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<mods:title id="2F22E8F03CC40438E3695CC25193F83D">A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group</mods:title>
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<mods:affiliation id="6C72403C06166478FEC8AC7600463901">Laboratorio de Entomología, Unidad de Ecología y Sistemática-UNESIS, Departamento de Biología, Pontificia Universidad Javeriana, Bogotá, Colombia. E-mail: pachyurus @ yahoo. com</mods:affiliation>
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<mods:affiliation id="2B9B98A40DA44F639F799E85B9136C37">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, Colombia. E-mail: aeflorezd @ unal. edu. co</mods:affiliation>
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Key to the Colombian species of
<taxonomicName id="9AA14D0CB566C74BFD9BD99C8BD9F9D3" ID-CoL="62BPN" authorityName="Thorell" authorityYear="1891" box="[558,677,1515,1541]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="6FD5EA9DB566C74BFD9BD99C8BD9F9D3" bold="true" box="[558,677,1515,1541]" italics="true" pageId="35" pageNumber="36">Ananteris</emphasis>
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1. Metasomal segment IV with six complete carinae (paired ventrolateral, dorsolateral and dorsosubmedian carinae), and ventrosubmedian and median lateral carinae absent (
<bibRefCitation id="39304B7EB566C74BFD2BDA388A0AF9B2" author="Botero-Trujillo" box="[670,886,1615,1636]" firstAuthor="Botero-Trujillo" pageId="35" pageNumber="36" pagination="287 - 299" refId="ref30969" refString="Botero-Trujillo, R. (2008 a) The scorpion genus Ananteris in Colombia: comments on the taxonomy and description of two new species (Scorpiones, Buthidae). Journal of Arachnology, 36, 287 - 299." type="journal article" year="2008" yearSuffix="a">Botero-Trujillo 2008a</bibRefCitation>
: figs. 5, 14); chela trichobothria
<emphasis id="6FD5EA9DB566C74BFB04DA388DBAF9B1" box="[1201,1222,1615,1639]" italics="true" pageId="35" pageNumber="36">
V
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and
<emphasis id="6FD5EA9DB566C74BFB43DA388C77F9B1" box="[1270,1291,1615,1639]" italics="true" pageId="35" pageNumber="36">
V
<subScript id="C12534CAB566C74BFAB1DA2B8C77F9B1" attach="left" box="[1284,1291,1628,1639]" fontSize="5" pageId="35" pageNumber="36">2</subScript>
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unaligned axially, with
<emphasis id="6FD5EA9DB566C74BFEF5DA198829F951" box="[320,341,1646,1671]" italics="true" pageId="35" pageNumber="36">
V
<subScript id="C12534CAB566C74BFEFBDA0B8829F951" attach="left" box="[334,341,1660,1671]" fontSize="5" pageId="35" pageNumber="36">2</subScript>
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located on an external position in relation to
<emphasis id="6FD5EA9DB566C74BFCBDDA198A61F951" box="[776,797,1646,1671]" italics="true" pageId="35" pageNumber="36">
V
<subScript id="C12534CAB566C74BFCA3DA0B8A61F951" attach="left" box="[790,797,1660,1671]" fontSize="5" pageId="35" pageNumber="36">1</subScript>
</emphasis>
(
<figureCitation id="C59A2A0AB566C74BFC9FDA198D75F955" box="[810,1033,1646,1667]" captionStart="FIGURES 4047" captionStartId="32.[151,263,1802,1825]" captionTargetBox="[151,1411,227,1748]" captionTargetId="figure-101@32.[151,1436,193,1781]" captionTargetPageId="32" captionText="FIGURES 4047. Right pedipalp hand, distribution of ventral trichobothria. 40. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 41. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 42. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 43. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). 44. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 45. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 46. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 47. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284810" httpUri="https://zenodo.org/record/5284810/files/figure.png" pageId="35" pageNumber="36">Figs. 40, 42, 4445, 47</figureCitation>
)................................ 2
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1 Metasomal segment IV with eight complete carinae (only median lateral carinae absent); chela trichobothria
<emphasis id="6FD5EA9DB566C74BFB43DAF98C77F970" box="[1270,1291,1678,1702]" italics="true" pageId="35" pageNumber="36">
V
<subScript id="C12534CAB566C74BFAB1DAEC8C77F970" attach="left" box="[1284,1291,1691,1702]" fontSize="5" pageId="35" pageNumber="36">1</subScript>
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and
<emphasis id="6FD5EA9DB566C74BFA8FDAF98C2CF970" box="[1338,1360,1678,1702]" italics="true" pageId="35" pageNumber="36">
V
<subScript id="C12534CAB566C74BFAFCDAEC8C2CF970" attach="left" box="[1353,1360,1691,1702]" fontSize="5" pageId="35" pageNumber="36">2</subScript>
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aligned axially (
<figureCitation id="C59A2A0AB566C74BFE9ADAD988DEF915" box="[303,418,1710,1731]" captionStart="FIGURES 4047" captionStartId="32.[151,263,1802,1825]" captionTargetBox="[151,1411,227,1748]" captionTargetId="figure-101@32.[151,1436,193,1781]" captionTargetPageId="32" captionText="FIGURES 4047. Right pedipalp hand, distribution of ventral trichobothria. 40. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 41. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 42. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 43. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). 44. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 45. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 46. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 47. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284810" httpUri="https://zenodo.org/record/5284810/files/figure.png" pageId="35" pageNumber="36">Figs. 41, 43</figureCitation>
