902)" masterLastPageNumber="323" masterPageNumber="301" pageNumber="304" updateTime="1698337942205" updateUser="plazi">
<mods:mods id="5A03C3E4A49E9094A62D88C5F7A7D0FD" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="7FF937C383A9E344A4AF5B3D5E88C6B0">
<mods:title id="6AAB7E8CF4141AEDD731A55319E99B84">A new genus and family of sejine mites (Acari, Parasitiformes, Mesostigmata, Sejoidea) based on new species from Lord Howe Island and Brazil, and a redescription of Sejus americanus (Banks, 1902)</mods:title>
</mods:titleInfo>
<mods:name id="B8F3736A49A203B8C861A2379360B6E4" type="personal">
<mods:role id="7841DD2F7FA984189C62A1EE4916DB65">
<mods:roleTerm id="AB8B60CA675442F649AC36061D3B1F51">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="315108D295E36FCA0CDC51E0886BD401">Walter, David Evans</mods:namePart>
</mods:name>
<mods:typeOfResource id="79CBE63D7AAD8BF6403CD1C9A75B8F8B">text</mods:typeOfResource>
<mods:relatedItem id="C7A7FE988FBDDF165CF5504E1B4FBA81" type="host">
<mods:titleInfo id="E6E134494B331218F125ED865C30B9CF">
<mods:title id="5E9515B252E74742C358B0356A350AE9">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="23ED884E76DAE790311D750FC4816ED4">
<mods:date id="BD78150A227B47FE13ACF47529358ED5">2013</mods:date>
<mods:detail id="9792337847D53BAF1ADA4370EAB95BB2" type="volume">
<mods:number id="03A351B34322A2049BD858D9955AC6D9">3691</mods:number>
</mods:detail>
<mods:detail id="FD697EF0D857021346748BD6376F5AF0" type="issue">
<mods:number id="5A908F2737B04F1510D4B52CDDA2A77D">3</mods:number>
</mods:detail>
<mods:extent id="7A3E4657E94603B8EEE89908BC1E69BE" unit="page">
<mods:start id="E36D9A3A9B34DC64981183470CAEDAE8">301</mods:start>
<mods:end id="8225618110E59AF3B2A88C55E117924A">323</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5CFBD7DEDD7A68593A76D12C0832F828">journal article</mods:classification>
<mods:identifier id="AD64367A26739B48DC410514ABA05C3A" type="DOI">10.11646/zootaxa.3691.3.1</mods:identifier>
<mods:identifier id="3EC140A26704E4C24DF228227B096121" type="GBIF-Dataset">ad034b98-4029-470c-9a6e-4b5d168b46e4</mods:identifier>
<mods:identifier id="D9310CC30FBC262E6014D99F114C944F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="469381FAE9737E62A299B1D1BF0DB81F" type="Zenodo-Dep">248009</mods:identifier>
<mods:identifier id="3140FAE0559B6DAEFC254AD20D867C29" type="ZooBank">88298B5D-2941-4A52-8C47-0330168EA240</mods:identifier>
</mods:mods>
<treatment id="094E480B5C2DFFD6FF3E4EB66BE0D70B" ID-DOI="http://doi.org/10.5281/zenodo.6159956" ID-GBIF-Taxon="119572730" ID-Zenodo-Dep="6159956" LSID="urn:lsid:plazi:treatment:094E480B5C2DFFD6FF3E4EB66BE0D70B" httpUri="http://treatment.plazi.org/id/094E480B5C2DFFD6FF3E4EB66BE0D70B" lastPageId="14" lastPageNumber="315" pageId="3" pageNumber="304">
<subSubSection id="C9FDAA965C2DFFDBFF3E4EB66B94D012" pageId="3" pageNumber="304" type="nomenclature">
<paragraph id="8158F91D5C2DFFDBFF3E4EB668F9D1D5" blockId="3.[151,472,720,747]" box="[151,472,720,747]" pageId="3" pageNumber="304">
<heading id="DA104E715C2DFFDBFF3E4EB668F9D1D5" bold="true" box="[151,472,720,747]" fontSize="11" level="1" pageId="3" pageNumber="304" reason="1">
<emphasis id="B393250F5C2DFFDBFF3E4EB668F9D1D5" bold="true" box="[151,472,720,747]" pageId="3" pageNumber="304">
<taxonomicName id="46E7829E5C2DFFDBFF3E4EB66842D1D4" box="[151,355,721,747]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="304" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B393250F5C2DFFDBFF3E4EB66842D1D4" bold="true" box="[151,355,721,747]" italics="true" pageId="3" pageNumber="304">Reginacharlottia</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A8A098745C2DFFDBFEC34EB768F9D1D5" box="[362,472,720,746]" pageId="3" pageNumber="304" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8158F91D5C2DFFDBFF3E4F7E6B94D012" blockId="3.[151,693,792,814]" box="[151,693,792,814]" pageId="3" pageNumber="304">
<typeStatus id="5E5C47BF5C2DFFDBFF3E4F7E69E9D011" box="[151,200,793,814]" pageId="3" pageNumber="304">Type</typeStatus>
species: 
<taxonomicName id="46E7829E5C2DFFDBFE8F4F7F6B7FD011" box="[294,606,792,814]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="304" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C2DFFDBFE8F4F7F6B7FD011" box="[294,606,792,814]" italics="true" pageId="3" pageNumber="304">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C2DFFDBFDCD4F7F6B94D012" bold="true" box="[612,693,792,813]" pageId="3" pageNumber="304">
<taxonomicNameLabel id="A8A098745C2DFFDBFDCD4F7F6B94D012" box="[612,693,792,813]" pageId="3" pageNumber="304" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C2DFFDBFF3E4F3D6C59D68D" pageId="3" pageNumber="304" type="diagnosis">
<paragraph id="8158F91D5C2DFFDBFF3E4F3D6C59D68D" blockId="3.[151,1437,858,2034]" pageId="3" pageNumber="304">
<emphasis id="B393250F5C2DFFDBFF3E4F3D6830D04C" bold="true" box="[151,273,858,883]" pageId="3" pageNumber="304">Diagnosis.</emphasis>
<emphasis id="B393250F5C2DFFDBFE8B4F3C6B19D04C" box="[290,568,859,883]" italics="true" pageId="3" pageNumber="304">Adults, nymphs, larvae</emphasis>
: Chelicera with edentate, regressed fixed digit covered by expanded, membranous cheliceral seta and elongate plumose interdigital excrescence (
<figureCitation id="19DCE5985C2DFFDBFC464F196D66D0A9" box="[1007,1095,894,918]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="3" pageNumber="304">Figs 14</figureCitation>
, 
<figureCitation id="19DCE5985C2DFFDBFBFD4F196D53D0A9" box="[1108,1138,894,918]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="3" pageNumber="304">21</figureCitation>
, 
<figureCitation id="19DCE5985C2DFFDBFBD64F196DE6D0A9" box="[1151,1223,894,918]" captionStart="FIGURES 24 – 27" captionStartId="10.[151,264,1780,1803]" captionTargetBox="[161,1430,198,1732]" captionTargetId="figure@10.[151,1436,193,1733]" captionTargetPageId="10" captionText="FIGURES 24 – 27. Reginacharlottia lordhowensis sp. n., adult male. 24 Subcapitulum. 25 Dorsal view of gnathosoma. 26 Lateral view of palp. 27 Distal palptibia and palptarsus (arrow points to spur on palpal apotele). Scale bars 24 – 26 = 10 µm; 27 = 1 µm." httpUri="https://zenodo.org/record/248015/files/figure.png" pageId="3" pageNumber="304">24–25</figureCitation>
) issuing from near base of movable digit; pilus dentilis minute; palpal apotele with two subequal tines and strong dorsal spur; hypostomal seta 
<emphasis id="B393250F5C2DFFDBFEF34FA16857D0E1" box="[346,374,966,990]" italics="true" pageId="3" pageNumber="304">h1</emphasis>
flattened, membranous (
<figureCitation id="19DCE5985C2DFFDBFD314FA16BC5D0E1" box="[664,740,966,990]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="3" pageNumber="304">Figs 5</figureCitation>
, 
<figureCitation id="19DCE5985C2DFFDBFD5D4FA16A35D0E1" box="[756,788,966,990]" captionStart="FIGURES 24 – 27" captionStartId="10.[151,264,1780,1803]" captionTargetBox="[161,1430,198,1732]" captionTargetId="figure@10.[151,1436,193,1733]" captionTargetPageId="10" captionText="FIGURES 24 – 27. Reginacharlottia lordhowensis sp. n., adult male. 24 Subcapitulum. 25 Dorsal view of gnathosoma. 26 Lateral view of palp. 27 Distal palptibia and palptarsus (arrow points to spur on palpal apotele). Scale bars 24 – 26 = 10 µm; 27 = 1 µm." httpUri="https://zenodo.org/record/248015/files/figure.png" pageId="3" pageNumber="304">24</figureCitation>
); tritosternal base with short lateral extensions (
<figureCitation id="19DCE5985C2DFFDBFAEF4FA16CAFD0E1" box="[1350,1422,966,990]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="3" pageNumber="304">Fig. 7</figureCitation>
), laciniae completely divided; postanal seta absent; legs I antenniform, tarsus I subcylindrical, without apotele (
<figureCitation id="19DCE5985C2DFFDBFAC34F8D6994D719" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="3" pageNumber="304">Figs 12</figureCitation>
, 
<figureCitation id="19DCE5985C2DFFDBFF6B4869682DD719" box="[194,268,1038,1062]" captionStart="FIGURES 28 – 30" captionStartId="11.[151,264,877,900]" captionTargetBox="[151,1436,193,855]" captionTargetId="figure@11.[151,1436,193,856]" captionTargetPageId="11" captionText="FIGURES 28 – 30. Reginacharlottia lordhowensis sp. n., adult female. 28 Leg I, telofemur, genu, tibia, tarsus. 29 Leg II, anterolateral view of distal telofemur (t), genu (g, al 1 missing), tibia, tarsus. 30 Habitus. Scale bars 28 – 29 = 50 µm." httpUri="https://zenodo.org/record/248016/files/figure.png" pageId="3" pageNumber="304">28, 30</figureCitation>
). 
<emphasis id="B393250F5C2DFFDBFE8948696B3ED719" box="[288,543,1038,1062]" italics="true" pageId="3" pageNumber="304">Adult male and female</emphasis>
: anal opening (
<figureCitation id="19DCE5985C2DFFDBFD6648696A38D719" box="[719,793,1038,1062]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="3" pageNumber="304">Figs 6</figureCitation>
, 
<figureCitation id="19DCE5985C2DFFDBFC8F48696A60D719" box="[806,833,1038,1062]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="3" pageNumber="304">11</figureCitation>
) on small shield overarched by pair of hypertrophied setae, ringed by accretion-like array of small setae and scale-like platelets; at least seven pairs of setae in sternogenital region (
<figureCitation id="19DCE5985C2DFFDBFEEC483168EAD751" box="[325,459,1110,1134]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="3" pageNumber="304">Figs 16, 17</figureCitation>
, 
<figureCitation id="19DCE5985C2DFFDBFE73483168D8D751" box="[474,505,1110,1134]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="3" pageNumber="304">35</figureCitation>
); 
<emphasis id="B393250F5C2DFFDBFDB948316B18D751" box="[528,569,1110,1134]" italics="true" pageId="3" pageNumber="304">av4</emphasis>
, 
<emphasis id="B393250F5C2DFFDBFDE048316B50D751" box="[585,625,1110,1134]" italics="true" pageId="3" pageNumber="304">pv4</emphasis>
on intercalary sclerite on tarsus IV; coxa II with three setae including small, bifurcate dorsal seta; genu I with eight or nine setae; soft cuticle bearing numerous setae with ventral rib and overlapping dorsal scales, each seta inserted on ribbed base (
<figureCitation id="19DCE5985C2DFFDBFCF448F96A95D789" box="[861,948,1182,1206]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="3" pageNumber="304">Fig. 17</figureCitation>
); corniculi setiform (
<figureCitation id="19DCE5985C2DFFDBFB0248F96DD1D789" box="[1195,1264,1182,1206]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="3" pageNumber="304">Fig. 5</figureCitation>
c); tritosternal laciniae free, base slightly expanded laterally; metapodal plate fused to peritrematal shield. 
<emphasis id="B393250F5C2DFFDBFB5448A46CB2D7E4" box="[1277,1427,1219,1243]" italics="true" pageId="3" pageNumber="304">Adult female</emphasis>
: podonotal-mesonotal shield truncate (
<figureCitation id="19DCE5985C2DFFDBFDFB48816B8AD7C1" box="[594,683,1254,1278]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="3" pageNumber="304">Fig. 34</figureCitation>
) to eroded (
<figureCitation id="19DCE5985C2DFFDBFCEE48816AAFD7C1" box="[839,910,1254,1278]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="3" pageNumber="304">Fig. 1</figureCitation>
) posteriorly, projecting vertex-like over the gnathosoma (
<figureCitation id="19DCE5985C2DFFDBFE86496D6855D61D" box="[303,372,1290,1314]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="3" pageNumber="304">Fig. 2</figureCitation>
), joined to broad peritrematal-metapodal shield (
<figureCitation id="19DCE5985C2DFFDBFC09496D6AD5D61D" box="[928,1012,1290,1314]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="3" pageNumber="304">Fig. 15</figureCitation>
); pygidial shield reduced to remnant; free lateral, mesonotal, and posteromarginal shields or platelets absent; genital shield (
<figureCitation id="19DCE5985C2DFFDBFBC749496D96D679" box="[1134,1207,1326,1350]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="3" pageNumber="304">Figs 3</figureCitation>
, 
<figureCitation id="19DCE5985C2DFFDBFB6F49496DC2D679" box="[1222,1251,1326,1350]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="3" pageNumber="304">35</figureCitation>
) reduced, straplike plate, nude or with 1–2 setae. 
<emphasis id="B393250F5C2DFFDBFD8D49346BBED654" box="[548,671,1363,1387]" italics="true" pageId="3" pageNumber="304">Adult male</emphasis>
: podonotal-mesonotal shield extended posteriorly to cover most of idiosoma, creased at level coxae IV (
<figureCitation id="19DCE5985C2DFFDBFD9D49116BA9D6B1" box="[564,648,1398,1422]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="3" pageNumber="304">Fig. 19</figureCitation>
); venter covered by large, angular shield bearing numerous setae (
<figureCitation id="19DCE5985C2DFFDBFAC749116996D68D" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="3" pageNumber="304">Fig. 20</figureCitation>
); genital opening on intercoxal shield bearing seven pairs of setae and covered by two nude valves (
<figureCitation id="19DCE5985C2DFFDBFABE49FD6C4BD68D" box="[1303,1386,1434,1458]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="3" pageNumber="304">Fig. 18</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C2DFFDCFF6E49D86D81D103" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304" type="discussion">
<paragraph id="8158F91D5C2DFFDBFF6E49D86C6AD424" blockId="3.[151,1437,858,2034]" pageId="3" pageNumber="304">
<emphasis id="B393250F5C2DFFDBFF6E49D8681AD6E8" bold="true" box="[199,315,1471,1495]" pageId="3" pageNumber="304">Remarks.</emphasis>
<taxonomicName id="46E7829E5C2DFFDBFEE849D968DED6E9" box="[321,511,1470,1494]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B393250F5C2DFFDBFEE849D968DED6E9" box="[321,511,1470,1494]" italics="true" pageId="3" pageNumber="304">Reginacharlottia</emphasis>
</taxonomicName>
contains a perplexing mix of apparently primitive and markedly aberrant character states. The plumose, feather boa-like process on the chelicera is otherwise unknown in the Sejoidea, but similar to excrescences found on the fixed digits in the Trigynaspida, the only other suborder of 
<taxonomicName id="46E7829E5C2DFFDBFBC74A616C2AD521" box="[1134,1291,1542,1566]" class="Arachnida" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="304" phylum="Arthropoda" rank="order">Mesostigmata</taxonomicName>
to retain the intercalary sclerite on tarsus IV bearing setae 
<emphasis id="B393250F5C2DFFDBFD0A4A4C6BEAD57C" box="[675,715,1579,1603]" italics="true" pageId="3" pageNumber="304">av4</emphasis>
, 
<emphasis id="B393250F5C2DFFDBFD724A4C6A22D57C" box="[731,771,1579,1603]" italics="true" pageId="3" pageNumber="304">pv4</emphasis>
(also found in the other orders of 
<taxonomicName id="46E7829E5C2DFFDBFB264A4D6C16D57D" box="[1167,1335,1578,1602]" class="Arachnida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Parasitiformes" pageId="3" pageNumber="304" phylum="Arthropoda" rank="order">Parasitiformes</taxonomicName>
). Female trigynaspids, however, are usually well sclerotised, and even the exceptions with reduced sclerotisation (e.g. 
<taxonomicName id="46E7829E5C2DFFDBFF3E4A156804D5B5" box="[151,293,1650,1674]" class="Arachnida" family="Davacaridae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="304" phylum="Arthropoda" rank="family">Davacaridae</taxonomicName>
) have several shields covering the genital opening. Also, the excrescences in the Trigynaspida can usually be traced to openings in the movable cheliceral digit. No such opening has been observed in 
<emphasis id="B393250F5C2DFFDBFF3E4ADC687AD5EC" box="[151,347,1723,1747]" italics="true" pageId="3" pageNumber="304">
<taxonomicName id="46E7829E5C2DFFDBFF3E4ADC6876D5EC" box="[151,343,1723,1747]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus" status="gen. nov.">Reginacharlottia</taxonomicName>
,
</emphasis>
and the excrescence appears to issue from the arthrodial membrane at the articulation of the movable and fixed digits, as do the interdigital membranes in the 
<taxonomicName id="46E7829E5C2DFFDBFC2F4AB96D6ED5C9" box="[902,1103,1758,1782]" class="Arachnida" family="Heterozerconidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="304" phylum="Arthropoda" rank="family">Heterozerconidae</taxonomicName>
. Until the homology of these structures can be tested, I use the term interdigital cheliceral excrescence for the structure in the new genus.
</paragraph>
<paragraph id="8158F91D5C2DFFDCFF6E4B416D47D25B" blockId="3.[151,1437,858,2034]" lastBlockId="4.[151,1437,151,716]" lastPageId="4" lastPageNumber="305" pageId="3" pageNumber="304">
With the exception of the cheliceral excrescence, however, the new genus is in substantial agreement with the diagnosis for Sejida (= Sejoidea) in Lindquist 
<emphasis id="B393250F5C2DFFDBFD194B2B6BC5D45C" box="[688,740,1867,1891]" italics="true" pageId="3" pageNumber="304">et al</emphasis>
. (2009) with these exceptions, Sejida (vs 
<taxonomicName id="46E7829E5C2DFFDBFB644B2C6CABD45C" box="[1229,1418,1867,1891]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="3" pageNumber="320" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B393250F5C2DFFDBFB644B2C6CABD45C" box="[1229,1418,1867,1891]" italics="true" pageId="3" pageNumber="304">Reginacharlottia</emphasis>
</taxonomicName>
): venter of female with large ventri-anal shield (reduced to small ventri-anal shield); metapodal plates usually free and well developed (fused to peritrematal shield); genital shield large, usually bearing two or more pairs of setae (genital shield a strap-like remnant usually nude, but rarely captures 1–2 setae); palpal apotele 2-tined (2 tines + large dorsal spur). The palpal apotele is a supposed remnant of the palpal claw complex. The basic construction of the tarsal claws in arachnids is composed of three parts, a pair of lateral claws and a central claw or pad. Usually the palptarsal apotele has a maximum of three tines in the 
<taxonomicName id="46E7829E5C2AFFDCFC974CDB6AFAD3EB" box="[830,987,188,212]" class="Arachnida" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="305" phylum="Arthropoda" rank="order">Mesostigmata</taxonomicName>
(four in some 
<taxonomicName id="46E7829E5C2AFFDCFB204CDB6C25D3EB" box="[1161,1284,188,212]" class="Arachnida" family="Veigaiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="305" phylum="Arthropoda" rank="family">Veigaiidae</taxonomicName>
), presumably representing the plesiotypic tridactyl condition. In the Sejida, and convergently in other groups of 
<taxonomicName id="46E7829E5C2AFFDCFB514CB86CB9D3C8" box="[1272,1432,223,247]" class="Arachnida" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="305" phylum="Arthropoda" rank="order">Mesostigmata</taxonomicName>
, this may be reduced to two tines and sometimes a basal spur is present that appears to be the remnant of the missing tine. However, although not previously reported, 
<taxonomicName id="46E7829E5C2AFFDCFD6D4D4F6D15D200" authority="Banks, 1902" authorityName="Banks" authorityYear="1902" box="[708,1076,295,320]" class="Arachnida" family="Sejidae" genus="Sejus" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="B393250F5C2AFFDCFD6D4D4F6AABD27F" box="[708,906,296,320]" italics="true" pageId="4" pageNumber="305">Sejus americanus</emphasis>
(Banks, 1902)
</taxonomicName>
, has a similar spur on its palpal apotele (
<figureCitation id="19DCE5985C2AFFDCFF5E4D2B686BD25B" box="[247,330,332,356]" captionStart="FIGURES 41 – 43" captionStartId="17.[151,275,1891,1916]" captionTargetBox="[168,1421,213,1866]" captionTargetId="figure@17.[151,1436,193,1870]" captionTargetPageId="17" captionText="FIGURES 41 – 43. Sejus americanus (Banks, 1902) adult female. 41 Hypostome showing membranous seta h 1. 42 Fixed and movable digits of chelicera. 43 Palptarsal apotele (arrow points to spur, middle tine ~ 25 µm long). Scale bars 41 = 10 µm, 42 = 50 µm." httpUri="https://zenodo.org/record/248020/files/figure.png" pageId="4" pageNumber="305">Fig. 43</figureCitation>
), so this character state is present in the traditional Sejida (= Sejoidea).
</paragraph>
<paragraph id="8158F91D5C2AFFDCFF6E4D086D81D103" blockId="4.[151,1437,151,716]" pageId="4" pageNumber="305">
Other character states also suggest that 
<taxonomicName id="46E7829E5C2AFFDCFD304D176A76D2B7" box="[665,855,368,392]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="320" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B393250F5C2AFFDCFD304D176A76D2B7" box="[665,855,368,392]" italics="true" pageId="4" pageNumber="305">Reginacharlottia</emphasis>
</taxonomicName>
belongs in the Sejoidea. For example, the strong tuberculate ornamentation of the shields (e.g. 
<figureCitation id="19DCE5985C2AFFDCFD0D4DF36A07D293" box="[676,806,404,428]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="4" pageNumber="305">Figs 22, 23</figureCitation>
) is characteristic of members of the Sejoidea, as is the podonotal-peritrematal shield in the female. Moreover, although the genital shield in the female is strongly reduced (
<figureCitation id="19DCE5985C2AFFDCFF364DBB69CAD2CB" box="[159,235,476,500]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="4" pageNumber="305">Fig. 3</figureCitation>
), it bears a modified region on the anterior border found in members of the Sejoidea where the spermatophore is attached (see Fig. 50). Additionally, the anterior hypostomal seta (
<emphasis id="B393250F5C2AFFDCFBEB4E676D7FD127" box="[1090,1118,512,536]" italics="true" pageId="4" pageNumber="305">h1</emphasis>
) is typically modified into a pair of closely inserted and largely membranous structures in the Sejoidea (e.g. 
<figureCitation id="19DCE5985C2AFFDCFBBA4E436D49D103" box="[1043,1128,548,572]" captionStart="FIGURES 41 – 43" captionStartId="17.[151,275,1891,1916]" captionTargetBox="[168,1421,213,1866]" captionTargetId="figure@17.[151,1436,193,1870]" captionTargetPageId="17" captionText="FIGURES 41 – 43. Sejus americanus (Banks, 1902) adult female. 41 Hypostome showing membranous seta h 1. 42 Fixed and movable digits of chelicera. 43 Palptarsal apotele (arrow points to spur, middle tine ~ 25 µm long). Scale bars 41 = 10 µm, 42 = 50 µm." httpUri="https://zenodo.org/record/248020/files/figure.png" pageId="4" pageNumber="305">Figs 41</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFBDD4E436DB2D103" box="[1140,1171,548,572]" captionStart="FIGURES 51 – 55" captionStartId="21.[151,264,1778,1801]" captionTargetBox="[157,1430,198,1747]" captionTargetId="figure@21.[151,1436,193,1757]" captionTargetPageId="21" captionText="FIGURES 51 – 55. Sejoidea, Queensland, Australia: 51 – 53 Epicroseius sp. (Sejidae). 51 Subcapitulum (arrow = h 1) 52 Chelicerae (c = corniculus, pd = pilus dentilis) 53 Male genital shield (gs arrow). 54 – 55 Asternolaelaps sp. (Ichthyostomatogasteridae). 54 Male genital shield (gs arrow). 55 Enlarged, bifurcate corniculus (arrow)." httpUri="https://zenodo.org/record/248023/files/figure.png" pageId="4" pageNumber="305">51</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C2AFFDCFF6E4E2F68A4D065" pageId="4" pageNumber="305" type="etymology">
<paragraph id="8158F91D5C2AFFDCFF6E4E2F6876D1F3" blockId="4.[151,1437,151,716]" pageId="4" pageNumber="305">
<emphasis id="B393250F5C2AFFDCFF6E4E2F686FD15F" bold="true" box="[199,334,584,608]" pageId="4" pageNumber="305">Etymology.</emphasis>
The new genus is feminine and named for Lord Richard Howe’s flagship the Queen Charlotte during the Battle of Ushant, an action claimed as a victory by both contestants, as an allusion to the confusing nature of characters displayed by the new genus and to the 
<typeStatus id="5E5C47BF5C2AFFDCFCED4EF76A55D197" box="[836,884,656,680]" pageId="4" pageNumber="305">type</typeStatus>
species locality: Lord Howe Island, New South 
<collectingCountry id="F9F0B98D5C2AFFDCFF3E4ED369FFD1F3" box="[151,222,692,716]" name="United Kingdom" pageId="4" pageNumber="305">Wales</collectingCountry>
, 
<collectingCountry id="F9F0B98D5C2AFFDCFF404ED36872D1F3" box="[233,339,692,716]" name="Australia" pageId="4" pageNumber="305">Australia</collectingCountry>
.
</paragraph>
<paragraph id="8158F91D5C2AFFDCFF3E4F7868A4D065" blockId="4.[151,592,799,858]" pageId="4" pageNumber="305">
<taxonomicName id="46E7829E5C2AFFDCFF3E4F786B28D006" box="[151,521,799,825]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C2AFFDCFF3E4F786B28D006" bold="true" box="[151,521,799,825]" italics="true" pageId="4" pageNumber="305">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C2AFFDCFDB94F466B71D006" bold="true" box="[528,592,801,825]" pageId="4" pageNumber="305">
<taxonomicNameLabel id="A8A098745C2AFFDCFDB94F466B71D006" box="[528,592,801,825]" pageId="4" pageNumber="305" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
<figureCitation id="19DCE5985C2AFFDCFF3E4F25680DD065" box="[151,300,834,858]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStart-2="FIGURES 11 – 14" captionStart-3="FIGURES 15 – 18" captionStart-4="FIGURES 19 – 23" captionStart-5="FIGURES 24 – 27" captionStart-6="FIGURES 28 – 30" captionStartId-0="5.[151,264,1927,1950]" captionStartId-1="6.[151,264,1919,1942]" captionStartId-2="7.[151,264,1831,1854]" captionStartId-3="8.[151,264,1755,1778]" captionStartId-4="9.[151,264,1850,1873]" captionStartId-5="10.[151,264,1780,1803]" captionStartId-6="11.[151,264,877,900]" captionTargetBox-0="[162,1430,198,1896]" captionTargetBox-1="[159,1431,195,1899]" captionTargetBox-2="[156,1436,203,1809]" captionTargetBox-3="[161,1429,211,1715]" captionTargetBox-4="[163,1430,199,1820]" captionTargetBox-5="[161,1430,198,1732]" captionTargetBox-6="[151,1436,193,855]" captionTargetId-0="figure@5.[151,1436,193,1906]" captionTargetId-1="figure@6.[151,1436,193,1906]" captionTargetId-2="figure@7.[151,1436,193,1810]" captionTargetId-3="figure@8.[151,1436,193,1734]" captionTargetId-4="figure@9.[151,1436,193,1829]" captionTargetId-5="figure@10.[151,1436,193,1733]" captionTargetId-6="figure@11.[151,1436,193,856]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="8" captionTargetPageId-4="9" captionTargetPageId-5="10" captionTargetPageId-6="11" captionText-0="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." captionText-1="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." captionText-2="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." captionText-3="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." captionText-4="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." captionText-5="FIGURES 24 – 27. Reginacharlottia lordhowensis sp. n., adult male. 24 Subcapitulum. 25 Dorsal view of gnathosoma. 26 Lateral view of palp. 27 Distal palptibia and palptarsus (arrow points to spur on palpal apotele). Scale bars 24 – 26 = 10 µm; 27 = 1 µm." captionText-6="FIGURES 28 – 30. Reginacharlottia lordhowensis sp. n., adult female. 28 Leg I, telofemur, genu, tibia, tarsus. 29 Leg II, anterolateral view of distal telofemur (t), genu (g, al 1 missing), tibia, tarsus. 30 Habitus. Scale bars 28 – 29 = 50 µm." httpUri-0="https://zenodo.org/record/248010/files/figure.png" httpUri-1="https://zenodo.org/record/248011/files/figure.png" httpUri-2="https://zenodo.org/record/248012/files/figure.png" httpUri-3="https://zenodo.org/record/248013/files/figure.png" httpUri-4="https://zenodo.org/record/248014/files/figure.png" httpUri-5="https://zenodo.org/record/248015/files/figure.png" httpUri-6="https://zenodo.org/record/248016/files/figure.png" pageId="4" pageNumber="305">Figures 1–30</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFE914F2568A0D065" box="[312,385,834,858]" captionStart="FIGURES 48 – 49" captionStartId="20.[151,264,1696,1719]" captionTargetBox="[156,1419,604,1668]" captionTargetId="figure@20.[151,1436,591,1675]" captionTargetPageId="20" captionText="FIGURES 48 – 49. Reginacharlottia lordhowensis sp. n. larva 48 Reconstruction of dorsal chaetotaxy (arrow points to posterior horn). 49 Anal region (pa = para-anal seta). Figure 50 Epicroseius sp. (Sejidae) Queensland, Australia, female genital shield with attached spermatophore. Scale bar 48 = 100 µm; 49 = 25 µm." httpUri="https://zenodo.org/record/248022/files/figure.png" pageId="4" pageNumber="305">48–49</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C2AFFDCFF3E4FED68AFD649" pageId="4" pageNumber="305" type="description">
<paragraph id="8158F91D5C2AFFDCFF3E4FED68AFD649" blockId="4.[151,1436,906,2010]" pageId="4" pageNumber="305">
<emphasis id="B393250F5C2AFFDCFF3E4FED68B9D09C" bold="true" box="[151,408,906,931]" pageId="4" pageNumber="305">Specimens examined.</emphasis>
<typeStatus id="5E5C47BF5C2AFFDCFE094FED6B29D09D" box="[416,520,906,930]" pageId="4" pageNumber="305" type="holotype">Holotype</typeStatus>
female, Lord Howe Island, New South Walter, 
<collectingCountry id="F9F0B98D5C2AFFDCFB824FED6DB4D09D" box="[1067,1173,906,930]" name="Australia" pageId="4" pageNumber="305">Australia</collectingCountry>
; ex leaf litter in closed rainforest 
<taxonomicName id="46E7829E5C2AFFDCFEA74FC968B9D0F9" box="[270,408,942,966]" class="Liliopsida" family="Cyperaceae" genus="Cleistocalyx" higherTaxonomySource="CoL" kingdom="Plantae" order="Poales" pageId="4" pageNumber="305" phylum="Tracheophyta" rank="genus">
<emphasis id="B393250F5C2AFFDCFEA74FC968B9D0F9" box="[270,408,942,966]" italics="true" pageId="4" pageNumber="305">Cleistocalyx</emphasis>
</taxonomicName>
/ 
<taxonomicName id="46E7829E5C2AFFDCFE094FC96B0ED0F9" box="[416,559,942,966]" class="Magnoliopsida" family="Oleaceae" genus="Chionanthus" kingdom="Plantae" order="Lamiales" pageId="4" pageNumber="305" phylum="Tracheophyta" rank="genus">
<emphasis id="B393250F5C2AFFDCFE094FC96B0ED0F9" box="[416,559,942,966]" italics="true" pageId="4" pageNumber="305">Chionanthus</emphasis>
</taxonomicName>
habitat (LHISO26L) on Boat Harbour walking trail 
<quantity id="461F54F85C2AFFDCFB0C4FC96DC4D0F9" box="[1189,1253,942,966]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="305" unit="m" value="400.0">400m</quantity>
before harbour (
<geoCoordinate id="E4D39FDA5C2AFFDCFF094FB5680AD0D5" box="[160,299,978,1002]" direction="south" orientation="latitude" pageId="4" pageNumber="305" precision="15" value="-31.558887">31°33' 32”S</geoCoordinate>
, 
<geoCoordinate id="E4D39FDA5C2AFFDCFE904FB568F4D0D5" box="[313,469,978,1002]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="15" value="159.09277">159°05' 34”E</geoCoordinate>
), 
<date id="F559DFDD5C2AFFDCFE424FB56BEAD0D5" box="[491,715,978,1002]" pageId="4" pageNumber="305" value="2000-11-21">21 November 2000</date>
. In Australian Museum, Sydney. 
<typeStatus id="5E5C47BF5C2AFFDCFBE14FB46D98D0D4" box="[1096,1209,979,1003]" pageId="4" pageNumber="305" type="paratype">Paratypes</typeStatus>
: 
<specimenCount id="97E132945C2AFFDCFB6C4FB56C60D0D5" box="[1221,1345,978,1002]" pageId="4" pageNumber="305" type="female">24 females</specimenCount>
(4 with larvae), 
<specimenCount id="97E132945C2AFFDCFF5D4F91687CD731" box="[244,349,1014,1038]" pageId="4" pageNumber="305" type="male">12 males</specimenCount>
, 2 deutonymphs, 1 protonymph, same data except from various litter 
<typeStatus id="5E5C47BF5C2AFFDCFBD24F906D97D730" box="[1147,1206,1015,1039]" pageId="4" pageNumber="305">types</typeStatus>
(sclerophyll, closed rainforest forest, palm) Boat Harbour, Erskine Valley, Dawson Point Ridge, Goat Home, Phillip Point, and Mount Gower, 
<date id="F559DFDD5C2AFFDCFF5B485968ECD769" box="[242,461,1086,1110]" pageId="4" pageNumber="305" value="2000-11-21">21 November 2000</date>
to 
<date id="F559DFDD5C2AFFDCFE5D48596BA0D769" box="[500,641,1086,1110]" pageId="4" pageNumber="305" value="2003-06-08">8 June 2003</date>
; 
<specimenCount id="97E132945C2AFFDCFD2648596BDDD769" box="[655,764,1086,1110]" pageId="4" pageNumber="305" type="female">2 females</specimenCount>
Intermediate Hill, Goat house walking track (
<geoCoordinate id="E4D39FDA5C2AFFDCFAA248596CB6D769" box="[1291,1431,1086,1110]" direction="south" orientation="latitude" pageId="4" pageNumber="305" precision="15" value="-31.554167">31°33' 15”S</geoCoordinate>
, 
<geoCoordinate id="E4D39FDA5C2AFFDCFF3E4805680ED745" box="[151,303,1122,1146]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="15" value="159.0825">159°04' 57”E</geoCoordinate>
) ex 
<taxonomicName id="46E7829E5C2AFFDCFEF748036B07D744" box="[350,550,1123,1147]" class="Insecta" family="Cerambycidae" genus="Howea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="fosteriana">
<emphasis id="B393250F5C2AFFDCFEF748036B07D744" box="[350,550,1123,1147]" italics="true" pageId="4" pageNumber="305">Howea fosteriana</emphasis>
</taxonomicName>
palm litter (LHI/JT/09L), 
<date id="F559DFDD5C2AFFDCFCE448056D37D745" box="[845,1046,1122,1146]" pageId="4" pageNumber="305" value="2001-02-23">23 February 2001</date>
, J. Tarnawski &amp; M. Shea; 
<specimenCount id="97E132945C2AFFDCFA9248056CBDD745" box="[1339,1436,1122,1146]" pageId="4" pageNumber="305" type="female">1 female</specimenCount>
ex moss, lichens, liverworts (LHI/JT/10L) near Goat House Cave, 
<date id="F559DFDD5C2AFFDCFCD648E16D68D7A1" box="[895,1097,1158,1182]" pageId="4" pageNumber="305" value="2001-02-23">23 February 2001</date>
; 
<specimenCount id="97E132945C2AFFDCFBFD48E16D8CD7A1" box="[1108,1197,1158,1182]" pageId="4" pageNumber="305" type="male">2 males</specimenCount>
, 1 deutonymph Little 
<collectingCountry id="F9F0B98D5C2AFFDCFF3E48CD69FAD7FD" box="[151,219,1194,1218]" name="Iceland" pageId="4" pageNumber="305">Island</collectingCountry>
behind Far Flats (
<geoCoordinate id="E4D39FDA5C2AFFDCFE1148CD6B19D7FD" box="[440,568,1194,1218]" direction="south" orientation="latitude" pageId="4" pageNumber="305" precision="15" value="-31.56889">31°34' 8”S</geoCoordinate>
, 
<geoCoordinate id="E4D39FDA5C2AFFDCFDE148CD6BC5D7FD" box="[584,740,1194,1218]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="15" value="159.08112">159°04' 52”E</geoCoordinate>
) ex 
<taxonomicName id="46E7829E5C2AFFDCFCB448CB6ACBD7FC" box="[797,1002,1195,1219]" class="Insecta" family="Cerambycidae" genus="Howea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="305" phylum="Arthropoda" rank="species" species="fosteriana">
<emphasis id="B393250F5C2AFFDCFCB448CB6ACBD7FC" box="[797,1002,1195,1219]" italics="true" pageId="4" pageNumber="305">Howea fosteriana</emphasis>
</taxonomicName>
litter (1HO21A), 
<date id="F559DFDD5C2AFFDCFB6948CD6C5FD7FD" box="[1216,1406,1194,1219]" pageId="4" pageNumber="305" value="2001-08-10">10 August 2001</date>
; 1 deutonymph 
<quantity id="461F54F85C2AFFDCFE8048A96851D7D9" box="[297,368,1230,1254]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="305" unit="m" value="100.0">100 m</quantity>
east Soldier Creek (
<geoCoordinate id="E4D39FDA5C2AFFDCFDFA48A96BFCD7D9" box="[595,733,1230,1254]" direction="south" orientation="latitude" pageId="4" pageNumber="305" precision="15" value="-31.581945">31°34' 55”S</geoCoordinate>
, 
<geoCoordinate id="E4D39FDA5C2AFFDCFD4048A96AA1D7D9" box="[745,896,1230,1254]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="15" value="159.08583">159°05' 09”E</geoCoordinate>
) ex 
<taxonomicName id="46E7829E5C2AFFDCFC1848A96D83D7D9" box="[945,1186,1230,1254]" class="Magnoliopsida" family="Myrtaceae" genus="Cleistocalyx" higherTaxonomySource="CoL" kingdom="Plantae" order="Myrtales" pageId="4" pageNumber="305" phylum="Tracheophyta" rank="species" species="fullageri">
<emphasis id="B393250F5C2AFFDCFC1848A96D83D7D9" box="[945,1186,1230,1254]" italics="true" pageId="4" pageNumber="305">Cleistocalyx fullageri</emphasis>
</taxonomicName>
litter (RATSCF7-3L), 
<date id="F559DFDD5C2AFFDCFF3E48956803D635" box="[151,290,1266,1290]" pageId="4" pageNumber="305" value="2003-06-08">8 June 2003</date>
; 1 protonymph, beside walking track, 3rd Tee (
<geoCoordinate id="E4D39FDA5C2AFFDCFC9F48956A9FD635" box="[822,958,1266,1290]" direction="south" orientation="latitude" pageId="4" pageNumber="305" precision="15" value="-31.553055">31°33' 11”S</geoCoordinate>
, 
<geoCoordinate id="E4D39FDA5C2AFFDCFC6248956D44D635" box="[971,1125,1266,1290]" direction="east" orientation="longitude" pageId="4" pageNumber="305" precision="15" value="159.0836">159°05' 01”E</geoCoordinate>
), 
<date id="F559DFDD5C2AFFDCFBD348956C33D635" box="[1146,1298,1266,1291]" pageId="4" pageNumber="305" value="2003-05-28">28 May 2003</date>
, RATGCFC 2-1, ex litter lowland mixed forest, I. Hutton, K. Lees. In Australian Museum, Sydney; Australian National Insect Collection, Canberra; Royal Alberta Museum, Alberta, 
<collectingCountry id="F9F0B98D5C2AFFDCFC8A495D6A5BD66D" box="[803,890,1338,1362]" name="Canada" pageId="4" pageNumber="305">Canada</collectingCountry>
; Acarology Laboratory, Ohio State University, Columbus, OH, 
<collectingCountry id="F9F0B98D5C2AFFDCFEE7493968ABD649" box="[334,394,1374,1398]" name="United States of America" pageId="4" pageNumber="305">USA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C2AFFDCFF6E49E56821D54D" pageId="4" pageNumber="305" type="diagnosis">
<paragraph id="8158F91D5C2AFFDCFF6E49E56A6FD515" blockId="4.[151,1436,906,2010]" pageId="4" pageNumber="305">
<emphasis id="B393250F5C2AFFDCFF6E49E5681BD6A4" bold="true" box="[199,314,1410,1435]" pageId="4" pageNumber="305">Diagnosis</emphasis>
. With the characteristics of the genus, podonotal-metanotal shield tubercles flattened, irregular, with marginal lamellae (
<figureCitation id="19DCE5985C2AFFDCFEC649C168E3D681" box="[367,450,1446,1470]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="4" pageNumber="305">Fig. 22</figureCitation>
), posterior margin of podonotal-metanotal shield eroded medially (
<figureCitation id="19DCE5985C2AFFDCFB1849C16DD2D681" box="[1201,1267,1446,1470]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="4" pageNumber="305">Fig. 1</figureCitation>
); peritrematalmetapodal shield with tuberculate ornamentation similar to podonotal-metanotal shield (
<figureCitation id="19DCE5985C2AFFDCFB2B49AD6DEDD6DD" box="[1154,1228,1482,1506]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="4" pageNumber="305">Figs 2</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFB7349AD6DD9D6DD" box="[1242,1272,1482,1506]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="4" pageNumber="305">15</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFAAC49AD6C04D6DD" box="[1285,1317,1482,1506]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="4" pageNumber="305">20</figureCitation>
); movable cheliceral digit ending in three minute teeth; soft cuticle with plicate-reticulate ornamentation (
<figureCitation id="19DCE5985C2AFFDCFB6A49896C16D539" box="[1219,1335,1518,1542]" captionStart-0="FIGURES 5 – 10" captionStart-1="FIGURES 11 – 14" captionStartId-0="6.[151,264,1919,1942]" captionStartId-1="7.[151,264,1831,1854]" captionTargetBox-0="[159,1431,195,1899]" captionTargetBox-1="[156,1436,203,1809]" captionTargetId-0="figure@6.[151,1436,193,1906]" captionTargetId-1="figure@7.[151,1436,193,1810]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." captionText-1="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri-0="https://zenodo.org/record/248011/files/figure.png" httpUri-1="https://zenodo.org/record/248012/files/figure.png" pageId="4" pageNumber="305">Figs 9–11</figureCitation>
). Genual setation I–IV, 
<date id="F559DFDD5C2AFFDCFE9D4A756895D515" box="[308,436,1554,1578]" pageId="4" pageNumber="305" value="2012-09-12" valueMax="2012-09-12" valueMin="2012-09-12">9-12-12-12</date>
, respectively; tibia IV with 11 setae.
</paragraph>
<paragraph id="8158F91D5C2AFFDCFF6E4A506821D54D" blockId="4.[151,1436,906,2010]" pageId="4" pageNumber="305">
<emphasis id="B393250F5C2AFFDCFF6E4A50683FD570" bold="true" box="[199,286,1591,1615]" pageId="4" pageNumber="305">Female</emphasis>
(n = 24). Body (
<figureCitation id="19DCE5985C2AFFDCFE4D4A516B6AD571" box="[484,587,1590,1614]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="4" pageNumber="305">Figs 1–2</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFDF04A516B57D571" box="[601,630,1590,1614]" captionStart="FIGURES 28 – 30" captionStartId="11.[151,264,877,900]" captionTargetBox="[151,1436,193,855]" captionTargetId="figure@11.[151,1436,193,856]" captionTargetPageId="11" captionText="FIGURES 28 – 30. Reginacharlottia lordhowensis sp. n., adult female. 28 Leg I, telofemur, genu, tibia, tarsus. 29 Leg II, anterolateral view of distal telofemur (t), genu (g, al 1 missing), tibia, tarsus. 30 Habitus. Scale bars 28 – 29 = 50 µm." httpUri="https://zenodo.org/record/248016/files/figure.png" pageId="4" pageNumber="305">30</figureCitation>
) sack-like 875–1200 µm long, 770–1100 wide (body expands as eggs develop).
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C2AFFD4FF6E4A196B7ED630" lastPageId="12" lastPageNumber="313" pageId="4" pageNumber="305" type="description">
<paragraph id="8158F91D5C2AFFDCFF6E4A196DB5D451" blockId="4.[151,1436,906,2010]" pageId="4" pageNumber="305">
<emphasis id="B393250F5C2AFFDCFF6E4A196839D5A9" box="[199,280,1662,1686]" italics="true" pageId="4" pageNumber="305">Cuticle</emphasis>
. Most sclerotised shields (podonotal-metanotal, peritrematal, ventral, pygidial) ornamented with flattened tubercles with irregular marginal lamellae interspersed with minute (1–3) denticles and surrounding raised muscle insertions (
<figureCitation id="19DCE5985C2AFFDCFE124AA16B03D5E1" box="[443,546,1734,1758]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="4" pageNumber="305">Figs 1–2</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFD984AA16B5DD5E1" box="[561,636,1734,1758]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="4" pageNumber="305">22–23</figureCitation>
); ventri-anal shield (
<figureCitation id="19DCE5985C2AFFDCFCC14AA16A95D5E1" box="[872,948,1734,1758]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="4" pageNumber="305">Figs 6</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFC6B4AA16AFCD5E1" box="[962,989,1734,1758]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="4" pageNumber="305">11</figureCitation>
) formed from scale-like platelets; legs with scale-like reticula with smooth or toothed margins (
<figureCitation id="19DCE5985C2AFFDCFCB84A8D6A42D43D" box="[785,867,1770,1794]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="4" pageNumber="305">Fig. 13</figureCitation>
). Soft cuticle plicate, ridges smooth to denticulate, larger denticles forming reticulate pattern (
<figureCitation id="19DCE5985C2AFFDCFD2A4B696BEFD419" box="[643,718,1806,1830]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="4" pageNumber="305">Figs 3</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFD744B696BD9D419" box="[733,760,1806,1830]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="4" pageNumber="305">11</figureCitation>
); numerous subdermal pores, hypertrichous. Setae (mostly 24–37 long, up to 20 wide) with ventral rachis and dorsal array of imbricate scale-like ornamentation resembling conifer-cones (
<figureCitation id="19DCE5985C2AFFDCFE944B3168B2D451" box="[317,403,1878,1902]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="4" pageNumber="305">Figs 11</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFE364B31689CD451" box="[415,445,1878,1902]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="4" pageNumber="305">17</figureCitation>
), each inserted on small, raised, ridged platelet (see 
<figureCitation id="19DCE5985C2AFFDCFBAE4B316D7DD451" box="[1031,1116,1878,1902]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="4" pageNumber="305">Figs 17</figureCitation>
, 
<figureCitation id="19DCE5985C2AFFDCFBC14B316DA9D451" box="[1128,1160,1878,1902]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="4" pageNumber="305">23</figureCitation>
).
</paragraph>
<paragraph id="8158F91D5C2AFFD3FF6E4B1C6C65D780" blockId="4.[151,1436,906,2010]" lastBlockId="11.[151,1436,975,2008]" lastPageId="11" lastPageNumber="312" pageId="4" pageNumber="305">
<emphasis id="B393250F5C2AFFDCFF6E4B1C6877D4AC" box="[199,342,1915,1939]" italics="true" pageId="4" pageNumber="305">Gnathosoma</emphasis>
. Chelicerae (
<figureCitation id="19DCE5985C2AFFDCFE424B1D6B1DD4AD" box="[491,572,1914,1938]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="4" pageNumber="305">Fig. 14</figureCitation>
) short (73 long including regressed fixed digit ~20–25), pilus dentilis minute setiform, cheliceral seta (~23 long) membranous, expanded and covering fixed digit; Movable digit 32–35 long, smooth, tapering to 3 small distal teeth. Feathery interdigital excrescence (37–45 long) apparently issuing from soft cuticle at juncture of digits and extending well beyond chelicera. Anterior hypostomatal seta 
<emphasis id="B393250F5C25FFD3FB764FA86DDAD0D8" box="[1247,1275,975,999]" italics="true" pageId="11" pageNumber="312">h1</emphasis>
(18–20 long) flattened, membranous (
<figureCitation id="19DCE5985C25FFD3FE044F946B36D734" box="[429,535,1011,1035]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStartId-0="5.[151,264,1927,1950]" captionStartId-1="6.[151,264,1919,1942]" captionTargetBox-0="[162,1430,198,1896]" captionTargetBox-1="[159,1431,195,1899]" captionTargetId-0="figure@5.[151,1436,193,1906]" captionTargetId-1="figure@6.[151,1436,193,1906]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." captionText-1="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri-0="https://zenodo.org/record/248010/files/figure.png" httpUri-1="https://zenodo.org/record/248011/files/figure.png" pageId="11" pageNumber="312">Figs 4–5</figureCitation>
); outer hypostomatal seta 
<emphasis id="B393250F5C25FFD3FCE14F946A45D734" box="[840,868,1011,1035]" italics="true" pageId="11" pageNumber="312">h2</emphasis>
(35–38) with 8–11 short branches distally; inner hypostomatal seta 
<emphasis id="B393250F5C25FFD3FED0487068B4D710" box="[377,405,1047,1071]" italics="true" pageId="11" pageNumber="312">h3</emphasis>
(43–50) flattened, whip-like; corniculus (13–15) setiform, curved, trifurcate distally; palpcoxal seta (17–25) simple. Deutosternal groove narrow, obsolescent, with six pairs of small teeth, slightly widened ventrally; three pairs semi-lunar arrays of denticles posteriad palpcoxal setae. Ve n tr al se ta 
<date id="F559DFDD5C25FFD3FB4348386C24D748" box="[1258,1285,1119,1143]" pageId="11" pageNumber="312">
<emphasis id="B393250F5C25FFD3FB4348386C24D748" box="[1258,1285,1119,1143]" italics="true" pageId="11" pageNumber="312">v1</emphasis>
</date>
(
<figureCitation id="19DCE5985C25FFD3FABB48386C57D748" box="[1298,1398,1119,1143]" captionStart-0="FIGURES 1 – 4" captionStart-1="FIGURES 5 – 10" captionStartId-0="5.[151,264,1927,1950]" captionStartId-1="6.[151,264,1919,1942]" captionTargetBox-0="[162,1430,198,1896]" captionTargetBox-1="[159,1431,195,1899]" captionTargetId-0="figure@5.[151,1436,193,1906]" captionTargetId-1="figure@6.[151,1436,193,1906]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." captionText-1="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri-0="https://zenodo.org/record/248010/files/figure.png" httpUri-1="https://zenodo.org/record/248011/files/figure.png" pageId="11" pageNumber="312">Figs 4–5</figureCitation>
) of palptrochanter (23–27) rake-like with 5–6 ventral tines. Palptarsal apophysis 2-tined and with short, dorsal spur (
<figureCitation id="19DCE5985C25FFD3FF3648C069CED780" box="[159,239,1191,1215]" captionStart="FIGURES 24 – 27" captionStartId="10.[151,264,1780,1803]" captionTargetBox="[161,1430,198,1732]" captionTargetId="figure@10.[151,1436,193,1733]" captionTargetPageId="10" captionText="FIGURES 24 – 27. Reginacharlottia lordhowensis sp. n., adult male. 24 Subcapitulum. 25 Dorsal view of gnathosoma. 26 Lateral view of palp. 27 Distal palptibia and palptarsus (arrow points to spur on palpal apotele). Scale bars 24 – 26 = 10 µm; 27 = 1 µm." httpUri="https://zenodo.org/record/248015/files/figure.png" pageId="11" pageNumber="312">Fig. 27</figureCitation>
arrow). Setation of palpal trochanter-femur-genu-tibia-tarsus: 
<date id="F559DFDD5C25FFD3FC0448C06D2ED780" box="[941,1039,1191,1215]" pageId="11" pageNumber="312" value="1914-05-02" valueMax="1914-06-02" valueMin="1914-05-02">2-5-6-14</date>
-15 (
<quantity id="461F54F85C25FFD3FBEA48C06D5CD780" box="[1091,1149,1191,1215]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="11" pageNumber="312" unit="in" value="10.0">10 in</quantity>
terminal cluster).
</paragraph>
<caption id="D598A9955C2BFFDDFF3E4BE06C11D4E4" httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="5" pageNumber="306" targetBox="[162,1430,198,1896]" targetPageId="5">
<paragraph id="8158F91D5C2BFFDDFF3E4BE06C11D4E4" blockId="5.[151,1437,1927,2011]" pageId="5" pageNumber="306">
<emphasis id="B393250F5C2BFFDDFF3E4BE06801D4A2" bold="true" box="[151,288,1927,1950]" pageId="5" pageNumber="306">FIGURES 1</emphasis>
– 
<emphasis id="B393250F5C2BFFDDFE854BEF681ED4A2" bold="true" box="[300,319,1928,1949]" pageId="5" pageNumber="306">4.</emphasis>
<taxonomicName id="46E7829E5C2BFFDDFEE24BEF6BA9D4A1" box="[331,648,1928,1950]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="306" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C2BFFDDFEE24BEF6BA9D4A1" box="[331,648,1928,1950]" italics="true" pageId="5" pageNumber="306">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A8A098745C2BFFDDFD3D4BEF6BF3D4A2" box="[660,722,1928,1949]" pageId="5" pageNumber="306" rank="species">
<emphasis id="B393250F5C2BFFDDFD3D4BEF6BF3D4A2" bold="true" box="[660,722,1928,1949]" pageId="5" pageNumber="306">sp. n.</emphasis>
</taxonomicNameLabel>
, adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta 
<emphasis id="B393250F5C2BFFDDFCC74BA26AA6D4E4" box="[878,903,1989,2011]" italics="true" pageId="5" pageNumber="306">h1</emphasis>
). Scale bars 1–2 = 100 µm; 3–4 = 10 µm.
</paragraph>
</caption>
<caption id="D598A9955C28FFDEFF3E4B186843D4CD" httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="6" pageNumber="307" targetBox="[159,1431,195,1899]" targetPageId="6">
<paragraph id="8158F91D5C28FFDEFF3E4B186843D4CD" blockId="6.[151,1437,1919,2034]" pageId="6" pageNumber="307">
<emphasis id="B393250F5C28FFDEFF3E4B18683FD4AA" bold="true" box="[151,286,1919,1942]" pageId="6" pageNumber="307">FIGURES 5</emphasis>
– 
<emphasis id="B393250F5C28FFDEFE854BE7686AD4AA" bold="true" box="[300,331,1920,1941]" pageId="6" pageNumber="307">10.</emphasis>
<taxonomicName id="46E7829E5C28FFDEFEFF4BE76BB3D4A9" box="[342,658,1920,1942]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="307" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C28FFDEFEFF4BE76BB3D4A9" box="[342,658,1920,1942]" italics="true" pageId="6" pageNumber="307">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C28FFDEFD344BE76BFBD4AA" bold="true" box="[669,730,1920,1941]" pageId="6" pageNumber="307">
<taxonomicNameLabel id="A8A098745C28FFDEFD344BE76BFBD4AA" box="[669,730,1920,1941]" pageId="6" pageNumber="307" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (
<emphasis id="B393250F5C28FFDEFDAE4BF86B01D48A" box="[519,544,1951,1973]" italics="true" pageId="6" pageNumber="307">h1</emphasis>
– 
<emphasis id="B393250F5C28FFDEFD844BF86B18D48A" box="[557,569,1951,1973]" italics="true" pageId="6" pageNumber="307">3</emphasis>
), ventral palptrochanteral seta (
<emphasis id="B393250F5C28FFDEFC3E4BF86A8ED48A" box="[919,943,1951,1973]" italics="true" pageId="6" pageNumber="307">v1</emphasis>
). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9–10 Variation in pygidial shield. Scale bars = 10 µm.
</paragraph>
</caption>
<caption id="D598A9955C29FFDFFF3E4B406A6ED445" httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="7" pageNumber="308" targetBox="[156,1436,203,1809]" targetPageId="7">
<paragraph id="8158F91D5C29FFDFFF3E4B406A6ED445" blockId="7.[151,1436,1831,1914]" pageId="7" pageNumber="308">
<emphasis id="B393250F5C29FFDFFF3E4B406809D403" bold="true" box="[151,296,1831,1854]" pageId="7" pageNumber="308">FIGURES 11</emphasis>
– 
<emphasis id="B393250F5C29FFDFFE9C4B40686FD403" bold="true" box="[309,334,1831,1852]" pageId="7" pageNumber="308">14</emphasis>
. 
<taxonomicName id="46E7829E5C29FFDFFEF44B406BB9D402" box="[349,664,1831,1853]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="308" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C29FFDFFEF44B406BB9D402" box="[349,664,1831,1853]" italics="true" pageId="7" pageNumber="308">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C29FFDFFD084B4F6BFCD402" bold="true" box="[673,733,1832,1853]" pageId="7" pageNumber="308">
<taxonomicNameLabel id="A8A098745C29FFDFFD084B4F6BFCD402" box="[673,733,1832,1853]" pageId="7" pageNumber="308" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm.
</paragraph>
</caption>
<caption id="D598A9955C26FFD0FF3E4ABC6B9AD411" httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="8" pageNumber="309" targetBox="[161,1429,211,1715]" targetPageId="8">
<paragraph id="8158F91D5C26FFD0FF3E4ABC6B9AD411" blockId="8.[151,1437,1755,1838]" pageId="8" pageNumber="309">
<emphasis id="B393250F5C26FFD0FF3E4ABC6808D5CF" bold="true" box="[151,297,1755,1778]" pageId="8" pageNumber="309">FIGURES 15</emphasis>
– 
<emphasis id="B393250F5C26FFD0FE9F4ABC6874D5CF" bold="true" box="[310,341,1755,1776]" pageId="8" pageNumber="309">18.</emphasis>
<taxonomicName id="46E7829E5C26FFD0FEF74ABC6BB9D5CE" box="[350,664,1755,1777]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C26FFD0FEF74ABC6BB9D5CE" box="[350,664,1755,1777]" italics="true" pageId="8" pageNumber="309">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C26FFD0FD094ABB6BFAD5CE" bold="true" box="[672,731,1756,1777]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A8A098745C26FFD0FD094ABB6BFAD5CE" box="[672,731,1756,1777]" pageId="8" pageNumber="309" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. 
<taxonomicName id="46E7829E5C26FFD0FD0C4A9D6AFED42F" box="[677,991,1786,1808]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="309" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C26FFD0FD0C4A9D6AFED42F" box="[677,991,1786,1808]" italics="true" pageId="8" pageNumber="309">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C26FFD0FC4E4A9D6D03D430" bold="true" box="[999,1058,1786,1807]" pageId="8" pageNumber="309">
<taxonomicNameLabel id="A8A098745C26FFD0FC4E4A9D6D03D430" box="[999,1058,1786,1807]" pageId="8" pageNumber="309" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm.
</paragraph>
</caption>
<caption id="D598A9955C27FFD1FF3E4B5D6CBDD4B1" httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="9" pageNumber="310" targetBox="[163,1430,199,1820]" targetPageId="9">
<paragraph id="8158F91D5C27FFD1FF3E4B5D6CBDD4B1" blockId="9.[151,1436,1850,1935]" pageId="9" pageNumber="310">
<emphasis id="B393250F5C27FFD1FF3E4B5D6808D46F" bold="true" box="[151,297,1850,1873]" pageId="9" pageNumber="310">FIGURES 19</emphasis>
– 
<emphasis id="B393250F5C27FFD1FE9F4B5C6877D46F" bold="true" box="[310,342,1851,1872]" pageId="9" pageNumber="310">23.</emphasis>
<taxonomicName id="46E7829E5C27FFD1FEF74B5C6BB8D46E" box="[350,665,1851,1873]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="310" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C27FFD1FEF74B5C6BB8D46E" box="[350,665,1851,1873]" italics="true" pageId="9" pageNumber="310">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C27FFD1FD084B5C6BFCD46F" bold="true" box="[673,733,1851,1872]" pageId="9" pageNumber="310">
<taxonomicNameLabel id="A8A098745C27FFD1FD084B5C6BFCD46F" box="[673,733,1851,1872]" pageId="9" pageNumber="310" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19–20 = 100 µm; 21–23 = 10 µm.
</paragraph>
</caption>
<caption id="D598A9955C24FFD2FF3E4A9369C9D477" httpUri="https://zenodo.org/record/248015/files/figure.png" pageId="10" pageNumber="311" targetBox="[161,1430,198,1732]" targetPageId="10">
<paragraph id="8158F91D5C24FFD2FF3E4A9369C9D477" blockId="10.[151,1436,1780,1864]" pageId="10" pageNumber="311">
<emphasis id="B393250F5C24FFD2FF3E4A93680BD435" bold="true" box="[151,298,1780,1803]" pageId="10" pageNumber="311">FIGURES 24</emphasis>
– 
<emphasis id="B393250F5C24FFD2FE9E4A926877D435" bold="true" box="[311,342,1781,1802]" pageId="10" pageNumber="311">27.</emphasis>
<taxonomicName id="46E7829E5C24FFD2FEC94A926BBDD434" box="[352,668,1781,1803]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="311" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C24FFD2FEC94A926BBDD434" box="[352,668,1781,1803]" italics="true" pageId="10" pageNumber="311">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C24FFD2FD0C4A926BC0D435" bold="true" box="[677,737,1781,1802]" pageId="10" pageNumber="311">
<taxonomicNameLabel id="A8A098745C24FFD2FD0C4A926BC0D435" box="[677,737,1781,1802]" pageId="10" pageNumber="311" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult male. 24 Subcapitulum. 25 Dorsal view of gnathosoma. 26 Lateral view of palp. 27 Distal palptibia and palptarsus (arrow points to spur on palpal apotele). Scale bars 24–26 = 10 µm; 27 = 1 µm.
</paragraph>
</caption>
<caption id="D598A9955C25FFD3FF3E4F0A6C0BD09D" httpUri="https://zenodo.org/record/248016/files/figure.png" pageId="11" pageNumber="312" targetBox="[151,1436,193,855]" targetPageId="11">
<paragraph id="8158F91D5C25FFD3FF3E4F0A6C0BD09D" blockId="11.[151,1436,877,930]" pageId="11" pageNumber="312">
<emphasis id="B393250F5C25FFD3FF3E4F0A680BD0BC" bold="true" box="[151,298,877,900]" pageId="11" pageNumber="312">FIGURES 28</emphasis>
– 
<emphasis id="B393250F5C25FFD3FE9E4F096876D0BC" bold="true" box="[311,343,878,899]" pageId="11" pageNumber="312">30.</emphasis>
<taxonomicName id="46E7829E5C25FFD3FEC84F096BBCD0BB" box="[353,669,878,900]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="312" phylum="Arthropoda" rank="species" species="lordhowensis" status="sp. nov.">
<emphasis id="B393250F5C25FFD3FEC84F096BBCD0BB" box="[353,669,878,900]" italics="true" pageId="11" pageNumber="312">Reginacharlottia lordhowensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C25FFD3FD0E4F096BC2D0BC" bold="true" box="[679,739,878,899]" pageId="11" pageNumber="312">
<taxonomicNameLabel id="A8A098745C25FFD3FD0E4F096BC2D0BC" box="[679,739,878,899]" pageId="11" pageNumber="312" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult female. 28 Leg I, telofemur, genu, tibia, tarsus. 29 Leg II, anterolateral view of distal telofemur (t), genu (g, 
<emphasis id="B393250F5C25FFD3FD3B4FEB6B92D09D" box="[658,691,908,930]" italics="true" pageId="11" pageNumber="312">al1</emphasis>
missing), tibia, tarsus. 30 Habitus. Scale bars 28–29 = 50 µm.
</paragraph>
</caption>
<paragraph id="8158F91D5C25FFD3FF6E48AB6BD1D6E0" blockId="11.[151,1436,975,2008]" pageId="11" pageNumber="312">
<emphasis id="B393250F5C25FFD3FF6E48AB6800D7DC" box="[199,289,1228,1251]" italics="true" pageId="11" pageNumber="312">Dorsum</emphasis>
. Prodorsal-metanotal and peritrematal shields (
<figureCitation id="19DCE5985C25FFD3FCEA48AC6A8DD7DC" box="[835,940,1227,1251]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="11" pageNumber="312">Figs 1–2</figureCitation>
, 
<figureCitation id="19DCE5985C25FFD3FC1248AC6AF9D7DC" box="[955,984,1227,1251]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="11" pageNumber="312">15</figureCitation>
) ornamented with large (mostly 10–15 long) flattened tubercles, angular to rounded in shape, and large, raised, finely punctate muscle insertions; vertex subrectangular with two pairs of setae on anterior margin, projecting over gnathosoma; prodorsal-metanotal shield bilobed posteriorly (lobes 500–550 long) with deep (100–150) medial incision; peritrematal shield very broad (to~ 100), extending posteriorly around coxa IV to capture metapodal plate; peritreme sinuate, extending along medial margin of shield from posterior corner of vertex to stigma at level of coxa IV, with post-stigmatal gutter extended posterior to coxa IV. Pygidial shield (
<figureCitation id="19DCE5985C25FFD3FDEF49C46B9AD684" box="[582,699,1443,1467]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="11" pageNumber="312">Figs 9–10</figureCitation>
) minute (18–37 long 
<date id="F559DFDD5C25FFD3FC1E49C36D35D684" box="[951,1044,1443,1467]" pageId="11" pageNumber="312" value="1945-10-21">x 21–45</date>
wide), without setae, with central tubercle divided into 
<date id="F559DFDD5C25FFD3FE2C49A06B3CD6E0" box="[389,541,1479,1503]" pageId="11" pageNumber="312" value="2012-03-02" valueMax="2012-04-02" valueMin="2012-03-02">2–3 and 4–12</date>
satellite tubercles.
</paragraph>
<paragraph id="8158F91D5C25FFD3FF6E498B6AF9D41C" blockId="11.[151,1436,975,2008]" pageId="11" pageNumber="312">
<emphasis id="B393250F5C25FFD3FF6E498B682DD53C" box="[199,268,1516,1539]" italics="true" pageId="11" pageNumber="312">Venter</emphasis>
. Sternal region (
<figureCitation id="19DCE5985C25FFD3FE62498C6B34D53C" box="[459,533,1515,1539]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="11" pageNumber="312">Figs 7</figureCitation>
, 
<figureCitation id="19DCE5985C25FFD3FD8B498C6B1ED53C" box="[546,575,1515,1539]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="11" pageNumber="312">16</figureCitation>
) with reticulate pattern, lightly sclerotised, reticulate and bearing 
<emphasis id="B393250F5C25FFD3FA87498C6C6ED53C" box="[1326,1359,1515,1539]" italics="true" pageId="11" pageNumber="312">st1</emphasis>
– 
<emphasis id="B393250F5C25FFD3FAF4498C6C4AD53C" box="[1373,1387,1515,1539]" italics="true" pageId="11" pageNumber="312">3</emphasis>
; 
<emphasis id="B393250F5C25FFD3FAD2498C6CBDD53C" box="[1403,1436,1515,1539]" italics="true" pageId="11" pageNumber="312">st1</emphasis>
thick, covered in barbs on ventral surface and distally; 
<emphasis id="B393250F5C25FFD3FD554A686A3CD518" box="[764,797,1551,1575]" italics="true" pageId="11" pageNumber="312">st2</emphasis>
(41) barbed, tapering; 
<emphasis id="B393250F5C25FFD3FBB74A686D1ED518" box="[1054,1087,1551,1575]" italics="true" pageId="11" pageNumber="312">st3</emphasis>
simple to lightly barbed; 
<emphasis id="B393250F5C25FFD3FAF64A686C5ED518" box="[1375,1407,1551,1575]" italics="true" pageId="11" pageNumber="312">st4</emphasis>
– 
<emphasis id="B393250F5C25FFD3FA274A776CBDD518" box="[1422,1436,1552,1575]" italics="true" pageId="11" pageNumber="312">7</emphasis>
simple, in soft cuticle; rarely spurious seta on small platelet carrying pore (stp3?). Genital shield (
<figureCitation id="19DCE5985C25FFD3FB744A546C3ED574" box="[1245,1311,1587,1611]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="11" pageNumber="312">Fig. 3</figureCitation>
) (100 long 
<date id="F559DFDD5C25FFD3FF3E4A3F69E9D550" box="[151,200,1623,1647]" pageId="11" pageNumber="312" value="1943-10">x 43</date>
wide) without setae or rarely with 
<emphasis id="B393250F5C25FFD3FDF84A3F6B53D550" box="[593,626,1624,1647]" italics="true" pageId="11" pageNumber="312">st7</emphasis>
captured asymmetrically, ornamented with small denticles and irregular tubercles, smooth area on anterior margin of shield for attachment of spermatophore, two pairs lyrifissures in soft cuticle. Anal opening (
<figureCitation id="19DCE5985C25FFD3FE0A4AF868D1D588" box="[419,496,1695,1719]" captionStart="FIGURES 1 – 4" captionStartId="5.[151,264,1927,1950]" captionTargetBox="[162,1430,198,1896]" captionTargetId="figure@5.[151,1436,193,1906]" captionTargetPageId="5" captionText="FIGURES 1 – 4. Reginacharlottia lordhowensis sp. n., adult female (scanning electron micrographs of previously slided specimen). 1 Dorsal habitus (arrow points to mesonotal element of podonotal-mesosnotal shield). 2 Ventral habitus. 3 Genital region. 4 Subcapitulum and left palp (arrow points to membranous seta h 1). Scale bars 1 – 2 = 100 µm; 3 – 4 = 10 µm." httpUri="https://zenodo.org/record/248010/files/figure.png" pageId="11" pageNumber="312">Figs 2</figureCitation>
, 
<figureCitation id="19DCE5985C25FFD3FDA84AF86B30D588" box="[513,529,1695,1719]" captionStart="FIGURES 5 – 10" captionStartId="6.[151,264,1919,1942]" captionTargetBox="[159,1431,195,1899]" captionTargetId="figure@6.[151,1436,193,1906]" captionTargetPageId="6" captionText="FIGURES 5 – 10. Reginacharlottia lordhowensis sp. n., adult female. 5 Lateral view distal hypostome, chelicerae, palps: corniculus (c), hypostomal setae (h 1 – 3), ventral palptrochanteral seta (v 1). 6 Ventri-anal shield surrounded by plicatedenticulate cuticle. 7 Sternogenital region (arrow points to spurious seta). 8 Endogynium. 9 – 10 Variation in pygidial shield. Scale bars = 10 µm." httpUri="https://zenodo.org/record/248011/files/figure.png" pageId="11" pageNumber="312">6</figureCitation>
, 
<figureCitation id="19DCE5985C25FFD3FD884AF86B1DD588" box="[545,572,1695,1719]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="11" pageNumber="312">11</figureCitation>
) in irregular conical shield (87 long x 100 wide) composed of scale-like platelets and outer array of 20 or more small platelets each bearing a smooth, tapering seta inserted on raised, ridged projection; inner region of scale-like platelets with anterior pair of flattened, tapering setae (90–92 long) projecting over opening, covered by pair of small (~25 long), nude valves.
</paragraph>
<paragraph id="8158F91D5C25FFD4FF6E4B576803D3EB" blockId="11.[151,1436,975,2008]" lastBlockId="12.[151,1437,151,2016]" lastPageId="12" lastPageNumber="313" pageId="11" pageNumber="312">
<emphasis id="B393250F5C25FFD3FF6E4B5769DDD478" box="[199,252,1840,1863]" italics="true" pageId="11" pageNumber="312">Legs</emphasis>
. (
<figureCitation id="19DCE5985C25FFD3FEBD4B48684FD478" box="[276,366,1839,1863]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="11" pageNumber="312">Figs 13</figureCitation>
, 
<figureCitation id="19DCE5985C25FFD3FED44B4868E5D478" box="[381,452,1839,1863]" captionStart="FIGURES 28 – 30" captionStartId="11.[151,264,877,900]" captionTargetBox="[151,1436,193,855]" captionTargetId="figure@11.[151,1436,193,856]" captionTargetPageId="11" captionText="FIGURES 28 – 30. Reginacharlottia lordhowensis sp. n., adult female. 28 Leg I, telofemur, genu, tibia, tarsus. 29 Leg II, anterolateral view of distal telofemur (t), genu (g, al 1 missing), tibia, tarsus. 30 Habitus. Scale bars 28 – 29 = 50 µm." httpUri="https://zenodo.org/record/248016/files/figure.png" pageId="11" pageNumber="312">28–30</figureCitation>
) Legs II–IV laterally flattened, strongly ornamented (
<figureCitation id="19DCE5985C25FFD3FB9D4B486DA8D478" box="[1076,1161,1839,1863]" captionStart="FIGURES 11 – 14" captionStartId="7.[151,264,1831,1854]" captionTargetBox="[156,1436,203,1809]" captionTargetId="figure@7.[151,1436,193,1810]" captionTargetPageId="7" captionText="FIGURES 11 – 14. Reginacharlottia lordhowensis sp. n., adult female. 11 Anal region showing ventri-anal shield, cuticular ornamentation, setae. 12 Tarsus I (arrow). 13 Anterolateral aspect of trochanter, femur and genu of leg III showing scale-like ornamentation. 14 Chelicera (light micrograph). Scale bars = 10 µm." httpUri="https://zenodo.org/record/248012/files/figure.png" pageId="11" pageNumber="312">Fig. 13</figureCitation>
); leg I (570) about half length of body, slender, antenniform, without apotele or acrotarsus (
<figureCitation id="19DCE5985C25FFD3FC234B346AFFD454" box="[906,990,1875,1899]" captionStart="FIGURES 28 – 30" captionStartId="11.[151,264,877,900]" captionTargetBox="[151,1436,193,855]" captionTargetId="figure@11.[151,1436,193,856]" captionTargetPageId="11" captionText="FIGURES 28 – 30. Reginacharlottia lordhowensis sp. n., adult female. 28 Leg I, telofemur, genu, tibia, tarsus. 29 Leg II, anterolateral view of distal telofemur (t), genu (g, al 1 missing), tibia, tarsus. 30 Habitus. Scale bars 28 – 29 = 50 µm." httpUri="https://zenodo.org/record/248016/files/figure.png" pageId="11" pageNumber="312">Fig. 28</figureCitation>
). Setation of tarsi: basitarsus II–IV each with four setae; telotarsi II–III with 14 setae, telotarsus IV with 14 plus 
<emphasis id="B393250F5C25FFD3FC124B106AC5D4B0" box="[955,996,1911,1935]" italics="true" pageId="11" pageNumber="312">av4</emphasis>
, 
<emphasis id="B393250F5C25FFD3FC5B4B106D3BD4B0" box="[1010,1050,1911,1935]" italics="true" pageId="11" pageNumber="312">pv4</emphasis>
on intercalary sclerite. Setation of legs I–IV, respectively, coxae: 2-3-2-1 (coxa II with small [9-10] dorsal seta divided distally); trochanters: 6-5-5-5; femora 
<date id="F559DFDD5C25FFD3FF474BD86870D4E8" box="[238,337,1983,2007]" pageId="11" pageNumber="312" value="2011-07-09" valueMax="2011-08-09" valueMin="2011-07-09">11-9-7-8</date>
; genua 
<date id="F559DFDD5C25FFD3FE014BD86B08D4E8" box="[424,553,1983,2007]" pageId="11" pageNumber="312" value="2012-09-12" valueMax="2012-09-12" valueMin="2012-09-12">9-12-12-12</date>
; tibiae: 8 (larval)- 
<date id="F559DFDD5C25FFD3FD534BD86A7CD4E8" box="[762,861,1983,2007]" pageId="11" pageNumber="312" value="2010-11-10">10-10-11</date>
; femora I–IV: 2-5/2-2; 2-4/2-1; 1-4/1-1; 2-(2+3)-1; genua I–IV: 2-2/1, 3/0-1; 2-3/1, 3/1-2; 2-3/1, 3/1-2; 2-3/1, 3/1-2; tibiae I-IV: 2-2/1, 2/0-1; 2-2/1, 2/1-2; 2-2/1, 2/1-2; 2-2/2, 2/1-2.
</paragraph>
<paragraph id="8158F91D5C22FFD4FF6E4C876821D128" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="313">
<emphasis id="B393250F5C22FFD4FF6E4C876825D3C7" bold="true" box="[199,260,224,248]" pageId="12" pageNumber="313">Male</emphasis>
(n = 12). Similar to female except dorsum covered by a single, large (640 long, 460 wide) shield, transverse fold present just anterior to the metapodal muscle insertions at level coxae IV (
<figureCitation id="19DCE5985C22FFD4FB274D636DC3D223" box="[1166,1250,260,284]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="12" pageNumber="313">Fig. 19</figureCitation>
). Sternitogenital shield (
<figureCitation id="19DCE5985C22FFD4FF474D406862D200" box="[238,323,295,319]" captionStart="FIGURES 15 – 18" captionStartId="8.[151,264,1755,1778]" captionTargetBox="[161,1429,211,1715]" captionTargetId="figure@8.[151,1436,193,1734]" captionTargetPageId="8" captionText="FIGURES 15 – 18. Reginacharlottia lordhowensis sp. n. Adult female. 15 Venter, legs and gnathosoma removed. 16 Sternal and genital region. 17 Prodorsal seta, lateral view. Reginacharlottia lordhowensis sp. n., adult male 18 Sternogenital region. Scale bars 15 = 500 µm; 17 = 10 µm; 16, 18 = 50 µm." httpUri="https://zenodo.org/record/248013/files/figure.png" pageId="12" pageNumber="313">Fig. 18</figureCitation>
) (152 long, 135 wide at endopodals II) reticulate to posterior margin of genital opening, then tuberculate; bearing seven pairs of setae, two pairs of lyrifissures (stp1, stp3; stp2 not present or obscure), genital opening covered by a pair of transverse, nude valves between coxae III; median gland opening just posterior to level of 
<emphasis id="B393250F5C22FFD4FF5A4DF36832D293" box="[243,275,404,428]" italics="true" pageId="12" pageNumber="313">st4</emphasis>
– 
<emphasis id="B393250F5C22FFD4FE884DF2680ED293" box="[289,303,405,428]" italics="true" pageId="12" pageNumber="313">5</emphasis>
pair; 
<emphasis id="B393250F5C22FFD4FED84DF368B3D293" box="[369,402,404,428]" italics="true" pageId="12" pageNumber="313">st1</emphasis>
(33) thick and coarsely barbed, 
<emphasis id="B393250F5C22FFD4FD554DF36A3DD293" box="[764,796,404,428]" italics="true" pageId="12" pageNumber="313">st2</emphasis>
– 
<emphasis id="B393250F5C22FFD4FC824DF36A18D293" box="[811,825,404,428]" italics="true" pageId="12" pageNumber="313">3</emphasis>
(25) barbed, tapering, 
<emphasis id="B393250F5C22FFD4FB934DF36D7BD293" box="[1082,1114,404,428]" italics="true" pageId="12" pageNumber="313">st4</emphasis>
– 
<emphasis id="B393250F5C22FFD4FBC04DF26D56D293" box="[1129,1143,405,428]" italics="true" pageId="12" pageNumber="313">7</emphasis>
(~15) with inflated bases, whip-like tips; 
<emphasis id="B393250F5C22FFD4FEED4DDF6845D2EF" box="[324,356,440,464]" italics="true" pageId="12" pageNumber="313">st4</emphasis>
– 
<emphasis id="B393250F5C22FFD4FEDA4DDF68A0D2F0" box="[371,385,440,463]" italics="true" pageId="12" pageNumber="313">5</emphasis>
insertions approximate, 
<emphasis id="B393250F5C22FFD4FD324DDF6B9AD2EF" box="[667,699,440,464]" italics="true" pageId="12" pageNumber="313">st6</emphasis>
– 
<emphasis id="B393250F5C22FFD4FD604DDF6BF6D2F0" box="[713,727,440,463]" italics="true" pageId="12" pageNumber="313">7</emphasis>
insertions variable. Large (228 long, 355 wide) ventral shield present encompassing anal shield and bearing numerous setae, ornamented with tooth-like tubercles and denticles (
<figureCitation id="19DCE5985C22FFD4FF364D9869D2D128" box="[159,243,511,535]" captionStart="FIGURES 19 – 23" captionStartId="9.[151,264,1850,1873]" captionTargetBox="[163,1430,199,1820]" captionTargetId="figure@9.[151,1436,193,1829]" captionTargetPageId="9" captionText="FIGURES 19 – 23. Reginacharlottia lordhowensis sp. n., adult male. 19 Dorsal habitus. 20 Ventri-anal region. 21 Chelicera showing excrescence (arrow points to membranous cheliceral seta). 22 Cuticle of podonotal shield showing tubercles, denticles, cluster of raised muscle insertions (arrow). 23 Cuticle of ventral shield. Scale bars 19 – 20 = 100 µm; 21 – 23 = 10 µm." httpUri="https://zenodo.org/record/248014/files/figure.png" pageId="12" pageNumber="313">Fig. 23</figureCitation>
).
</paragraph>
<paragraph id="8158F91D5C22FFD4FF6E4E436D7BD198" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="313">
<emphasis id="B393250F5C22FFD4FF6E4E436842D103" bold="true" box="[199,355,548,572]" pageId="12" pageNumber="313">Deutonymph</emphasis>
(n = 2). Structure of gnathosoma, cuticular ornamentation, form of idiosomal setae and ornamentation of plates similar to adults. Podonotal shield truncate and separate from broad mesonotal plate bearing either 6 or 25 setae. Sternal region with five pairs of setae, but without distinct shield. Anal shield similar to adults, but with three pairs of small setae and overarching pair of hypertrophied setae.
</paragraph>
<paragraph id="8158F91D5C22FFD4FF6E4ED36AA6D008" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="313">
<emphasis id="B393250F5C22FFD4FF6E4ED3687CD1F3" bold="true" box="[199,349,692,716]" pageId="12" pageNumber="313">Protonymph</emphasis>
(n = 1). Similar to other stages but palpal trochanter with one seta; podonotal shield truncate posteriorly; a pair of small, nude mesonotal scutellae present; sternal region with weakly differentiated shield bearing 
<emphasis id="B393250F5C22FFD4FF5E4E9B6839D02B" box="[247,280,764,788]" italics="true" pageId="12" pageNumber="313">st1</emphasis>
– 
<emphasis id="B393250F5C22FFD4FE8F4E9B6815D02B" box="[294,308,764,788]" italics="true" pageId="12" pageNumber="313">3</emphasis>
; anal shield similar to deutonymph, but with two pairs of small setae and overarching pair of hypertrophied setae; tarsus IV with 
<emphasis id="B393250F5C22FFD4FD8E4F476B71D007" box="[551,592,800,824]" italics="true" pageId="12" pageNumber="313">av4</emphasis>
, 
<emphasis id="B393250F5C22FFD4FDF44F476BA7D007" box="[605,646,800,824]" italics="true" pageId="12" pageNumber="313">pv4</emphasis>
on intercalary sclerite.
