treatments-xml/data/D5/AA/E2/D5AAE2668C795040A997B57F4BB0D001.xml
2024-06-21 12:53:16 +02:00

192 lines
20 KiB
XML

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak &amp; Panha, Somsak" docDate="2022" docId="D5AAE2668C795040A997B57F4BB0D001" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Rhaphaulus lorraini Pfeiffer 1856" docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Panha, Somsak</mods:namePart>
<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand &amp; Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-08-25</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1119</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>115</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="199839049" LSID="urn:lsid:plazi:treatment:D5AAE2668C795040A997B57F4BB0D001" httpUri="http://treatment.plazi.org/id/D5AAE2668C795040A997B57F4BB0D001" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="D5AAE266-8C79-5040-A997-B57F4BB0D001" authority="Pfeiffer, 1856" authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus lorraini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorraini">Rhaphaulus lorraini Pfeiffer, 1856</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">Fig. 13A, B</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus lorraini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorraini">Rhaphaulus lorraini</taxonomicName>
Pfeiffer, 1856a: 36. Type locality: Pulo Penang [Penang Island, Penang State, Malaysia].
<bibRefCitation author="Pfeiffer, L" journalOrPublisher="Proceedings of the Zoological Society of London" pageId="0" pageNumber="1" refId="B125" refString="Pfeiffer, L, 1856b. Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles, extramarines nouvelles, ou peu connues. Beschreibung und Abbidung neuer order kritischer Land- und Suesswasser Mollusken. Tome 1, Lief 1-12, pp. 1-138, pls 1-36. [Lief. 5-8: 49-90, pls 13-24 (1856)] [Published in parts, dates follow Johnson (1969)]" title="Novitates Conchologicae. Series prima. Mollusca extramarina. Descriptions et figures de coquilles, extramarines nouvelles, ou peu connues. Beschreibung und Abbidung neuer order kritischer Land- und Suesswasser Mollusken. Tome 1, Lief 1 - 12, pp. 1 - 138, pls 1 - 36. [Lief. 5 - 8: 49 - 90, pls 13 - 24 (1856)] [Published in parts, dates follow Johnson (1969)]" year="1856 b">Pfeiffer 1856b</bibRefCitation>
: 75, pl. 20, figs 21, 22.
<bibRefCitation author="von Martens, E" journalOrPublisher="Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, Berlin" pageId="0" pageNumber="1" refId="B187" refString="von Martens, E, 1867. Die Preussische Expedition nach Ost-Asien. Verlag der Koeniglichen Geheimen Ober-Hofbuchdruckerei, Berlin" title="Die Preussische Expedition nach Ost-Asien." year="1867">von Martens 1867</bibRefCitation>
: 155.
<bibRefCitation DOI="https://doi.org/10.5962/bhl.part.14301" author="de, Morgan J" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="0" pageNumber="1" pagination="353 - 428" refId="B32" refString="de, Morgan J, 1885. Mollusques terrestres &amp; fluviatiles du royaume de Perak et des pays voisins (presqu'ile Malaise). Bulletin de la Societe Zoologique de France 10: 353 - 428, DOI: https://doi.org/10.5962/bhl.part.14301" title="Mollusques terrestres &amp; fluviatiles du royaume de Perak et des pays voisins (presqu'ile Malaise)." url="https://doi.org/10.5962/bhl.part.14301" volume="10" year="1885">de Morgan 1885</bibRefCitation>
: 413.
<bibRefCitation author="Smith, EA" journalOrPublisher="Proceedings of the Malacological Society of London" pageId="0" pageNumber="1" pagination="17 - 19" refId="B145" refString="Smith, EA, 1898. Description of Rhaphaulus perakensis, n.sp., with a list of the known species of the genus. Proceedings of the Malacological Society of London 3: 17 - 19" title="Description of Rhaphaulus perakensis, n. sp., with a list of the known species of the genus." volume="3" year="1898">Smith 1898</bibRefCitation>
: 18, figs 3, 4.
<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
: 276.
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
: 33, Penang.
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 42, West Malaysia.
<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2014
</bibRefCitation>
: 572, fig. 9.
<bibRefCitation author="Bavay, A" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" refId="B8" refString="BEDO (2017) Land Snails: Checklist of Molluscan Biodiversity in Thailand.BEDO, Bangkok, 300 pp. [in Thai]" year="1904">BEDO 2017</bibRefCitation>
: 96.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus lorainii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorainii">Rhaphaulus lorainii</taxonomicName>
[sic]-Sowerby I 1866:
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1853" family="Pupinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" rank="family">Pupinidae</taxonomicName>
, pl. 2 (pl. 264),
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">Rhaphaulus</taxonomicName>
, fig. 5.
<bibRefCitation author="Reeve, L" journalOrPublisher="Journal of Systematic Palaeontology" pageId="0" pageNumber="1" refId="B139" refString="Reeve, L, 1878. Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1-10. Lovell Reeve &amp; Co, London, [without pagination]." title="Conchologia Iconica: Illustrations of the shells of molluscous animals. Volume 20, Monograph of the genus Pupinidae, pls 1 - 10. Lovell Reeve &amp; Co, London, [without pagination]." year="1878">Reeve 1878</bibRefCitation>
:
<taxonomicName authorityName="L. Pfeiffer" authorityYear="1853" family="Pupinidae" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="" order="Architaenioglossa" pageId="0" pageNumber="1" rank="family">Pupinidae</taxonomicName>
, pl. 10, sp. 96.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus lorrainii" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorrainii">Rhaphaulus lorrainii</taxonomicName>
[sic]-
<bibRefCitation author="Habe, T" journalOrPublisher="Nature and Life in Southeast Asia" pageId="0" pageNumber="1" pagination="111 - 127" refId="B63" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and Life in Southeast Asia 4: 111 - 127" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe 1965</bibRefCitation>
: 115, 116, pl. 2, fig. 12, as a synonym of
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">Rhaphaulus chrysalis</taxonomicName>
, Khao Chong, Trang Province, peninsular Thailand.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName baseAuthorityName="Pfeiffer" baseAuthorityYear="1853" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus chrysalis" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="chrysalis">Rhaphaulus chrysalis</taxonomicName>
?-
<bibRefCitation author="Maassen, WJM" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B87" refString="Maassen, WJM, 2001. A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1-155." title="A preliminary checklist of the non-marine molluscs of West-Malaysia, A handlist. De Kreukel (Supplement): 1 - 155." year="2001">Maassen 2001</bibRefCitation>
: 42, West Malaysia.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
NHMUK 20130454 (3 shells; Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">13A</figureCitation>
) from Pulo Penang.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
CUMZ 12162 (1 shell; Fig.
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. A, B Rhaphaulus lorraini: A syntype NHMUK 20130454 from Pulo Penang and B specimen CUMZ 12162 from Kiriwong (Tham Kope) Temple, Phang Nga C Rhaphaulus perakensis, syntype NHMUK 1897.3.15.41 from Larut, Perak D Rhaphaulus ascendens, syntype UMZC I. 100025 from Patalung, Malay Peninsula E, F Rhaphaulus tonkinensis E holotype HNHM 98757 and F specimen CUMZ 12163 / 1 from Luang Cave, Chiang Rai. Photo: B. Pall-Gergely (E)" figureDoi="10.3897/zookeys.1119.85400.figure13" httpUri="https://binary.pensoft.net/fig/735374" pageId="0" pageNumber="1">13B</figureCitation>
) from Kiriwong (Tham Kope) Temple, Thap Put District, Phang Nga Province,
<date value="2009-01-16">16 Jan. 2009</date>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell ovate; body whorls bulging. Tube cylindrical, pointing upward and forward.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Pfeiffer" authorityYear="1856" class="Gastropoda" family="Pupinidae" genus="Rhaphaulus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Rhaphaulus lorraini" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="lorraini">
<emphasis italics="true" pageId="0" pageNumber="1">Rhaphaulus lorraini</emphasis>
</taxonomicName>
can be distinguished from all other species from mainland Southeast Asia by a cylindrical tube pointing upward and forward.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
Malaysia and southern Thailand (
<bibRefCitation author="Laidlaw, FF" journalOrPublisher="Journal of the Malaysian Branch of the Royal Asiatic Society" pageId="0" pageNumber="1" pagination="25 - 37" refId="B82" refString="Laidlaw, FF, 1928. A list of the land and fresh-water mollusca of the Malay Peninsula with notes. Journal of the Malaysian Branch of the Royal Asiatic Society 6: 25 - 37" title="A list of the land and fresh-water mollusca of the Malay Peninsula with notes." volume="6" year="1928">Laidlaw 1928</bibRefCitation>
;
<bibRefCitation author="Pall-Gergely, B" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="1" pagination="565 - 573" refId="B111" refString="Pall-Gergely, B, Hunyadi, A, Maassen, WJM, 2014. Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae). Journal of Conchology 41: 565 - 573" title="Review on Rhaphaulus L. Pfeiffer 1856 and Streptaulus Benson 1857 species with description of Rh. tonkinensis n. sp. from Vietnam (Gastropoda: Pupinidae)." volume="41" year="2014">
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
et al. 2014
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
It is possible that
<taxonomicName lsidName="R. chrysalis" pageId="0" pageNumber="1" rank="species" species="chrysalis">
<emphasis italics="true" pageId="0" pageNumber="1">R. chrysalis</emphasis>
</taxonomicName>
sensu
<bibRefCitation author="Habe, T" journalOrPublisher="Nature and Life in Southeast Asia" pageId="0" pageNumber="1" pagination="111 - 127" refId="B63" refString="Habe, T, 1965. Operculated land molluscs from Southeast Asia. Nature and Life in Southeast Asia 4: 111 - 127" title="Operculated land molluscs from Southeast Asia." volume="4" year="1965">Habe (1965)</bibRefCitation>
from Khao Chong, Trang Province, southern Thailand is
<taxonomicName lsidName="R. lorraini" pageId="0" pageNumber="1" rank="species" species="lorraini">
<emphasis italics="true" pageId="0" pageNumber="1">R. lorraini</emphasis>
</taxonomicName>
. This species is distributed in the Malay Peninsula and is disjunct from
<taxonomicName lsidName="R. chrysalis" pageId="0" pageNumber="1" rank="species" species="chrysalis">
<emphasis italics="true" pageId="0" pageNumber="1">R. chrysalis</emphasis>
</taxonomicName>
, which is distributed in northeastern India and Myanmar. See also remarks in
<taxonomicName lsidName="R. ascendens" pageId="0" pageNumber="1" rank="species" species="ascendens">
<emphasis italics="true" pageId="0" pageNumber="1">R. ascendens</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>