129 lines
17 KiB
XML
129 lines
17 KiB
XML
<document id="A68E14F356C58F20B990D72D4046B39A" ID-DOI="10.11646/zootaxa.4576.3.6" ID-GBIF-Dataset="7abdc62c-c350-49dd-8060-4ed64da07362" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715772" ID-ZooBank="56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603665960" checkinUser="plazi" docAuthor="Muona, Jyrki" docDate="2019" docId="03D387E9FFAF7777FF6FEBF1FA89FE0A" docLanguage="en" docName="zootaxa.4576.3.6.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrannothroscus Muona 2019, new genus" docType="treatment" docVersion="5" lastPageNumber="530" masterDocId="FFEAFF91FFA7777EFFF8EE40FF81FFA7" masterDocTitle="Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species" masterLastPageNumber="543" masterPageNumber="521" pageNumber="529" updateTime="1698730058212" updateUser="plazi">
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<mods:title id="FD7B0BC102689A64562B45CFF7D07AF9">Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species</mods:title>
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<mods:namePart id="79C67A2D5EFA53B65AC0E0F337CB5337">Muona, Jyrki</mods:namePart>
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<mods:title id="6A9BA4BA48260B477B36CD10A0D2A24C">Zootaxa</mods:title>
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<mods:part id="C750333CD113F67A113773F70C1DF8E1">
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<mods:date id="A2F65FD07C72283F21C3557A96F4B9C7">2019</mods:date>
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<mods:number id="8D606D1763E2D4DE6AE9EC41562426C4">2019-04-03</mods:number>
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<mods:number id="B471CC6D4B89AC45F4197FC919F563E4">4576</mods:number>
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<treatment id="03D387E9FFAF7777FF6FEBF1FA89FE0A" ID-DOI="http://doi.org/10.5281/zenodo.3716788" ID-GBIF-Taxon="162639248" ID-Zenodo-Dep="3716788" LSID="urn:lsid:plazi:treatment:03D387E9FFAF7777FF6FEBF1FA89FE0A" httpUri="http://treatment.plazi.org/id/03D387E9FFAF7777FF6FEBF1FA89FE0A" lastPageId="9" lastPageNumber="530" pageId="8" pageNumber="529">
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<subSubSection id="C3606574FFAF7776FF6FEBF1FE6CFA6C" box="[151,493,1457,1483]" pageId="8" pageNumber="529" type="nomenclature">
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<paragraph id="8BC536FFFFAF7776FF6FEBF1FE6CFA6C" blockId="8.[151,493,1457,1483]" box="[151,493,1457,1483]" pageId="8" pageNumber="529">
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<heading id="D08D8193FFAF7776FF6FEBF1FE6CFA6C" bold="true" box="[151,493,1457,1483]" fontSize="11" level="1" pageId="8" pageNumber="529" reason="1">
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<emphasis id="B90EEAEDFFAF7776FF6FEBF1FE6CFA6C" bold="true" box="[151,493,1457,1483]" pageId="8" pageNumber="529">
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<taxonomicName id="4C7A4D7CFFAF7776FF6FEBF1FEE4FA6C" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[151,357,1457,1483]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFAF7776FF6FEBF1FEE4FA6C" bold="true" box="[151,357,1457,1483]" italics="true" pageId="8" pageNumber="529">Tyrannothroscus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D5796FFAF7776FE94EBF1FE6CFA6C" box="[364,493,1457,1483]" pageId="8" pageNumber="529" rank="genus">new genus</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAF7776FF6FEBB7FDF9F9AA" box="[151,632,1527,1550]" pageId="8" pageNumber="529" type="materials_examined">
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<paragraph id="8BC536FFFFAF7776FF6FEBB7FDF9F9AA" blockId="8.[151,632,1527,1550]" box="[151,632,1527,1550]" pageId="8" pageNumber="529">
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<emphasis id="B90EEAEDFFAF7776FF6FEBB7FE9DF9AA" bold="true" box="[151,284,1527,1549]" pageId="8" pageNumber="529">
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<typeStatus id="54C1885DFFAF7776FF6FEBB7FF4DF9AA" box="[151,204,1527,1549]" pageId="8" pageNumber="529">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="4C7A4D7CFFAF7776FED2EBB7FE79F9A9" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[298,504,1527,1550]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B90EEAEDFFAF7776FED2EBB7FE79F9A9" box="[298,504,1527,1550]" italics="true" pageId="8" pageNumber="529">Tyrannothroscus rex</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D5796FFAF7776FE07EBB8FDF9F9AA" box="[511,632,1527,1549]" pageId="8" pageNumber="529" rank="species">new species</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAF7776FF6FE87AFACCF830" pageId="8" pageNumber="529" type="diagnosis">
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<paragraph id="8BC536FFFFAF7776FF6FE87AFBFBF9F4" blockId="8.[151,1436,1594,2015]" box="[151,1146,1594,1619]" pageId="8" pageNumber="529">
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<emphasis id="B90EEAEDFFAF7776FF6FE87AFE8BF9F4" bold="true" box="[151,266,1594,1619]" pageId="8" pageNumber="529">Diagnosis</emphasis>
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. Elateroidea synapomorphy: pro-mesothoracic joint with clicking mechanism.