) (unaligned in
<taxonomicName id="9AA14D0CB566C74BFD81DAD98BC8F915" ID-CoL="66LCV" authorityName="Botero-Trujillo" authorityYear="2009" box="[564,692,1710,1731]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="volschenki">
<emphasis id="6FD5EA9DB566C74BFD81DAD98BC8F915" box="[564,692,1710,1731]" italics="true" pageId="35" pageNumber="36">A. volschenki</emphasis>
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and slightly unaligned in
<taxonomicName id="9AA14D0CB566C74BFC1BDAD98D75F915" ID-CoL="66LC5" authority="Botero-Trujillo &amp; Flórez, 2011" authorityName="Botero-Trujillo &amp; Flórez" authorityYear="2011" box="[942,1033,1710,1731]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ochoai" status="sp. nov.">
<emphasis id="6FD5EA9DB566C74BFC1BDAD98D75F915" box="[942,1033,1710,1731]" italics="true" pageId="35" pageNumber="36">A. ochoai</emphasis>
</taxonomicName>
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<taxonomicNameLabel id="74E657E6B566C74BFBA5DAD98D21F915" box="[1040,1117,1710,1731]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
)
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......................... 6
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2. Metasomal carinal formula10:10:10:6:3, with ventrolateral carinae absent on segment V (
<bibRefCitation id="39304B7EB566C74BFB8CDABC8C6CF936" author="Botero-Trujillo" box="[1081,1296,1739,1760]" firstAuthor="Botero-Trujillo" pageId="35" pageNumber="36" pagination="287 - 299" refId="ref30969" refString="Botero-Trujillo, R. (2008 a) The scorpion genus Ananteris in Colombia: comments on the taxonomy and description of two new species (Scorpiones, Buthidae). Journal of Arachnology, 36, 287 - 299." type="journal article" year="2008" yearSuffix="a">Botero-Trujillo 2008a</bibRefCitation>
: fig. 15); carapace slightly concave anteriorly and lacking median projection (
<figureCitation id="C59A2A0AB566C74BFC22DA9F8A95F92B" box="[919,1001,1768,1789]" captionStart="FIGURES 3235" captionStartId="29.[151,263,1283,1306]" captionTargetBox="[266,1321,199,1252]" captionTargetId="figure-334@29.[264,1322,193,1262]" captionTargetPageId="29" captionText="FIGURES 3235. Anterior margin of carapace and chelicerae. 32. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 33. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100779). 34. Ananteris dorae Botero-Trujillo, 2008, adult male (MPUJ-SCO-385). 35. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814)." figureDoi="http://doi.org/10.5281/zenodo.5284804" httpUri="https://zenodo.org/record/5284804/files/figure.png" pageId="35" pageNumber="36">Fig. 34</figureCitation>
); fixed finger trichobothria in the order
<emphasis id="6FD5EA9DB566C74BFF54DB718984F8CD" box="[225,248,1798,1819]" italics="true" pageId="35" pageNumber="36">eb</emphasis>
:
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:
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:
<emphasis id="6FD5EA9DB566C74BFEF6DB718822F8CD" box="[323,350,1798,1819]" italics="true" pageId="35" pageNumber="36">est</emphasis>
:
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:
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, with
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distinctly basal to
<emphasis id="6FD5EA9DB566C74BFD2FDB718BC9F8CD" box="[666,693,1798,1819]" italics="true" pageId="35" pageNumber="36">est</emphasis>
(
<figureCitation id="C59A2A0AB566C74BFD74DB718A75F8CD" box="[705,777,1798,1819]" captionStart="FIGURES 4851" captionStartId="33.[151,263,1857,1880]" captionTargetBox="[180,1406,245,1790]" captionTargetId="figure-68@33.[151,1436,193,1836]" captionTargetPageId="33" captionText="FIGURES 4851. Right pedipalp chela, distribution of external and dorsal trichobothria. 48. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 49. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 50. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 51. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284814" httpUri="https://zenodo.org/record/5284814/files/figure.png" pageId="35" pageNumber="36">Fig. 50</figureCitation>
); pectines with 1214 teeth; hemispermatophore with three lobes, of which the basal is armed with three minute spines (
<figureCitation id="C59A2A0AB566C74BFD79DB548A6EF8EE" box="[716,786,1827,1848]" captionStart="FIGURES 6063" captionStartId="38.[151,263,1792,1815]" captionTargetBox="[184,1410,214,1744]" captionTargetId="figure-98@38.[151,1436,193,1771]" captionTargetPageId="38" captionText="FIGURES 6063. Hemispermatophore in ectal aspect and detail of the capsular region in ventral aspect (left for Ananteris dorae Botero-Trujillo, 2008, inverted; right for others). 60. Ananteris arcadioi Botero-Trujillo, 2008 (ICN-As-774). 61. Ananteris columbiana Lourenço, 1991 (ICN-As-446). 62. Ananteris dorae Botero-Trujillo, 2008 (ICN-As-816). 63. Ananteris ehrlichi Lourenço, 1994 (IAvH-E 100815). Scale bars = 0.5 mm (0.25 mm for A. dorae). Abbreviations: il = internal lobe, el = external lobe, ml = median lobe, bl = basal lobe." figureDoi="http://doi.org/10.5281/zenodo.5284828" httpUri="https://zenodo.org/record/5284828/files/figure.png" pageId="35" pageNumber="36">Fig. 62</figureCitation>
)..............................................
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<emphasis id="6FD5EA9DB566C74BFAFCDB548CE7F8EE" box="[1353,1435,1827,1848]" italics="true" pageId="35" pageNumber="36">A. dorae</emphasis>
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2 Metasomal carinal formula 10:10:10:6:5, with ventrolateral carinae present on segment V; carapace with well-developed anteromedian projection (
<figureCitation id="C59A2A0AB566C74BFE12DB29888AF8A5" box="[423,502,1886,1907]" captionStart="FIGURES 3235" captionStartId="29.[151,263,1283,1306]" captionTargetBox="[266,1321,199,1252]" captionTargetId="figure-334@29.[264,1322,193,1262]" captionTargetPageId="29" captionText="FIGURES 3235. Anterior margin of carapace and chelicerae. 32. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 33. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100779). 34. Ananteris dorae Botero-Trujillo, 2008, adult male (MPUJ-SCO-385). 35. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814)." figureDoi="http://doi.org/10.5281/zenodo.5284804" httpUri="https://zenodo.org/record/5284804/files/figure.png" pageId="35" pageNumber="36">Figs. 32</figureCitation>
,
<figureCitation id="C59A2A0AB566C74BFDB5DB298B1EF8A5" box="[512,610,1886,1907]" captionStart="FIGURES 3639" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 3639. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="35" pageNumber="36">3637, 39</figureCitation>
); fixed finger trichobothria in the order
<emphasis id="6FD5EA9DB566C74BFC69DB298A8EF8A5" box="[988,1010,1886,1907]" italics="true" pageId="35" pageNumber="36">eb</emphasis>
:
<emphasis id="6FD5EA9DB566C74BFC4FDB298D66F8A5" box="[1018,1050,1886,1907]" italics="true" pageId="35" pageNumber="36">esb</emphasis>
:
<emphasis id="6FD5EA9DB566C74BFB95DB298D47F8A5" box="[1056,1083,1886,1907]" italics="true" pageId="35" pageNumber="36">est</emphasis>
/
<emphasis id="6FD5EA9DB566C74BFBF7DB298D26F8A5" box="[1090,1114,1886,1907]" italics="true" pageId="35" pageNumber="36">db</emphasis>
:
<emphasis id="6FD5EA9DB566C74BFBD5DB298D0DF8A5" box="[1120,1137,1886,1907]" italics="true" pageId="35" pageNumber="36">et</emphasis>
:
<emphasis id="6FD5EA9DB566C74BFBCDDB298DF7F8A5" box="[1144,1163,1886,1907]" italics="true" pageId="35" pageNumber="36">dt</emphasis>
, with
<emphasis id="6FD5EA9DB566C74BFB73DB298D9DF8A5" box="[1222,1249,1886,1907]" italics="true" pageId="35" pageNumber="36">est</emphasis>
beside
<emphasis id="6FD5EA9DB566C74BFA9CDB298C3DF8A5" box="[1321,1345,1886,1907]" italics="true" pageId="35" pageNumber="36">db</emphasis>
or nearly so (
<figureCitation id="C59A2A0AB566C74BFEB6DB0C882FF846" box="[259,339,1915,1936]" captionStart="FIGURES 4851" captionStartId="33.[151,263,1857,1880]" captionTargetBox="[180,1406,245,1790]" captionTargetId="figure-68@33.[151,1436,193,1836]" captionTargetPageId="33" captionText="FIGURES 4851. Right pedipalp chela, distribution of external and dorsal trichobothria. 48. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 49. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 50. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 51. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284814" httpUri="https://zenodo.org/record/5284814/files/figure.png" pageId="35" pageNumber="36">Figs. 48</figureCitation>
,
<figureCitation id="C59A2A0AB566C74BFEE8DB0C88BCF846" box="[349,448,1915,1936]" captionStart="FIGURES 5255" captionStartId="34.[151,263,1859,1882]" captionTargetBox="[179,1408,232,1790]" captionTargetId="figure-71@34.[151,1436,193,1838]" captionTargetPageId="34" captionText="FIGURES 5255. Right pedipalp chela, distribution of external and dorsal trichobothria. 52. Ananteris meridana González- Sponga, 2006, adult female (ICN-As-321). 53. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 54. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 55. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO- 390). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284818" httpUri="https://zenodo.org/record/5284818/files/figure.png" pageId="35" pageNumber="36">5253, 55</figureCitation>
); pectines with 15 or more teeth; hemispermatophore with three lobes lacking any spines (
<figureCitation id="C59A2A0AB566C74BFA96DB0C8C0FF846" box="[1315,1395,1915,1936]" captionStart="FIGURES 6063" captionStartId="38.[151,263,1792,1815]" captionTargetBox="[184,1410,214,1744]" captionTargetId="figure-98@38.[151,1436,193,1771]" captionTargetPageId="38" captionText="FIGURES 6063. Hemispermatophore in ectal aspect and detail of the capsular region in ventral aspect (left for Ananteris dorae Botero-Trujillo, 2008, inverted; right for others). 60. Ananteris arcadioi Botero-Trujillo, 2008 (ICN-As-774). 61. Ananteris columbiana Lourenço, 1991 (ICN-As-446). 62. Ananteris dorae Botero-Trujillo, 2008 (ICN-As-816). 63. Ananteris ehrlichi Lourenço, 1994 (IAvH-E 100815). Scale bars = 0.5 mm (0.25 mm for A. dorae). Abbreviations: il = internal lobe, el = external lobe, ml = median lobe, bl = basal lobe." figureDoi="http://doi.org/10.5281/zenodo.5284828" httpUri="https://zenodo.org/record/5284828/files/figure.png" pageId="35" pageNumber="36">Figs. 60</figureCitation>
,
<figureCitation id="C59A2A0AB566C74BFAC8DB0C8862F87B" captionStart="FIGURES 6467" captionStartId="39.[151,264,1792,1815]" captionTargetBox="[199,1415,211,1744]" captionTargetId="figure-110@39.[151,1436,193,1771]" captionTargetPageId="39" captionText="FIGURES 6467. Hemispermatophore in ectal aspect and detail of the capsular region in ventral aspect (left for Ananteris solimariae sp. nov., inverted; right for others). 64. Ananteris meridana González-Sponga, 2006 (ICN-As-435). 65. Ananteris myriamae Botero-Trujillo, 2007 (MPUJ-SCO-382). 66. Ananteris ochoai sp. nov., paratype (ICN-As-815). 67. Ananteris solimariae sp. nov., paratype (ICN-As-779). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284832" httpUri="https://zenodo.org/record/5284832/files/figure.png" pageId="35" pageNumber="36">64, 65, 67</figureCitation>
).............................................................................................. 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB566C74BFF22DBC18CE0F83E" pageId="35" pageNumber="36">
<paragraph id="5D1E368FB566C74BFF22DBC18CE0F81D" blockId="35.[151,1436,1585,2024]" box="[151,1436,1974,1995]" pageId="35" pageNumber="36">
<keyLead id="E65095BAB566C74BFF22DBC18CE0F81D" box="[151,1436,1974,1995]" pageId="35" pageNumber="36">
3. Dorsal surface of chelicerae predominantly yellow, with brown color restricted to the anterior region (
<figureCitation id="C59A2A0AB566C74BFB04DBC18C58F81D" box="[1201,1316,1974,1995]" captionStart="FIGURES 3639" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 3639. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="35" pageNumber="36">Figs. 36, 39</figureCitation>
)........4
</keyLead>
</paragraph>
<paragraph id="5D1E368FB566C74BFF22DBA48CE0F83E" blockId="35.[151,1436,1585,2024]" box="[151,1436,2003,2024]" pageId="35" pageNumber="36">
<keyLead id="E65095BAB566C74BFF22DBA48CE0F83E" box="[151,1436,2003,2024]" pageId="35" pageNumber="36">
3 Dorsal surface of chelicerae with reticulations (
<figureCitation id="C59A2A0AB566C74BFD12DBA48B84F83E" box="[679,760,2003,2024]" captionStart="FIGURES 3235" captionStartId="29.[151,263,1283,1306]" captionTargetBox="[266,1321,199,1252]" captionTargetId="figure-334@29.[264,1322,193,1262]" captionTargetPageId="29" captionText="FIGURES 3235. Anterior margin of carapace and chelicerae. 32. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 33. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100779). 34. Ananteris dorae Botero-Trujillo, 2008, adult male (MPUJ-SCO-385). 35. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814)." figureDoi="http://doi.org/10.5281/zenodo.5284804" httpUri="https://zenodo.org/record/5284804/files/figure.png" pageId="35" pageNumber="36">Figs. 32</figureCitation>
,
<figureCitation id="C59A2A0AB566C74BFCB7DBA48A62F83E" box="[770,798,2003,2024]" captionStart="FIGURES 3639" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 3639. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="35" pageNumber="36">37</figureCitation>
).................................................... 5
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</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DCE08CE0FEE9" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DCE08CE0FF31" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DCE08CE0FF31" pageId="36" pageNumber="37">
4. Ventrosubmedian carinae on metasomal segment III incomplete, only present in the basal half; pectines with 1719 teeth on males and females; coxosternal region with diffuse brownish pigmentation more conspicuous on coxapophyses I and coxae I II...........................................................................................
<taxonomicName id="9AA14D0CB561C74CFAF5DCA58CE0FF31" ID-CoL="66LBP" authorityName="Gonzalez-Sponga" authorityYear="2006" box="[1344,1436,210,231]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="meridana">
<emphasis id="6FD5EA9DB561C74CFAF5DCA58CE0FF31" box="[1344,1436,210,231]" italics="true" pageId="36" pageNumber="37">meridana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22DC988CE0FEE9" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DC988CE0FEE9" pageId="36" pageNumber="37">
4 Ventrosubmedian carinae on metasomal segment III complete; pectines with 1921 teeth on males and 1820 on females; coxosternal region almost entirely yellow, with only some brownish areas restricted to the anterior margin of coxae............................................................................................
<taxonomicName id="9AA14D0CB561C74CFB7DDD5D8C35FEE9" ID-CoL="66LCL" authority="Botero-Trujillo &amp; Flórez, 2011" authorityName="Botero-Trujillo &amp; Flórez" authorityYear="2011" box="[1224,1353,298,319]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="solimariae" status="sp. nov.">
<emphasis id="6FD5EA9DB561C74CFB7DDD5D8C35FEE9" box="[1224,1353,298,319]" italics="true" pageId="36" pageNumber="37">A. solimariae</emphasis>
</taxonomicName>
<emphasis id="6FD5EA9DB561C74CFAFADD5D8CE0FEE9" bold="true" box="[1359,1436,298,319]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="74E657E6B561C74CFAFADD5D8CE0FEE9" box="[1359,1436,298,319]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DD308CE7FE39" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DD308CE0FE41" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DD308CE0FE41" pageId="36" pageNumber="37">
5. Dorsal surface of chelicerae with dense and complete reticular pattern (
<figureCitation id="C59A2A0AB561C74CFC39DD308AA8FE8A" box="[908,980,327,348]" captionStart="FIGURES 3639" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 3639. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="36" pageNumber="37">Fig. 37</figureCitation>
); pedipalp hand uniformly yellow and spotless; coxosternal region with conspicuous brown areas (Botero-Trujillo 2007: fig. 4); pectines with 17 teeth on males and 1516 on females.....................................................................................
<taxonomicName id="9AA14D0CB561C74CFAABDDF58CE0FE41" ID-CoL="66LC2" authorityName="Botero-Trujillo" authorityYear="2007" box="[1310,1436,386,407]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="myriamae">
<emphasis id="6FD5EA9DB561C74CFAABDDF58CE0FE41" box="[1310,1436,386,407]" italics="true" pageId="36" pageNumber="37">A. myriamae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22DDE88CE7FE39" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DDE88CE7FE39" pageId="36" pageNumber="37">
5 Dorsal surface of chelicerae with incomplete reticular pattern (
<figureCitation id="C59A2A0AB561C74CFCF7DDE88AF7FE62" box="[834,907,415,436]" captionStart="FIGURES 3235" captionStartId="29.[151,263,1283,1306]" captionTargetBox="[266,1321,199,1252]" captionTargetId="figure-334@29.[264,1322,193,1262]" captionTargetPageId="29" captionText="FIGURES 3235. Anterior margin of carapace and chelicerae. 32. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 33. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100779). 34. Ananteris dorae Botero-Trujillo, 2008, adult male (MPUJ-SCO-385). 35. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814)." figureDoi="http://doi.org/10.5281/zenodo.5284804" httpUri="https://zenodo.org/record/5284804/files/figure.png" pageId="36" pageNumber="37">Fig. 32</figureCitation>
); pedipalp hand most usually with conspicuous brown areas, infrequently completely yellow; coxosternal region entirely yellow and spotless; pectines with 1721 teeth on males and 1618 on females.............................................................................
<taxonomicName id="9AA14D0CB561C74CFA84DDAD8CE7FE39" ID-CoL="66LJC" authorityName="Botero-Trujillo" authorityYear="2008" box="[1329,1435,474,495]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="arcadioi">
<emphasis id="6FD5EA9DB561C74CFA84DDAD8CE7FE39" box="[1329,1435,474,495]" italics="true" pageId="36" pageNumber="37">A. arcadioi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DD808CE0FD51" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DD808CE0FD9C" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DD808CE0FD9C" pageId="36" pageNumber="37">
6. Chela trichobothria
<emphasis id="6FD5EA9DB561C74CFE17DD8F88CBFDC6" box="[418,439,504,528]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFE05DE7288CBFDC6" attach="left" box="[432,439,517,528]" fontSize="5" pageId="36" pageNumber="37">1</subScript>
</emphasis>
and
<emphasis id="6FD5EA9DB561C74CFE52DD8F8880FDC6" box="[487,508,504,528]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFE40DE728880FDC6" attach="left" box="[501,508,517,528]" fontSize="5" pageId="36" pageNumber="37">2</subScript>
</emphasis>
unaligned axially, with
<emphasis id="6FD5EA9DB561C74CFD50DD8F8B87FDC6" box="[741,763,504,528]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFD41DE728B87FDC6" attach="left" box="[756,763,517,528]" fontSize="5" pageId="36" pageNumber="37">2</subScript>
</emphasis>
located on an external position in relation to
<emphasis id="6FD5EA9DB561C74CFB06DD8F8DB5FDC6" box="[1203,1225,504,528]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFB77DE728DB5FDC6" attach="left" box="[1218,1225,517,528]" fontSize="5" pageId="36" pageNumber="37">1</subScript>
</emphasis>
(
<figureCitation id="C59A2A0AB561C74CFB63DD808C62FDDA" box="[1238,1310,503,524]" captionStart="FIGURES 4047" captionStartId="32.[151,263,1802,1825]" captionTargetBox="[151,1411,227,1748]" captionTargetId="figure-101@32.[151,1436,193,1781]" captionTargetPageId="32" captionText="FIGURES 4047. Right pedipalp hand, distribution of ventral trichobothria. 40. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 41. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 42. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 43. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). 44. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 45. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 46. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 47. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284810" httpUri="https://zenodo.org/record/5284810/files/figure.png" pageId="36" pageNumber="37">Fig. 46</figureCitation>
; Botero-Trujillo 2009: fig. 25); fixed finger trichobothria in the order
<emphasis id="6FD5EA9DB561C74CFCBCDE608A5CFDFA" box="[777,800,535,556]" italics="true" pageId="36" pageNumber="37">eb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFC92DE608A3BFDFA" box="[807,839,535,556]" italics="true" pageId="36" pageNumber="37">esb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFCF8DE608A14FDFA" box="[845,872,535,556]" italics="true" pageId="36" pageNumber="37">est</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFCDADE608AFBFDFA" box="[879,903,535,556]" italics="true" pageId="36" pageNumber="37">db</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFC38DE608AE2FDFA" box="[909,926,535,556]" italics="true" pageId="36" pageNumber="37">et</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFC10DE608AC4FDFA" box="[933,952,535,556]" italics="true" pageId="36" pageNumber="37">dt</emphasis>
, with
<emphasis id="6FD5EA9DB561C74CFC41DE608D73FDFA" box="[1012,1039,535,556]" italics="true" pageId="36" pageNumber="37">est</emphasis>
notably basal to
<emphasis id="6FD5EA9DB561C74CFB06DE608DB7FDFA" box="[1203,1227,535,556]" italics="true" pageId="36" pageNumber="37">db</emphasis>
(
<figureCitation id="C59A2A0AB561C74CFB62DE608C62FDFA" box="[1239,1310,535,556]" captionStart="FIGURES 5255" captionStartId="34.[151,263,1859,1882]" captionTargetBox="[179,1408,232,1790]" captionTargetId="figure-71@34.[151,1436,193,1838]" captionTargetPageId="34" captionText="FIGURES 5255. Right pedipalp chela, distribution of external and dorsal trichobothria. 52. Ananteris meridana González- Sponga, 2006, adult female (ICN-As-321). 53. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 54. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 55. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO- 390). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284818" httpUri="https://zenodo.org/record/5284818/files/figure.png" pageId="36" pageNumber="37">Fig. 54</figureCitation>
; Botero-Trujillo 2009: fig. 24)...................................................................................... 7
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22DE258CE0FD51" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DE258CE0FD51" pageId="36" pageNumber="37">
6 Chela trichobothria
<emphasis id="6FD5EA9DB561C74CFE16DE2588C4FDBC" box="[419,440,594,618]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFE04DE2888C4FDBC" attach="left" box="[433,440,607,618]" fontSize="5" pageId="36" pageNumber="37">1</subScript>
</emphasis>
and
<emphasis id="6FD5EA9DB561C74CFE5CDE258882FDBC" box="[489,510,594,618]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFE42DE288882FDBC" attach="left" box="[503,510,607,618]" fontSize="5" pageId="36" pageNumber="37">2</subScript>
</emphasis>
aligned axially (