</paragraph>
<paragraph id="8158F91D5C22FFD4FF6E4F236B7ED630" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="313">
<emphasis id="B393250F5C22FFD4FF6E4F236857D063" bold="true" box="[199,374,836,860]" pageId="12" pageNumber="313">Egg and larva.</emphasis>
(
<figureCitation id="19DCE5985C22FFD4FE2C4F236B24D063" box="[389,517,836,860]" captionStart="FIGURES 48 – 49" captionStartId="20.[151,264,1696,1719]" captionTargetBox="[156,1419,604,1668]" captionTargetId="figure@20.[151,1436,591,1675]" captionTargetPageId="20" captionText="FIGURES 48 – 49. Reginacharlottia lordhowensis sp. n. larva 48 Reconstruction of dorsal chaetotaxy (arrow points to posterior horn). 49 Anal region (pa = para-anal seta). Figure 50 Epicroseius sp. (Sejidae) Queensland, Australia, female genital shield with attached spermatophore. Scale bar 48 = 100 µm; 49 = 25 µm." httpUri="https://zenodo.org/record/248022/files/figure.png" pageId="12" pageNumber="313">Figs 48, 49</figureCitation>
) Female with up to 12 large (&gt;300 long) eggs with fully developed larvae. Larval idiosoma (~330 long) encased in lightly sclerotised, reticulate cuticle except smooth in intercoxal region. Dorsal seta 
<emphasis id="B393250F5C22FFD4FF644FE869C3D098" box="[205,226,911,935]" italics="true" pageId="12" pageNumber="313">j1</emphasis>
(86 long) densely barbed in distal 3/4th, widely separated (35 between insertions) (
<figureCitation id="19DCE5985C22FFD4FB194FE86C29D098" box="[1200,1288,911,935]" captionStart="FIGURES 48 – 49" captionStartId="20.[151,264,1696,1719]" captionTargetBox="[156,1419,604,1668]" captionTargetId="figure@20.[151,1436,591,1675]" captionTargetPageId="20" captionText="FIGURES 48 – 49. Reginacharlottia lordhowensis sp. n. larva 48 Reconstruction of dorsal chaetotaxy (arrow points to posterior horn). 49 Anal region (pa = para-anal seta). Figure 50 Epicroseius sp. (Sejidae) Queensland, Australia, female genital shield with attached spermatophore. Scale bar 48 = 100 µm; 49 = 25 µm." httpUri="https://zenodo.org/record/248022/files/figure.png" pageId="12" pageNumber="313">Fig. 48</figureCitation>
); other setae roughened with incipient barbs; 
<emphasis id="B393250F5C22FFD4FDBF4FD46B0AD0F4" box="[534,555,947,971]" italics="true" pageId="12" pageNumber="313">j3</emphasis>
(132 long, 28 between insertions), 
<emphasis id="B393250F5C22FFD4FC714FD46ACFD0F4" box="[984,1006,947,971]" italics="true" pageId="12" pageNumber="313">j4</emphasis>
– 
<emphasis id="B393250F5C22FFD4FC554FD46D2BD0F5" box="[1020,1034,947,970]" italics="true" pageId="12" pageNumber="313">5</emphasis>
, 
<emphasis id="B393250F5C22FFD4FBB44FD46D14D0F4" box="[1053,1077,947,971]" italics="true" pageId="12" pageNumber="313">z2</emphasis>
, 
<emphasis id="B393250F5C22FFD4FBE04FD46D43D0F4" box="[1097,1122,947,971]" italics="true" pageId="12" pageNumber="313">z4</emphasis>
, 
<emphasis id="B393250F5C22FFD4FBDC4FD46DAFD0F4" box="[1141,1166,947,971]" italics="true" pageId="12" pageNumber="313">s4</emphasis>
, 
<emphasis id="B393250F5C22FFD4FB084FD46D9AD0F4" box="[1185,1211,947,971]" italics="true" pageId="12" pageNumber="313">s6</emphasis>
, 
<emphasis id="B393250F5C22FFD4FB644FD46DC6D0F4" box="[1229,1255,947,971]" italics="true" pageId="12" pageNumber="313">J2</emphasis>
– 
<emphasis id="B393250F5C22FFD4FB5F4FD46C25D0F5" box="[1270,1284,947,970]" italics="true" pageId="12" pageNumber="313">5</emphasis>
, 
<emphasis id="B393250F5C22FFD4FABE4FD46C15D0F4" box="[1303,1332,947,971]" italics="true" pageId="12" pageNumber="313">Z3</emphasis>
– 
<emphasis id="B393250F5C22FFD4FAEA4FD46C70D0F5" box="[1347,1361,947,970]" italics="true" pageId="12" pageNumber="313">5</emphasis>
, 
<emphasis id="B393250F5C22FFD4FACD4FD46C5ED0F4" box="[1380,1407,947,971]" italics="true" pageId="12" pageNumber="313">S3</emphasis>
– 
<emphasis id="B393250F5C22FFD4FA274FD46CBDD0F4" box="[1422,1436,947,971]" italics="true" pageId="12" pageNumber="313">4</emphasis>
similarly ornamented (70–105 long), 
<emphasis id="B393250F5C22FFD4FD934FB06B76D0D1" box="[570,599,983,1006]" italics="true" pageId="12" pageNumber="313">S5</emphasis>
(30) simple; 
<emphasis id="B393250F5C22FFD4FD454FB06A23D0D0" box="[748,770,983,1007]" italics="true" pageId="12" pageNumber="313">j6</emphasis>
apparently absent; median horn-like pygidial tubercle (15– 20 high) present between 
<emphasis id="B393250F5C22FFD4FE6A4F9C68FFD72C" box="[451,478,1019,1043]" italics="true" pageId="12" pageNumber="313">J4</emphasis>
– 
<emphasis id="B393250F5C22FFD4FE454F9C6B27D72D" box="[492,518,1019,1042]" italics="true" pageId="12" pageNumber="313">J5</emphasis>
(
<figureCitation id="19DCE5985C22FFD4FDB14F9D6B4AD72D" box="[536,619,1018,1042]" captionStart="FIGURES 48 – 49" captionStartId="20.[151,264,1696,1719]" captionTargetBox="[156,1419,604,1668]" captionTargetId="figure@20.[151,1436,591,1675]" captionTargetPageId="20" captionText="FIGURES 48 – 49. Reginacharlottia lordhowensis sp. n. larva 48 Reconstruction of dorsal chaetotaxy (arrow points to posterior horn). 49 Anal region (pa = para-anal seta). Figure 50 Epicroseius sp. (Sejidae) Queensland, Australia, female genital shield with attached spermatophore. Scale bar 48 = 100 µm; 49 = 25 µm." httpUri="https://zenodo.org/record/248022/files/figure.png" pageId="12" pageNumber="313">Fig. 48</figureCitation>
arrow). Sternal setae 
<emphasis id="B393250F5C22FFD4FCC24F9C6AADD72C" box="[875,908,1019,1043]" italics="true" pageId="12" pageNumber="313">st1</emphasis>
– 
<emphasis id="B393250F5C22FFD4FC334F9C6A89D72C" box="[922,936,1019,1043]" italics="true" pageId="12" pageNumber="313">3</emphasis>
(25–32 long) acicular, weakly barbed; 
<emphasis id="B393250F5C22FFD4FAD94F9C6CBDD72C" box="[1392,1436,1019,1043]" italics="true" pageId="12" pageNumber="313">JV1</emphasis>
similar (32 long); 
<emphasis id="B393250F5C22FFD4FEC04847689FD708" box="[361,446,1055,1079]" italics="true" pageId="12" pageNumber="313">JV (5?)</emphasis>
(46) barbed. Anal opening (
<figureCitation id="19DCE5985C22FFD4FCA148786A7DD708" box="[776,860,1055,1079]" captionStart="FIGURES 48 – 49" captionStartId="20.[151,264,1696,1719]" captionTargetBox="[156,1419,604,1668]" captionTargetId="figure@20.[151,1436,591,1675]" captionTargetPageId="20" captionText="FIGURES 48 – 49. Reginacharlottia lordhowensis sp. n. larva 48 Reconstruction of dorsal chaetotaxy (arrow points to posterior horn). 49 Anal region (pa = para-anal seta). Figure 50 Epicroseius sp. (Sejidae) Queensland, Australia, female genital shield with attached spermatophore. Scale bar 48 = 100 µm; 49 = 25 µm." httpUri="https://zenodo.org/record/248022/files/figure.png" pageId="12" pageNumber="313">Fig. 49</figureCitation>
) simple, para-anal setae (70 long) weakly barbed, inserted posteriad opening, postanal seta absent. Chelicerae (~60 long), interdigital membrane just passing (~4) tip movable digit (15 long) possibly edentate; setae 
<emphasis id="B393250F5C22FFD4FD1348006BF7D740" box="[698,726,1127,1151]" italics="true" pageId="12" pageNumber="313">h1</emphasis>
(~10 long) tapering, acuminate, slightly inflated (candle-flame shaped); 
<emphasis id="B393250F5C22FFD4FEA948EC683DD79C" box="[256,284,1163,1187]" italics="true" pageId="12" pageNumber="313">h2</emphasis>
(~23 long) curved medially, barbed on outer distal side; corniculus minute (7–9), with three distal cusps. Deutosternal gutter obsolescent, with three rows of two denticles each. Tritosternum with simple columnar base, two thick laciniae (~35 long). Dorsal setae on palpgenu-tibia thick, blunt, curved (resemble solenidia); palptarsal apotele with strong basal spur.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C22FFD4FF6E497C6D2DD60D" box="[199,1036,1306,1331]" pageId="12" pageNumber="313" type="etymology">
<paragraph id="8158F91D5C22FFD4FF6E497C6D2DD60D" blockId="12.[151,1437,151,2016]" box="[199,1036,1306,1331]" pageId="12" pageNumber="313">
<emphasis id="B393250F5C22FFD4FF6E497C686FD60C" bold="true" box="[199,334,1307,1331]" pageId="12" pageNumber="313">Etymology.</emphasis>
The species name is genitive and refers to the island of origin.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C22FFD5FF6E49586819D2B9" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313" type="discussion">
<paragraph id="8158F91D5C22FFD4FF6E49586A9CD535" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="313">
<emphasis id="B393250F5C22FFD4FF6E4958681AD668" bold="true" box="[199,315,1343,1367]" pageId="12" pageNumber="313">Remarks.</emphasis>
Adult males and females of the new species were relatively frequent in soil-litter extractions (and one pit-fall trap sample), but never abundant compared with other 
<taxonomicName id="46E7829E5C22FFD4FC2D49056D04D645" box="[900,1061,1378,1402]" class="Arachnida" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="313" phylum="Arthropoda" rank="order">Mesostigmata</taxonomicName>
. The association of the following species with a large arthropod and the highly modified chelicerae and reduced legs I of both species in the new genus suggest that these mites may be associated with some larger animal. For example, the reduced legs I are reminiscent of some trigynaspid mites that are able to jump and are associated with beetles: 
<taxonomicName id="46E7829E5C22FFD4FB7C49A8685DD535" authority="Walter" authorityName="Walter" class="Arachnida" family="Saltiseiidae" genus="Saltiseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="hunteri">
<emphasis id="B393250F5C22FFD4FB7C49A86CBDD6D8" box="[1237,1436,1487,1511]" italics="true" pageId="12" pageNumber="313">Saltiseius hunteri</emphasis>
Walter (Saltiseiidae)
</taxonomicName>
and 
<emphasis id="B393250F5C22FFD4FE1B49946B87D534" box="[434,678,1523,1547]" italics="true" pageId="12" pageNumber="313">Micromegislus bakeri</emphasis>
Trägårdh (Walter 2000).
</paragraph>
<paragraph id="8158F91D5C22FFD4FF6E4A7068CAD415" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="313">
Other Sejoidea are known to disperse on insects associated with woody habitats. For example, deutonymphs of 
<taxonomicName id="46E7829E5C22FFD4FF3E4A5C68FED56D" authority="Berlese" authorityName="Berlese" box="[151,479,1594,1619]" class="Arachnida" family="Sejidae" genus="Uropodella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="lacinata">
<emphasis id="B393250F5C22FFD4FF3E4A5C685ED56C" box="[151,383,1595,1619]" italics="true" pageId="12" pageNumber="313">Uropodella lacinata</emphasis>
Berlese
</taxonomicName>
have been reported to be phoretic on the tenebrionid beetle 
<taxonomicName id="46E7829E5C22FFD4FB114A5B688BD548" authority="(Herbst) (Kethley 1983)" authorityName="(Herbst) (Kethley" authorityYear="1983" class="Insecta" family="Tenebrionidae" genus="Neatus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="313" phylum="Arthropoda" rank="species" species="tenebrioides">
<emphasis id="B393250F5C22FFD4FB114A5B6CBDD56C" box="[1208,1436,1595,1619]" italics="true" pageId="12" pageNumber="313">Neatus tenebrioides</emphasis>
(Herbst) (Kethley 1983)
</taxonomicName>
and deutonymphs of 
<taxonomicName id="46E7829E5C22FFD4FD0E4A386BD8D548" box="[679,761,1631,1655]" class="Arachnida" family="Sejidae" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="313" phylum="Arthropoda" rank="family">Sejidae</taxonomicName>
on cerambycid beetles (Halliday 2012). The deutonymphs attach to their beetle carriers via an anal stalk produced by a complex anal opening (Kethley 1983). Only two deutonymphs of the new species have been discovered and neither possessed a modified anal opening. However, the free mesonotal plate in one deutonymph was much larger than in the other specimen, and bore about four times as many setae. Given the difference in dorsal sclerotisation in the adults, these deutonymphs may represent the male and female, respectively.
</paragraph>
<paragraph id="8158F91D5C22FFD4FF6E4B506C6AD4A8" blockId="12.[151,1437,151,2016]" pageId="12" pageNumber="313">
Species of 
<taxonomicName id="46E7829E5C22FFD4FEE84B5068C8D470" box="[321,489,1847,1871]" class="Arachnida" family="Sejidae" genus="Asternolaelaps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="12" pageNumber="313" phylum="Arthropoda" rank="genus">
<emphasis id="B393250F5C22FFD4FEE84B5068C8D470" box="[321,489,1847,1871]" italics="true" pageId="12" pageNumber="313">Asternolaelaps</emphasis>
</taxonomicName>
are often associated with mammal nests (Lindquist 
<emphasis id="B393250F5C22FFD4FB874B5F6D7FD470" box="[1070,1118,1847,1871]" italics="true" pageId="12" pageNumber="313">et al</emphasis>
. 2009). Other than seabirds, however, Lord Howe Island has only a few rare endemic vertebrates (two lizards and a bat) and none that make nests in the closed forest litter habitats in which the mites were most frequently encountered (Hutton 1986).
</paragraph>
<paragraph id="8158F91D5C22FFD5FF6E4BC5681DD3C8" blockId="12.[151,1437,151,2016]" lastBlockId="13.[151,1436,151,247]" lastPageId="13" lastPageNumber="314" pageId="12" pageNumber="313">
The presence of fully developed eggs within four of the females examined and the egg burster-like pygidial tubercle on the larva suggest that hatching may occur at or very soon after oviposition. 
<emphasis id="B393250F5C22FFD4FB0D4BA06CBDD4E0" box="[1188,1436,1991,2015]" italics="true" pageId="12" pageNumber="313">Micromegislus bakeri</emphasis>
Trägårdh (Parantenulidae) is reported to give birth to active larvae on its host ground beetles (Nickel &amp; Elzinga 1970). Egg bursters have been reported in the Prostigmata (Marchiando 1981), but not to my knowledge in the 
<taxonomicName id="46E7829E5C23FFD5FF3E4CB86819D3C8" box="[151,312,223,247]" class="Arachnida" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="order">Mesostigmata</taxonomicName>
.
</paragraph>
<paragraph id="8158F91D5C23FFD5FF3E4D2B6819D2B9" blockId="13.[151,569,332,390]" pageId="13" pageNumber="314">
<taxonomicName id="46E7829E5C23FFD5FF3E4D2B68D2D259" box="[151,499,332,358]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="braziliensis" status="sp. nov.">
<emphasis id="B393250F5C23FFD5FF3E4D2B68D2D259" bold="true" box="[151,499,332,358]" italics="true" pageId="13" pageNumber="314">Reginacharlottia braziliensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A8A098745C23FFD5FE534D2A6B18D25A" box="[506,569,333,357]" pageId="13" pageNumber="314" rank="species">
<emphasis id="B393250F5C23FFD5FE534D2A6B18D25A" bold="true" box="[506,569,333,357]" pageId="13" pageNumber="314">sp. n.</emphasis>
</taxonomicNameLabel>
<figureCitation id="19DCE5985C23FFD5FF3E4D096819D2B9" box="[151,312,366,390]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="13" pageNumber="314">Figures 31–35</figureCitation>
</paragraph>
</subSubSection>
<caption id="D598A9955C23FFD5FF3E4B1C6D42D4D2" httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="13" pageNumber="314" targetBox="[157,1405,448,1894]" targetPageId="13">
<paragraph id="8158F91D5C23FFD5FF3E4B1C6D42D4D2" blockId="13.[151,1436,1915,2029]" pageId="13" pageNumber="314">
<emphasis id="B393250F5C23FFD5FF3E4B1C680BD4AF" bold="true" box="[151,298,1915,1938]" pageId="13" pageNumber="314">FIGURES 31</emphasis>
– 
<emphasis id="B393250F5C23FFD5FE9E4B1C6876D4AF" bold="true" box="[311,343,1915,1936]" pageId="13" pageNumber="314">35.</emphasis>
<taxonomicName id="46E7829E5C23FFD5FEC94B1C6BABD4AE" box="[352,650,1915,1937]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="314" phylum="Arthropoda" rank="species" species="brasiliensis" status="sp. nov.">
<emphasis id="B393250F5C23FFD5FEC94B1C6BABD4AE" box="[352,650,1915,1937]" italics="true" pageId="13" pageNumber="314">Reginacharlottia brasiliensis</emphasis>
</taxonomicName>
<emphasis id="B393250F5C23FFD5FD3D4B1B6BF1D4AE" bold="true" box="[660,720,1916,1937]" pageId="13" pageNumber="314">
<taxonomicNameLabel id="A8A098745C23FFD5FD3D4B1B6BF1D4AE" box="[660,720,1916,1937]" pageId="13" pageNumber="314" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, adult female. 31 Holotype female on amblypygid 
<taxonomicName id="46E7829E5C23FFD5FB594B1C6C56D4AE" box="[1264,1399,1915,1937]" class="Arachnida" family="Phrynichidae" genus="Trichodamon" kingdom="Animalia" order="Amblypygi" pageId="13" pageNumber="314" phylum="Arthropoda" rank="genus">
<emphasis id="B393250F5C23FFD5FB594B1C6C56D4AE" box="[1264,1399,1915,1937]" italics="true" pageId="13" pageNumber="314">Trichodamon</emphasis>
</taxonomicName>
aff 
<emphasis id="B393250F5C23FFD5FF3E4BFD69F1D48F" box="[151,208,1946,1968]" italics="true" pageId="13" pageNumber="314">froesi</emphasis>
(photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm.
</paragraph>
</caption>
<subSubSection id="C9FDAA965C20FFD6FF3E4CF06D02D3C8" pageId="14" pageNumber="315" type="description">
<paragraph id="8158F91D5C20FFD6FF3E4CF06D02D3C8" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="315">
<emphasis id="B393250F5C20FFD6FF3E4CF068BDD38F" bold="true" box="[151,412,151,176]" pageId="14" pageNumber="315">Specimens examined.</emphasis>
<typeStatus id="5E5C47BF5C20FFD6FE004CF06B30D390" box="[425,529,151,175]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
female, ex 
<taxonomicName id="46E7829E5C20FFD6FD0F4CFF6AA1D38F" box="[678,896,152,176]" class="Arachnida" family="Phrynichidae" genus="Trichodamon" kingdom="Animalia" order="Amblypygi" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="froes">
<emphasis id="B393250F5C20FFD6FD0F4CFF6AA1D38F" box="[678,896,152,176]" italics="true" pageId="14" pageNumber="315">Trichodamon froes</emphasis>
</taxonomicName>
i Mello-Leitão, 1940 (Arachnida: 
<taxonomicName id="46E7829E5C20FFD6FABC4CF06CB8D390" box="[1301,1433,151,175]" class="Arachnida" kingdom="Animalia" order="Amblypygi" pageId="14" pageNumber="315" phylum="Arthropoda" rank="order">Amblypygi</taxonomicName>
: Phyrnidae), Caverna Agua Fina, Carinhanha, Bahia State, 
<collectingCountry id="F9F0B98D5C20FFD6FC8D4CDB6A4AD3EB" box="[804,875,188,212]" name="Brazil" pageId="14" pageNumber="315">Brazil</collectingCountry>
, 
<date id="F559DFDD5C20FFD6FCDF4CDB6D2FD3EB" box="[886,1038,188,213]" pageId="14" pageNumber="315" value="2012-05-29">29 May 2012</date>
, M.E. Bichuett &amp; J.E. Gallão. 
<typeStatus id="5E5C47BF5C20FFD6FACD4CDA6CBDD3EA" box="[1380,1436,189,213]" pageId="14" pageNumber="315">Type</typeStatus>
deposited in Department of Zoology, UNESP, Sao Paulo State University, 
<collectingCountry id="F9F0B98D5C20FFD6FC7E4CB86D3ED3C8" box="[983,1055,223,247]" name="Brazil" pageId="14" pageNumber="315">Brazil</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C20FFD6FF6E4D646CB2D198" pageId="14" pageNumber="315" type="diagnosis">
<paragraph id="8158F91D5C20FFD6FF6E4D646CB2D198" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="315">
<emphasis id="B393250F5C20FFD6FF6E4D646860D223" bold="true" box="[199,321,259,284]" pageId="14" pageNumber="315">Diagnosis.</emphasis>
With the characteristics of the genus; podonotal-metanotal shield (635 long) densely setose and tuberculate: cuticular tubercles polygonal to substellate (
<figureCitation id="19DCE5985C20FFD6FCB04D406A4CD200" box="[793,877,295,319]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="14" pageNumber="315">Fig. 34</figureCitation>
), more or less contiguous; ca. 30 pairs of sigillar tubercles in podonotal region, metanotal region with two clusters of six sigillar tubercles; metapodal plates fused and joined to podonotal region at transverse fold; peritrematal-metapodal shield with fishscale-like reticulate ornamentation (
<figureCitation id="19DCE5985C20FFD6FEE34DF36881D293" box="[330,416,404,428]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="14" pageNumber="315">Fig. 32</figureCitation>
), large gland opening about mid-shield; movable cheliceral digit (
<figureCitation id="19DCE5985C20FFD6FB3E4DF36DCBD293" box="[1175,1258,404,428]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="14" pageNumber="315">Fig. 33</figureCitation>
) ending in five minute teeth; soft cuticle with smoothly net-like ornamentation of narrowly elongate hexagons (
<figureCitation id="19DCE5985C20FFD6FB7A4DD06C07D2F0" box="[1235,1318,439,463]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="14" pageNumber="315">Fig. 32</figureCitation>
), irregular polygons, and substellate cells. Tectum with smooth, rounded anterior margin. Genital shield (~105 long) with polygonal reticulation, surrounded by striate cuticle and seven pairs of setae (
<figureCitation id="19DCE5985C20FFD6FBBE4D986D4CD128" box="[1047,1133,511,535]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="14" pageNumber="315">Fig. 35</figureCitation>
). Dorsal setae 22–30 long with ventral vane and dorsal conifer cone-like ornamentation, ventral setae similar, but strongly tapered, acuminate, barbs short. Leg setation: coxa I–IV: 3-3-2-1; trochanters I-IV: 6-5-5-5; femur I-IV: 11 (v = short spine) - 10 (2 basifemur) - 7-8 (1 on basifemur); genu I-IV: 8-11 (2-3/1, 3/0-2) - 10 (2-2/1, 2/1-2) - 11 (1-2/1, 3/1-2); tibia I-IV: 8-10 (2-2/1, 2/1-2) - 10 (2-2/1, 2/1-2) - 10 (2-2/1, 2/1-2); tarsus IV with intercalary sclerite bearing 
<emphasis id="B393250F5C20FFD6FA844EF76C77D197" box="[1325,1366,656,680]" italics="true" pageId="14" pageNumber="315">av4</emphasis>
, 
<emphasis id="B393250F5C20FFD6FACD4EF76CADD197" box="[1380,1420,656,680]" italics="true" pageId="14" pageNumber="315">pv4</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C9FDAA965C20FFD6FF6E4ED36BE0D70B" pageId="14" pageNumber="315" type="discussion">
<paragraph id="8158F91D5C20FFD6FF6E4ED36BE0D70B" blockId="14.[151,1437,151,1077]" pageId="14" pageNumber="315">
<emphasis id="B393250F5C20FFD6FF6E4ED3681AD1F3" bold="true" box="[199,315,692,716]" pageId="14" pageNumber="315">Remarks.</emphasis>
Although only a single female specimen has been collected and the slide preparation is in poor condition (the body has burst, the sternal region cannot be observed, the subcapitulum is only partially visible, the pygidium is apparently absent), it seems appropriate to provide a diagnosis and name for the species to encourage its further study. The Brazilian specimen shares the diagnostic characteristics of 
<taxonomicName id="46E7829E5C20FFD6FB914F476DD7D007" box="[1080,1270,800,824]" class="Arachnida" family="Uropodellidae" genus="Reginacharlottia" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="14" pageNumber="320" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B393250F5C20FFD6FB914F476DD7D007" box="[1080,1270,800,824]" italics="true" pageId="14" pageNumber="315">Reginacharlottia</emphasis>
</taxonomicName>
, including the unique cheliceral modifications, setiform corniculus, palpal apotele with dorsal spur, rake-like ventral seta on the palptrochanter, uniquely formed ventri-anal shield, and conifer cone-like dorsal setae. However, it differs from the 
<typeStatus id="5E5C47BF5C20FFD6FF3E4FEA69E6D09A" box="[151,199,909,933]" pageId="14" pageNumber="315">type</typeStatus>
species in the details of the ornamentation of the shields, the metanotal region (a single continuous metanotal plate bearing both pairs of muscle insertions is fused to the podonotal shield, see 
<figureCitation id="19DCE5985C20FFD6FB8A4FC86D57D0F8" box="[1059,1142,943,967]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="14" pageNumber="315">Fig. 34</figureCitation>
), the ornamentation of the soft cuticle (jigsaw-like patterns of deformed stellate cells or compressed hexagons, see 
<figureCitation id="19DCE5985C20FFD6FB694FB36C39D0D3" box="[1216,1304,980,1004]" captionStart="FIGURES 31 – 35" captionStartId="13.[151,264,1915,1938]" captionTargetBox="[157,1405,448,1894]" captionTargetId="figure@13.[151,1435,446,1894]" captionTargetPageId="13" captionText="FIGURES 31 – 35. Reginacharlottia brasiliensis sp. n., adult female. 31 Holotype female on amblypygid Trichodamon aff froesi (photo by Adriano Gambarini). 32 Details of ornamentation of soft cuticle and peritrematal shield (bottom). 33 Venter of palps and chelicerae (arrow points to interdigital membrane). 34 Podonotal shield showing distribution of sigillae and details of ornamentation. 35 Genital shield and associated setae. Scale bars 33, 35 = 50 µm; 34 = 500 µm." httpUri="https://zenodo.org/record/248017/files/figure.png" pageId="14" pageNumber="315">Fig. 32</figureCitation>
), the more extensive distribution of tapered ventral setae, and some details of the leg setation, especially of the more reduced setation of the genua (
<date id="F559DFDD5C20FFD6FE26487B6B2AD70B" box="[399,523,1052,1076]" pageId="14" pageNumber="315" value="2011-08-11" valueMax="2011-08-10" valueMin="2011-08-11">8-11-10-11</date>
vs 
<date id="F559DFDD5C20FFD6FD9B487B6B95D70B" box="[562,692,1052,1076]" pageId="14" pageNumber="315" value="2012-09-12" valueMax="2012-09-12" valueMin="2012-09-12">9-12-12-12</date>
).
</paragraph>
</subSubSection>
</treatment>
</document>