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</paragraph>
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<paragraph id="8BC536FFFFAF7776FF6FE81EFED1F8E8" blockId="8.[151,1436,1594,2015]" pageId="8" pageNumber="529">
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<taxonomicName id="4C7A4D7CFFAF7776FF6FE81EFE95F9D0" authorityName="Laporte" authorityYear="1840" box="[151,276,1630,1655]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="family">Throscidae</taxonomicName>
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synapomorphies: a2 as wide as a3–a4, antennal grooves running along notosternal suture and then bending laterally along proleg cavity (
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<figureCitation id="13412A7AFFAF7776FDB2E8C2FD0EF93C" box="[586,655,1666,1691]" captionStart="FIGURE 6" captionStartId="10.[151,250,873,895]" captionTargetBox="[151,1436,193,860]" captionTargetId="figure@10.[151,1436,193,860]" captionTargetPageId="10" captionText="FIGURE 6. Tyrannothroscus rex n. sp., ventral view. A = antennal groove; MtG = metathoracic groove; AtG = abdominal tarsal groove, SAR = supra-antennal ridge." figureDoi="http://doi.org/10.5281/zenodo.3715784" httpUri="https://zenodo.org/record/3715784/files/figure.png" pageId="8" pageNumber="529">Fig. 6</figureCitation>
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). Synapomorphies of
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<emphasis id="B90EEAEDFFAF7776FC73E8C2FBA3F93C" box="[907,1058,1666,1691]" italics="true" pageId="8" pageNumber="529">Rhomboaspis</emphasis>
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+
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<taxonomicName id="4C7A4D7CFFAF7776FBBAE8C2FB7CF93C" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[1090,1277,1666,1691]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFAF7776FBBAE8C2FB7CF93C" box="[1090,1277,1666,1691]" italics="true" pageId="8" pageNumber="529">Tyrannothroscus</emphasis>
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</taxonomicName>
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+ Potergini +
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<taxonomicName id="4C7A4D7CFFAF7776FF6FE8E6FE86F918" box="[151,263,1702,1727]" class="Insecta" family="Throscidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="tribe" tribe="Throscini">Throscini</taxonomicName>
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: antennal grooves and proleg impressions separated by septum, metaventrite with tarsal grooves, abdomen with tarsal grooves. Synapomorphies of
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<emphasis id="B90EEAEDFFAF7776FD3EE88AFCDCF944" box="[710,861,1738,1763]" italics="true" pageId="8" pageNumber="529">Rhomboaspis</emphasis>
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+
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<taxonomicName id="4C7A4D7CFFAF7776FC82E88AFBB4F944" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[890,1077,1738,1763]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFAF7776FC82E88AFBB4F944" box="[890,1077,1738,1763]" italics="true" pageId="8" pageNumber="529">Tyrannothroscus</emphasis>
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</taxonomicName>
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: elytra with strong and sharp, at most minutely punctate striae, body form very wide.