<figureCitation id="C59A2A0AB561C74CFD17DE258A66FDB1" box="[674,794,594,615]" captionStart="FIGURES 4047" captionStartId="32.[151,263,1802,1825]" captionTargetBox="[151,1411,227,1748]" captionTargetId="figure-101@32.[151,1436,193,1781]" captionTargetPageId="32" captionText="FIGURES 4047. Right pedipalp hand, distribution of ventral trichobothria. 40. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 41. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 42. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 43. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). 44. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 45. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 46. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 47. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284810" httpUri="https://zenodo.org/record/5284810/files/figure.png" pageId="36" pageNumber="37">Figs. 41, 43</figureCitation>
); fixed finger trichobothria either
<emphasis id="6FD5EA9DB561C74CFBD3DE258D00FDB1" box="[1126,1148,594,615]" italics="true" pageId="36" pageNumber="37">eb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB36DE258DDFFDB1" box="[1155,1187,594,615]" italics="true" pageId="36" pageNumber="37">esb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB1FDE258DBEFDB1" box="[1194,1218,594,615]" italics="true" pageId="36" pageNumber="37">db</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB7CDE258D98FDB1" box="[1225,1252,594,615]" italics="true" pageId="36" pageNumber="37">est</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB5FDE258D87FDB1" box="[1258,1275,594,615]" italics="true" pageId="36" pageNumber="37">et</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFAB7DE258C69FDB1" box="[1282,1301,594,615]" italics="true" pageId="36" pageNumber="37">dt</emphasis>
or
<emphasis id="6FD5EA9DB561C74CFA82DE258C31FDB1" box="[1335,1357,594,615]" italics="true" pageId="36" pageNumber="37">eb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFAE1DE258C08FDB1" box="[1364,1396,594,615]" italics="true" pageId="36" pageNumber="37">esb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFACEDE258CEAFDB1" box="[1403,1430,594,615]" italics="true" pageId="36" pageNumber="37">est</emphasis>
/
<emphasis id="6FD5EA9DB561C74CFF54DE058985FD51" box="[225,249,626,647]" italics="true" pageId="36" pageNumber="37">db</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFF4ADE05886CFD51" box="[255,272,626,647]" italics="true" pageId="36" pageNumber="37">et</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFEA2DE058856FD51" box="[279,298,626,647]" italics="true" pageId="36" pageNumber="37">dt</emphasis>
..............................................................................................8
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DEF88CE0FD28" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DEF88CE7FD17" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DEF88CE7FD17" pageId="36" pageNumber="37">
7. Pectines with 15 teeth on males and 1516 on females; metasomal carinal formula 10:8:8:8:5, with segments IIIV lacking median lateral carinae; chela trichobothria
<emphasis id="6FD5EA9DB561C74CFDC9DEDB8BEDFD13" box="[636,657,684,709]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFD3FDECD8BEDFD13" attach="left" box="[650,657,698,709]" fontSize="5" pageId="36" pageNumber="37">1</subScript>
</emphasis>
and
<emphasis id="6FD5EA9DB561C74CFD0ADEDB8BA9FD13" box="[703,725,684,709]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFD7BDECD8BA9FD13" attach="left" box="[718,725,698,709]" fontSize="5" pageId="36" pageNumber="37">2</subScript>
</emphasis>
markedly unaligned axially (Botero-Trujillo 2009: fig. 25) ..
<taxonomicName id="9AA14D0CB561C74CFAA9DEDB8CE7FD17" ID-CoL="66LCV" authorityName="Botero-Trujillo" authorityYear="2009" box="[1308,1435,684,705]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="volschenki">
<emphasis id="6FD5EA9DB561C74CFAA9DEDB8CE7FD17" box="[1308,1435,684,705]" italics="true" pageId="36" pageNumber="37">A. volschenki</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22DEBB8CE0FD28" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DEBB8CE0FD28" pageId="36" pageNumber="37">
7 Pectines with 2023 teeth on males and 2223 on females; metasomal carinal formula 10:10:10:8:5, with only segment IV lacking median lateral carinae; chela trichobothria
<emphasis id="6FD5EA9DB561C74CFD7DDE9D8BA2FCD4" box="[712,734,746,770]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFD62DE808BA2FCD4" attach="left" box="[727,734,759,770]" fontSize="5" pageId="36" pageNumber="37">1</subScript>
</emphasis>
and
<emphasis id="6FD5EA9DB561C74CFCB9DE9D8A5EFCD4" box="[780,802,746,770]" italics="true" pageId="36" pageNumber="37">
V
<subScript id="C12534CAB561C74CFCAEDE808A5EFCD4" attach="left" box="[795,802,759,770]" fontSize="5" pageId="36" pageNumber="37">2</subScript>
</emphasis>
only slightly unaligned axially (
<figureCitation id="C59A2A0AB561C74CFBEDDE9E8DE2FD28" box="[1112,1182,745,766]" captionStart="FIGURES 4047" captionStartId="32.[151,263,1802,1825]" captionTargetBox="[151,1411,227,1748]" captionTargetId="figure-101@32.[151,1436,193,1781]" captionTargetPageId="32" captionText="FIGURES 4047. Right pedipalp hand, distribution of ventral trichobothria. 40. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 41. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 42. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 43. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). 44. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 45. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 46. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 47. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284810" httpUri="https://zenodo.org/record/5284810/files/figure.png" pageId="36" pageNumber="37">Fig. 46</figureCitation>
).....
<taxonomicName id="9AA14D0CB561C74CFB58DE9D8C35FD29" ID-CoL="66LC5" authority="Botero-Trujillo &amp; Flórez, 2011" authorityName="Botero-Trujillo &amp; Flórez" authorityYear="2011" box="[1261,1353,746,767]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ochoai" status="sp. nov.">
<emphasis id="6FD5EA9DB561C74CFB58DE9D8C35FD29" box="[1261,1353,746,767]" italics="true" pageId="36" pageNumber="37">A. ochoai</emphasis>
</taxonomicName>
<emphasis id="6FD5EA9DB561C74CFAFADE9E8CE0FD28" bold="true" box="[1359,1436,745,766]" pageId="36" pageNumber="37">
<taxonomicNameLabel id="74E657E6B561C74CFAFADE9E8CE0FD28" box="[1359,1436,745,766]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DF7E8CE7FCEA" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DF7E8CE0FCC8" blockId="36.[151,1437,151,1180]" box="[151,1436,777,798]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DF7E8CE0FCC8" box="[151,1436,777,798]" pageId="36" pageNumber="37">8. Metasomal carinal formula 10:10:10:8:5, with median lateral carinae on segment III only present anteriorly.............. 9</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22DF508CE7FCEA" blockId="36.[151,1437,151,1180]" box="[151,1435,807,828]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DF508CE7FCEA" box="[151,1435,807,828]" pageId="36" pageNumber="37">8 Metasomal carinal formula 10:10:8:8:5, with median lateral carinae absent on segment III........................... 11</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DF338CE7FC67" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DF338CE7FCA0" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DF338CE7FCA0" pageId="36" pageNumber="37">
9. Fixed finger trichobothria in the order
<emphasis id="6FD5EA9DB561C74CFDEBDF338B09FC8F" box="[606,629,836,857]" italics="true" pageId="36" pageNumber="37">eb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFDC9DF338BE0FC8F" box="[636,668,836,857]" italics="true" pageId="36" pageNumber="37">esb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFD17DF338BC1FC8F" box="[674,701,836,857]" italics="true" pageId="36" pageNumber="37">est</emphasis>
/
<emphasis id="6FD5EA9DB561C74CFD71DF338BA0FC8F" box="[708,732,836,857]" italics="true" pageId="36" pageNumber="37">db</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFD57DF338B8FFC8F" box="[738,755,836,857]" italics="true" pageId="36" pageNumber="37">et</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFD4FDF338A71FC8F" box="[762,781,836,857]" italics="true" pageId="36" pageNumber="37">dt</emphasis>
, with
<emphasis id="6FD5EA9DB561C74CFCF8DF338A14FC8F" box="[845,872,836,857]" italics="true" pageId="36" pageNumber="37">est</emphasis>
located beside
<emphasis id="6FD5EA9DB561C74CFBB7DF338D66FC8F" box="[1026,1050,836,857]" italics="true" pageId="36" pageNumber="37">db</emphasis>
or nearly so; pectines with 21 or more teeth............................................................................................... 10
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22DF088CE7FC67" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DF088CE7FC67" pageId="36" pageNumber="37">
9 Fixed finger trichobothria in the order
<emphasis id="6FD5EA9DB561C74CFDE7DF088B15FC42" box="[594,617,895,916]" italics="true" pageId="36" pageNumber="37">eb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFDDADF088BF3FC42" box="[623,655,895,916]" italics="true" pageId="36" pageNumber="37">esb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFD20DF088BD1FC42" box="[661,685,895,916]" italics="true" pageId="36" pageNumber="37">db</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFD01DF088BB3FC42" box="[692,719,895,916]" italics="true" pageId="36" pageNumber="37">est</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFD60DF088B9AFC42" box="[725,742,895,916]" italics="true" pageId="36" pageNumber="37">et</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFD58DF088A7CFC42" box="[749,768,895,916]" italics="true" pageId="36" pageNumber="37">dt</emphasis>
, with
<emphasis id="6FD5EA9DB561C74CFC89DF088A28FC42" box="[828,852,895,916]" italics="true" pageId="36" pageNumber="37">db</emphasis>
distinctly basal to
<emphasis id="6FD5EA9DB561C74CFBBCDF088D58FC42" box="[1033,1060,895,916]" italics="true" pageId="36" pageNumber="37">est</emphasis>
(
<figureCitation id="C59A2A0AB561C74CFB85DF088D04FC42" box="[1072,1144,895,916]" captionStart="FIGURES 4851" captionStartId="33.[151,263,1857,1880]" captionTargetBox="[180,1406,245,1790]" captionTargetId="figure-68@33.[151,1436,193,1836]" captionTargetPageId="33" captionText="FIGURES 4851. Right pedipalp chela, distribution of external and dorsal trichobothria. 48. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 49. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 50. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 51. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284814" httpUri="https://zenodo.org/record/5284814/files/figure.png" pageId="36" pageNumber="37">Fig. 49</figureCitation>
); pectines with 1620 teeth on males and 1519 on females..................................................................
<taxonomicName id="9AA14D0CB561C74CFAA5DFEB8CE7FC67" ID-CoL="66LL8" authorityName="Lourenco" authorityYear="1991" box="[1296,1435,924,945]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="columbiana">
<emphasis id="6FD5EA9DB561C74CFAA5DFEB8CE7FC67" box="[1296,1435,924,945]" italics="true" pageId="36" pageNumber="37">A. columbiana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DFCE8CE7FC3A" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DFCE8CE0FC18" blockId="36.[151,1437,151,1180]" box="[151,1436,953,974]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DFCE8CE0FC18" box="[151,1436,953,974]" pageId="36" pageNumber="37">
10. Ocular carinae smooth; pectines with 2124 teeth on males (females unknown)...........................
<taxonomicName id="9AA14D0CB561C74CFA93DFCE8CE0FC18" ID-CoL="66KZS" authorityName="Lourenco &amp; Florez" authorityYear="1989" box="[1318,1436,953,974]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="gorgonae">
<emphasis id="6FD5EA9DB561C74CFA93DFCE8CE0FC18" box="[1318,1436,953,974]" italics="true" pageId="36" pageNumber="37">A. gorgonae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22DFA08CE7FC3A" blockId="36.[151,1437,151,1180]" box="[151,1435,983,1004]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DFA08CE7FC3A" box="[151,1435,983,1004]" pageId="36" pageNumber="37">
10 Ocular carinae granulose; pectines with 2223 teeth on males and 2122 on females........................
<taxonomicName id="9AA14D0CB561C74CFA99DFA08CE7FC3A" ID-CoL="66LCR" authorityName="Teruel &amp; Garcia" authorityYear="2007" box="[1324,1435,983,1004]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="tolimana">
<emphasis id="6FD5EA9DB561C74CFA99DFA08CE7FC3A" box="[1324,1435,983,1004]" italics="true" pageId="36" pageNumber="37">A. tolimana</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22DF838CE7FBB7" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22DF838CE0FBF0" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22DF838CE0FBF0" pageId="36" pageNumber="37">
11. Pedipalp hand with conspicuous brown areas; fixed finger trichobothria in the order
<emphasis id="6FD5EA9DB561C74CFBBDDF838D63FBDF" box="[1032,1055,1012,1033]" italics="true" pageId="36" pageNumber="37">eb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB93DF838D3AFBDF" box="[1062,1094,1012,1033]" italics="true" pageId="36" pageNumber="37">esb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFBF9DF838D1BFBDF" box="[1100,1127,1012,1033]" italics="true" pageId="36" pageNumber="37">est</emphasis>
/
<emphasis id="6FD5EA9DB561C74CFBD8DF838DF9FBDF" box="[1133,1157,1012,1033]" italics="true" pageId="36" pageNumber="37">db</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB39DF838DE1FBDF" box="[1164,1181,1012,1033]" italics="true" pageId="36" pageNumber="37">et</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB11DF838DCBFBDF" box="[1188,1207,1012,1033]" italics="true" pageId="36" pageNumber="37">dt</emphasis>
, with
<emphasis id="6FD5EA9DB561C74CFB46DF838C72FBDF" box="[1267,1294,1012,1033]" italics="true" pageId="36" pageNumber="37">est</emphasis>
located beside
<emphasis id="6FD5EA9DB561C74CFF54D8668985FBF0" box="[225,249,1041,1062]" italics="true" pageId="36" pageNumber="37">db</emphasis>
or nearly so (
<figureCitation id="C59A2A0AB561C74CFEC8D86688B8FBF0" box="[381,452,1041,1062]" captionStart="FIGURES 4851" captionStartId="33.[151,263,1857,1880]" captionTargetBox="[180,1406,245,1790]" captionTargetId="figure-68@33.[151,1436,193,1836]" captionTargetPageId="33" captionText="FIGURES 4851. Right pedipalp chela, distribution of external and dorsal trichobothria. 48. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 49. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 50. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 51. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284814" httpUri="https://zenodo.org/record/5284814/files/figure.png" pageId="36" pageNumber="37">Fig. 51</figureCitation>
); pectines with 2324 teeth on males and 2123 on females.........................
<taxonomicName id="9AA14D0CB561C74CFA83D8668CE0FBF0" ID-CoL="66LKN" authorityName="Lourenco" authorityYear="1994" box="[1334,1436,1041,1062]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="ehrlichi">
<emphasis id="6FD5EA9DB561C74CFA83D8668CE0FBF0" box="[1334,1436,1041,1062]" italics="true" pageId="36" pageNumber="37">A. ehrlichi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22D8588CE7FBB7" blockId="36.[151,1437,151,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22D8588CE7FBB7" pageId="36" pageNumber="37">
11 Pedipalp hand completely yellow; fixed finger trichobothria in the order
<emphasis id="6FD5EA9DB561C74CFC23D8588AD1FB92" box="[918,941,1071,1092]" italics="true" pageId="36" pageNumber="37">eb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFC01D8588AA8FB92" box="[948,980,1071,1092]" italics="true" pageId="36" pageNumber="37">esb</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFC6FD8588A8EFB92" box="[986,1010,1071,1092]" italics="true" pageId="36" pageNumber="37">db</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFC4CD8588D68FB92" box="[1017,1044,1071,1092]" italics="true" pageId="36" pageNumber="37">est</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFBAFD8588D57FB92" box="[1050,1067,1071,1092]" italics="true" pageId="36" pageNumber="37">et</emphasis>
:
<emphasis id="6FD5EA9DB561C74CFB87D8588D39FB92" box="[1074,1093,1071,1092]" italics="true" pageId="36" pageNumber="37">dt</emphasis>
, with
<emphasis id="6FD5EA9DB561C74CFB35D8588DE4FB92" box="[1152,1176,1071,1092]" italics="true" pageId="36" pageNumber="37">db</emphasis>
distinctly basal to
<emphasis id="6FD5EA9DB561C74CFAF9D8588C1BFB92" box="[1356,1383,1071,1092]" italics="true" pageId="36" pageNumber="37">est</emphasis>
; pectines with less than 20 teeth............................................................................. 12
</keyLead>
</paragraph>
</keyStep>
<keyStep id="E6552E2AB561C74CFF22D81E8CE7FB4A" pageId="36" pageNumber="37">
<paragraph id="5D1E368FB561C74CFF22D81E8CE0FBA8" blockId="36.[151,1437,151,1180]" box="[151,1436,1129,1150]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22D81E8CE0FBA8" box="[151,1436,1129,1150]" pageId="36" pageNumber="37">
12. Pectines with 1719 teeth on males (female unknown).................................................
<taxonomicName id="9AA14D0CB561C74CFA8FD81E8CE0FBA8" ID-CoL="66KZR" authorityName="Botero-Trujillo" authorityYear="2009" box="[1338,1436,1129,1150]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="faguasi">
<emphasis id="6FD5EA9DB561C74CFA8FD81E8CE0FBA8" box="[1338,1436,1129,1150]" italics="true" pageId="36" pageNumber="37">A. faguasi</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="5D1E368FB561C74CFF22D8F08CE7FB4A" blockId="36.[151,1437,151,1180]" box="[151,1435,1159,1180]" pageId="36" pageNumber="37">
<keyLead id="E65095BAB561C74CFF22D8F08CE7FB4A" box="[151,1435,1159,1180]" pageId="36" pageNumber="37">
12 Pectines with 16 teeth on males and 1516 on females..................................................
<taxonomicName id="9AA14D0CB561C74CFAF9D8F08CE7FB4A" ID-CoL="66LB2" authorityName="Lourenco" authorityYear="1999" box="[1356,1435,1159,1180]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="leilae">
<emphasis id="6FD5EA9DB561C74CFAF9D8F08CE7FB4A" box="[1356,1435,1159,1180]" italics="true" pageId="36" pageNumber="37">A. leilae</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</document>