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<taxonomicName id="4C7A4D7CFFAF7776FCE8E8AEFC4AF8A0" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[784,971,1774,1799]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFAF7776FCE8E8AEFC4AF8A0" box="[784,971,1774,1799]" italics="true" pageId="8" pageNumber="529">Tyrannothroscus</emphasis>
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</taxonomicName>
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apomorphies: head with strong carinae above antennal insertions (
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<figureCitation id="13412A7AFFAF7776FE2BE952FD9DF88C" box="[467,540,1810,1835]" captionStart="FIGURE 7" captionStartId="10.[151,250,1949,1971]" captionTargetBox="[162,1431,968,1932]" captionTargetId="figure@10.[162,1431,968,1932]" captionTargetPageId="10" captionText="FIGURE 7. Tyrannothroscus rex n. sp., head, ventrolateral view, slightly frontally. A = antennal groove; E = eye; F = frontoclypeal region; C = labrum; PL = ventrally bent fron margin of prosternum; PP = prosternal process; SAR = supraantennal ridge." figureDoi="http://doi.org/10.5281/zenodo.3715788" httpUri="https://zenodo.org/record/3715788/files/figure.png" pageId="8" pageNumber="529">Fig. 7</figureCitation>
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), metasternal discrimen well developed, meso- and metatarsomeres 4 lobed beneath (
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<figureCitation id="13412A7AFFAF7776FF05E976FEC3F8E8" box="[253,322,1846,1871]" captionStart="FIGURE 8" captionStartId="11.[151,250,1081,1103]" captionTargetBox="[152,1435,210,1031]" captionTargetId="figure@11.[152,1435,210,1031]" captionTargetPageId="11" captionText="FIGURE 8. Tyrannothroscus rex n. sp., metatarsus. t4 = lobed tarsomere 4." figureDoi="http://doi.org/10.5281/zenodo.3715790" httpUri="https://zenodo.org/record/3715790/files/figure.png" pageId="8" pageNumber="529">Fig. 8</figureCitation>
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).
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</paragraph>
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<paragraph id="8BC536FFFFAF7776FF3FE91AFACCF830" blockId="8.[151,1436,1594,2015]" pageId="8" pageNumber="529">Diagnostic other characters: pronotal lateral carinae complete, head without median carina, antennal insertions located in fossae, antennae with apical, symmetrical club, prosternal process basally straight in lateral view.</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAF7777FF3FE9E2FBFCFE2F" lastPageId="9" lastPageNumber="530" pageId="8" pageNumber="529" type="description">
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<paragraph id="8BC536FFFFAF7777FF3FE9E2FBFCFE2F" blockId="8.[151,1436,1594,2015]" lastBlockId="9.[151,1436,151,429]" lastPageId="9" lastPageNumber="530" pageId="8" pageNumber="529">
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<emphasis id="B90EEAEDFFAF7776FF3FE9E2FED0F81C" bold="true" box="[199,337,1954,1979]" pageId="8" pageNumber="529">Description</emphasis>
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. Body wide, flattened, pronotum strongly narrowing anteriorly (
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<figureCitation id="13412A7AFFAF7776FBBEE9E2FB18F81C" box="[1094,1177,1954,1979]" captionStart="FIGURE 6" captionStartId="10.[151,250,873,895]" captionTargetBox="[151,1436,193,860]" captionTargetId="figure@10.[151,1436,193,860]" captionTargetPageId="10" captionText="FIGURE 6. Tyrannothroscus rex n. sp., ventral view. A = antennal groove; MtG = metathoracic groove; AtG = abdominal tarsal groove, SAR = supra-antennal ridge." figureDoi="http://doi.org/10.5281/zenodo.3715784" httpUri="https://zenodo.org/record/3715784/files/figure.png" pageId="8" pageNumber="529">Figs. 6</figureCitation>
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,
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<figureCitation id="13412A7AFFAF7776FB51E9E3FB38F81C" box="[1193,1209,1955,1979]" captionStart="FIGURE 9" captionStartId="11.[151,250,1998,2020]" captionTargetBox="[151,1436,1177,1948]" captionTargetId="figure@11.[151,1436,1177,1948]" captionTargetPageId="11" captionText="FIGURE 9. Tyrannothroscus rex n. sp., dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3715792" httpUri="https://zenodo.org/record/3715792/files/figure.png" pageId="8" pageNumber="529">9</figureCitation>
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). Head with strong supraocular ridges uniting in mid-line, frontoclypeus narrow, elongate, eyes large, antennae stout, a1 wide and short, a2 shorter than a3 and a4 but as wide, a9–a11 forming a club. Clypeus well slerotized, small. Anterior edge of prosternum strongly developed, projecting downwards as a wide arched extension. Metaventrite large and wide, cavity for pro- and mesolegs well-developed, ridged around and with separate sections for both legs, proleg area exceptionally wide, metatarsal grooves long, lateroposteriorly directed (
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<figureCitation id="13412A7AFFAE7777FC01EF44FBC4FEBA" box="[1017,1093,260,285]" captionStart="FIGURE 6" captionStartId="10.[151,250,873,895]" captionTargetBox="[151,1436,193,860]" captionTargetId="figure@10.[151,1436,193,860]" captionTargetPageId="10" captionText="FIGURE 6. Tyrannothroscus rex n. sp., ventral view. A = antennal groove; MtG = metathoracic groove; AtG = abdominal tarsal groove, SAR = supra-antennal ridge." figureDoi="http://doi.org/10.5281/zenodo.3715784" httpUri="https://zenodo.org/record/3715784/files/figure.png" pageId="9" pageNumber="530">Fig. 6</figureCitation>
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). Prosternal process slightly narrowed in middle, diverging strongly posteriorly, lateral carniae complete and apex rounded (
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<figureCitation id="13412A7AFFAE7777FAEDEF67FAE8FEE7" box="[1301,1385,295,320]" captionStart="FIGURE 1" captionStartId="2.[151,250,1200,1222]" captionTargetBox="[172,1421,396,1189]" captionTargetId="figure@2.[172,1421,396,1189]" captionTargetPageId="2" captionText="FIGURE 1. Pactopus horni LeConte. Head and thorax, ventrolateral view. A = antennal groove; F = depression for mesofemur; M = mesothoracic tarsal groove; mc = mesocoxa; pc = procoxa; S = septum; T = depression for mesotibia." figureDoi="http://doi.org/10.5281/zenodo.3715774" httpUri="https://zenodo.org/record/3715774/files/figure.png" pageId="9" pageNumber="530">Figs. 1</figureCitation>
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,
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<figureCitation id="13412A7AFFAE7777FA86EF67FA0FFEE7" box="[1406,1422,295,320]" captionStart="FIGURE 6" captionStartId="10.[151,250,873,895]" captionTargetBox="[151,1436,193,860]" captionTargetId="figure@10.[151,1436,193,860]" captionTargetPageId="10" captionText="FIGURE 6. Tyrannothroscus rex n. sp., ventral view. A = antennal groove; MtG = metathoracic groove; AtG = abdominal tarsal groove, SAR = supra-antennal ridge." figureDoi="http://doi.org/10.5281/zenodo.3715784" httpUri="https://zenodo.org/record/3715784/files/figure.png" pageId="9" pageNumber="530">6</figureCitation>
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). Metacoxae with prominent median apex, abdominal tarsal grooves extending to ventrite four. Elytral striae sharp, interstices flat, wide. Tibiae simple, without tarsal grooves, tarsi delicate, t4 lobed below.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF3FEFD4FA89FE0A" box="[199,1288,404,429]" pageId="9" pageNumber="530" type="etymology">
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<paragraph id="8BC536FFFFAE7777FF3FEFD4FA89FE0A" blockId="9.[151,1436,151,429]" box="[199,1288,404,429]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF3FEFD4FEC7FE0A" bold="true" box="[199,326,404,429]" pageId="9" pageNumber="530">Etymology</emphasis>
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.
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<taxonomicName id="4C7A4D7CFFAE7777FEACEFD4FD8EFE0A" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[340,527,404,429]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="529" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B90EEAEDFFAE7777FEACEFD4FD8EFE0A" box="[340,527,404,429]" italics="true" pageId="9" pageNumber="530">Tyrannothroscus</emphasis>
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</taxonomicName>
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refers to the relatively large size of this throscid, the largest known.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |