418 lines
60 KiB
XML
418 lines
60 KiB
XML
<document id="14600DE103838AA7C4B9E9ACE7ADD095" ID-DOI="10.11646/zootaxa.2904.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5284774" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629728268910" checkinUser="felipe" docAuthor="Botero-Trujillo, Ricardo & Flórez, Eduardo" docDate="2011" docId="D5088799B55CC773FF22DD3B8CE7F845" docLanguage="en" docName="zt02904p044.pdf" docOrigin="Zootaxa 2904 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2904.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Ananteris ochoai Botero-Trujillo & Flórez 2011, sp. nov." docType="treatment" docVersion="4" lastPageNumber="28" masterDocId="2931FFE1B545C768FFB5DC77897CFFD6" masterDocTitle="A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group" masterLastPageNumber="44" masterPageNumber="1" pageNumber="26" updateTime="1699102918901" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="420F86AA58531E5896C228E1673C59D6">A revisionary approach of Colombian Ananteris (Scorpiones, Buthidae): two new species, a new synonymy, and notes on the value of trichobothria and hemispermatophore for the taxonomy of the group</mods:title>
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<mods:roleTerm id="DF3C21DB46258F0037BFA7C58CB6D5C6">Author</mods:roleTerm>
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<mods:namePart id="4611D68A6EFD39865BEFB26F6D5B9B17">Botero-Trujillo, Ricardo</mods:namePart>
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<mods:affiliation id="9EE17BBFB77EBF17891BEFD1C3B8B8F7">Laboratorio de Entomología, Unidad de Ecología y Sistemática-UNESIS, Departamento de Biología, Pontificia Universidad Javeriana, Bogotá, Colombia. E-mail: pachyurus @ yahoo. com</mods:affiliation>
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<mods:namePart id="4CA6DD5148D90700624D4421E4252A2F">Flórez, Eduardo</mods:namePart>
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<mods:affiliation id="82DDBB41ABAB583FBFF2FDD388615A47">Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá, Colombia. E-mail: aeflorezd @ unal. edu. co</mods:affiliation>
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<mods:typeOfResource id="5603DEC245F4A459C23FB795173EDC88">text</mods:typeOfResource>
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<mods:title id="D35E37BC92D96A733835D642ED0D2B7A">Zootaxa</mods:title>
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<mods:date id="050752B9824B8C8B25C1B5002BD3D6AA">2011</mods:date>
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<mods:number id="1A1615BDC734FFB8C48DF78683EA2874">2011-06-02</mods:number>
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<mods:number id="3A142B680F5A2EDECAF8AEE689A4CD40">2904</mods:number>
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<paragraph id="5D1E368FB55CC771FF22DD3B88B7FEB3" blockId="25.[151,728,331,390]" box="[151,459,331,358]" pageId="25" pageNumber="26">
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<heading id="065681E3B55CC771FF22DD3B88B7FEB3" bold="true" box="[151,459,331,358]" fontSize="11" level="1" pageId="25" pageNumber="26" reason="1">
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<emphasis id="6FD5EA9DB55CC771FF22DD3B88B7FEB3" bold="true" box="[151,459,331,358]" pageId="25" pageNumber="26">
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<taxonomicName id="9AA14D0CB55CC771FF22DD3B881AFEB0" ID-CoL="66LC5" authority="Botero-Trujillo & Flórez, 2011" authorityName="Botero-Trujillo & Flórez" authorityYear="2011" box="[151,358,332,358]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="ochoai" status="sp. nov.">
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<emphasis id="6FD5EA9DB55CC771FF22DD3B881AFEB0" bold="true" box="[151,358,332,358]" italics="true" pageId="25" pageNumber="26">Ananteris ochoai</emphasis>
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<taxonomicNameLabel id="74E657E6B55CC771FEDBDD3C88B7FEB3" box="[366,459,331,357]" pageId="25" pageNumber="26" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="15BB6504B55CC771FF22DD198BA4FE50" box="[151,728,366,390]" pageId="25" pageNumber="26" type="description">
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<paragraph id="5D1E368FB55CC771FF22DD198BA4FE50" blockId="25.[151,728,331,390]" box="[151,728,366,390]" pageId="25" pageNumber="26">
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<figureCitation id="C59A2A0AB55CC771FF22DD19887CFE50" box="[151,256,366,390]" captionStart="FIGURE 1" captionStartId="6.[151,250,1822,1845]" captionTargetBox="[177,1415,197,1775]" captionTargetId="figure-117@6.[174,1418,195,1778]" captionTargetPageId="6" captionText="FIGURE 1. Known distribution of Colombian Ananteris. () Ananteris arcadioi Botero-Trujillo, 2008. () Ananteris columbiana Lourenço, 1991. () Ananteris dorae Botero-Trujillo, 2008. () A. dorae and Ananteris ochoai sp. nov. ()Ananteris ehrlichi Lourenço, 1994. () Ananteris faguasi Botero-Trujillo, 2009. () Ananteris gorgonae Lourenço & Flórez, 1989. () Ananteris leilae Lourenço, 1999. () Ananteris meridana González-Sponga, 2006 (also type locality in Venezuela). () Ananteris myriamae Botero-Trujillo, 2007. () Ananteris solimariae sp. nov. ()Ananteris tolimana Teruel & García, 2007. () Ananteris volschenki Botero-Trujillo, 2009." figureDoi="http://doi.org/10.5281/zenodo.5284778" httpUri="https://zenodo.org/record/5284778/files/figure.png" pageId="25" pageNumber="26">Figures 1</figureCitation>
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,
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<figureCitation id="C59A2A0AB55CC771FEBBDD19882AFE50" box="[270,342,366,390]" captionStart="FIGURES 24–27" captionStartId="26.[151,264,1519,1542]" captionTargetBox="[154,1433,180,1491]" captionTargetId="figure-217@26.[151,1436,174,1498]" captionTargetPageId="26" captionText="FIGURES 24–27. Ananteris ochoai sp. nov. 24–25. Holotype male, dorsal and ventral aspects (MPUJ-SCO-388). 26–27. Allotype female, dorsal and ventral aspects (MPUJ-SCO-389). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5284798" httpUri="https://zenodo.org/record/5284798/files/figure.png" pageId="25" pageNumber="26">24–27</figureCitation>
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,
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<figureCitation id="C59A2A0AB55CC771FED4DD198803FE50" box="[353,383,366,390]" captionStart="FIGURES 36–39" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 36–39. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="25" pageNumber="26">38</figureCitation>
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,
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<figureCitation id="C59A2A0AB55CC771FE39DD1988D6FE50" box="[396,426,366,390]" captionStart="FIGURES 40–47" captionStartId="32.[151,263,1802,1825]" captionTargetBox="[151,1411,227,1748]" captionTargetId="figure-101@32.[151,1436,193,1781]" captionTargetPageId="32" captionText="FIGURES 40–47. Right pedipalp hand, distribution of ventral trichobothria. 40. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 41. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 42. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 43. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). 44. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 45. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 46. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 47. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284810" httpUri="https://zenodo.org/record/5284810/files/figure.png" pageId="25" pageNumber="26">46</figureCitation>
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,
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<figureCitation id="C59A2A0AB55CC771FE03DD1988A8FE50" box="[438,468,366,390]" captionStart="FIGURES 52–55" captionStartId="34.[151,263,1859,1882]" captionTargetBox="[179,1408,232,1790]" captionTargetId="figure-71@34.[151,1436,193,1838]" captionTargetPageId="34" captionText="FIGURES 52–55. Right pedipalp chela, distribution of external and dorsal trichobothria. 52. Ananteris meridana González- Sponga, 2006, adult female (ICN-As-321). 53. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 54. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 55. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO- 390). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284818" httpUri="https://zenodo.org/record/5284818/files/figure.png" pageId="25" pageNumber="26">54</figureCitation>
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,
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<figureCitation id="C59A2A0AB55CC771FE55DD198B54FE50" box="[480,552,366,390]" captionStart="FIGURES 56–57" captionStartId="35.[151,263,1091,1114]" captionTargetBox="[152,1435,198,1061]" captionTargetId="figure-698@35.[151,1436,193,1070]" captionTargetPageId="35" captionText="FIGURES 56–57. Two views of the collection locality of Ananteris ochoai sp. nov. and Ananteris dorae Botero-Trujillo, 2008 in Río Ñambi Natural Reserve, Nariño Department. Photographs by José A. Ochoa." figureDoi="http://doi.org/10.5281/zenodo.5284822" httpUri="https://zenodo.org/record/5284822/files/figure.png" pageId="25" pageNumber="26">56–57</figureCitation>
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,
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<figureCitation id="C59A2A0AB55CC771FD81DD198B2FFE50" box="[564,595,366,390]" captionStart="FIGURES 64–67" captionStartId="39.[151,264,1792,1815]" captionTargetBox="[199,1415,211,1744]" captionTargetId="figure-110@39.[151,1436,193,1771]" captionTargetPageId="39" captionText="FIGURES 64–67. Hemispermatophore in ectal aspect and detail of the capsular region in ventral aspect (left for Ananteris solimariae sp. nov., inverted; right for others). 64. Ananteris meridana González-Sponga, 2006 (ICN-As-435). 65. Ananteris myriamae Botero-Trujillo, 2007 (MPUJ-SCO-382). 66. Ananteris ochoai sp. nov., paratype (ICN-As-815). 67. Ananteris solimariae sp. nov., paratype (ICN-As-779). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284832" httpUri="https://zenodo.org/record/5284832/files/figure.png" pageId="25" pageNumber="26">66</figureCitation>
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;
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<tableCitation id="10230334B55CC771FDEBDD198BC0FE50" box="[606,700,366,390]" captionStart="TABLE 1" captionStartId="2.[151,235,295,318]" captionText="TABLE 1. Checklist of the Colombian species of Ananteris with indication to the biogeographic regions (sensu Hernández- Camacho et al. 1992) and departments where they are known from. Asterisk (*) indicates departments for which the genus is herein recorded for the first time." pageId="25" pageNumber="26">Tables 1</tableCitation>
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,
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<tableCitation id="10230334B55CC771FD7FDD198BA4FE50" box="[714,728,366,390]" captionStart="TABLE 2" captionStartId="5.[151,235,155,178]" captionText="TABLE 2. Meristic data for Ananteris species. Measurements in millimeters. 1Sum of prosoma, mesosoma and metasoma. 2Sum of tergites I–VII. 3Sum of metasomal segments I–V and telson. 4Sum of femur, patella and chela. 5Measured from the commissure of the junction with the movable finger to the finger tip. 6Measured along the marginal lamella. See for other measurements: Lourenço (1991) and Teruel & Roncallo (2008) for A. columbiana, Lourenço (1994) for A. ehrlichi, Botero-Trujillo (2007) for A. myriamae, Botero-Trujillo (2008a) for A. arcadioi and A. dorae." pageId="25" pageNumber="26">2</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="15BB6504B55CC771FF22DDC08DBBFD54" pageId="25" pageNumber="26" type="materials_examined">
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<emphasis id="6FD5EA9DB55CC771FF22DDC0883AFE19" bold="true" box="[151,326,438,463]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Type material.</emphasis>
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<materialsCitation id="EDC93CD2B55CC771FEFBDDC18A56FDC0" collectingDate="2008-09-11" collectionCode="UV, R" collectorName="Path El Barro & O. Tovar & J. P. Botero & Botero-Trujillo, J & J. A. Ochoa" country="Colombia" county="Barbacoas" elevation="1397" latitude="1.2861111" location="Rio Nambi Natural Reserve" longLatPrecision="21" longitude="-78.07444" municipality="Path El Barro" pageId="25" pageNumber="26" specimenCount="1" stateProvince="Narino" typeStatus="holotype">
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<typeStatus id="821A882DB55CC771FEFBDDC188C9FE18" box="[334,437,438,462]" pageId="25" pageNumber="26" type="holotype">Holotype</typeStatus>
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male (adult):
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</emphasis>
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<collectingCountry id="25B6761FB55CC771FDEFDDC18B88FE18" box="[602,756,438,462]" name="Colombia" pageId="25" pageNumber="26">COLOMBIA</collectingCountry>
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:
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<collectingRegion id="9F65F86DB55CC771FCB5DDC08A9DFE18" box="[768,993,439,462]" country="Colombia" name="Narino" pageId="25" pageNumber="26">Nariño Department</collectingRegion>
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:
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</emphasis>
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<collectingCounty id="B47F4E03B55CC771FC45DDC18D1BFE18" box="[1008,1127,438,462]" pageId="25" pageNumber="26">Barbacoas</collectingCounty>
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,
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<collectorName id="F0545359B55CC771FBC1DDC18C6DFE18" box="[1140,1297,438,462]" pageId="25" pageNumber="26">
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<collectingMunicipality id="BD7AACF5B55CC771FBC1DDC18C6DFE18" box="[1140,1297,438,462]" pageId="25" pageNumber="26">Path El Barro</collectingMunicipality>
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</collectorName>
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,
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<location id="587E6054B55CC771FAAADDC1882CFE24" LSID="urn:lsid:plazi:treatment:D5088799B55CC773FF22DD3B8CE7F845:587E6054B55CC771FAAADDC1882CFE24" country="Colombia" county="Barbacoas" latitude="1.2861111" longLatPrecision="21" longitude="-78.07444" municipality="Path El Barro" name="Rio Nambi Natural Reserve" pageId="25" pageNumber="26" stateProvince="Narino">Río Ñambi Natural Reserve</location>
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,
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<geoCoordinate id="38955048B55CC771FEE8DDAD8896FE24" box="[349,490,474,498]" degrees="01" direction="north" minutes="17" orientation="latitude" pageId="25" pageNumber="26" precision="15" seconds="10" value="1.2861111">01°17’10’’N</geoCoordinate>
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,
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<elevation id="D68CD1BCB55CC771FD26DDAD8A6FFE24" box="[659,787,474,498]" metricMagnitude="3" metricUnit="m" metricValue="1.3969999999999998" pageId="25" pageNumber="26" unit="m" value="1397.0">
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asl
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</elevation>
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,
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<collectionCode id="3BB0AE4AB55CC771FC95DDAC8A34FE24" box="[800,840,475,498]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="25" pageNumber="26">UV</collectionCode>
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light,
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<date id="291F104FB55CC771FC26DDAD8D0DFE24" box="[915,1137,474,498]" pageId="25" pageNumber="26" value="2008-09-11">
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<collectingDate id="395BE9A7B55CC771FC26DDAD8D0DFE24" box="[915,1137,474,498]" pageId="25" pageNumber="26" value="2008-09-11">11 September 2008</collectingDate>
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</date>
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,
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<collectorName id="F0545359B55CC771FBCBDDAD8D98FE24" box="[1150,1252,474,498]" pageId="25" pageNumber="26">O. Tovar</collectorName>
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,
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<collectorName id="F0545359B55CC771FB44DDAC8C0AFE24" box="[1265,1398,475,498]" pageId="25" pageNumber="26">J. P. Botero</collectorName>
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,
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<collectionCode id="3BB0AE4AB55CC771FA37DDAC8CE4FE24" box="[1410,1432,475,498]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="25" pageNumber="26">R</collectionCode>
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.
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<collectorName id="F0545359B55CC771FF22DD898821FDC0" box="[151,349,510,534]" pageId="25" pageNumber="26">Botero-Trujillo, J</collectorName>
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. Souza &
|
||
<collectorName id="F0545359B55CC771FE7ADD888B2EFDC0" box="[463,594,510,534]" pageId="25" pageNumber="26">J. A. Ochoa</collectorName>
|
||
(MPUJ-SCO-388)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="EDC93CD2B55CC771FC81DD898886FDEC" country="Colombia" location="COLOMBIA" pageId="25" pageNumber="26" specimenCount="2" typeStatus="allotype">
|
||
<emphasis id="6FD5EA9DB55CC771FC81DD898D43FDC0" box="[820,1087,510,534]" italics="true" pageId="25" pageNumber="26">
|
||
<typeStatus id="821A882DB55CC771FC81DD898AEEFDC0" box="[820,914,510,534]" pageId="25" pageNumber="26" type="allotype">Allotype</typeStatus>
|
||
female (adult):
|
||
</emphasis>
|
||
<collectingCountry id="25B6761FB55CC771FBF3DD898D9CFDC0" box="[1094,1248,510,534]" name="Colombia" pageId="25" pageNumber="26">COLOMBIA</collectingCountry>
|
||
: identical data to the
|
||
<typeStatus id="821A882DB55CC771FF75DE55885EFDEC" box="[192,290,546,570]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
(MPUJ-SCO-389)
|
||
</materialsCitation>
|
||
.
|
||
<emphasis id="6FD5EA9DB55CC771FDB1DE548B02FDEC" box="[516,638,547,570]" italics="true" pageId="25" pageNumber="26">
|
||
<typeStatus id="821A882DB55CC771FDB1DE548B05FDEC" box="[516,633,547,570]" pageId="25" pageNumber="26" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="4BA7FD06B55CC771FD33DE558B86FDEC" box="[646,762,546,570]" pageId="25" pageNumber="26" type="subadult">1 subadult</specimenCount>
|
||
male and
|
||
<specimenCount id="4BA7FD06B55CC771FCD8DE558AC6FDEC" box="[877,954,546,570]" pageId="25" pageNumber="26" type="adult">2 adult</specimenCount>
|
||
females with identical data to the
|
||
<typeStatus id="821A882DB55CC771FA8FDE558CE0FDEC" box="[1338,1436,546,570]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
(ICN-As-799).
|
||
<specimenCount id="4BA7FD06B55CC771FEF6DE3188ECFD88" box="[323,400,582,606]" pageId="25" pageNumber="26" type="adult">3 adult</specimenCount>
|
||
males, 1 small(?) male,
|
||
<specimenCount id="4BA7FD06B55CC771FD2ADE318A6EFD88" box="[671,786,582,606]" pageId="25" pageNumber="26" type="subadult">1 subadult</specimenCount>
|
||
male and
|
||
<materialsCitation id="EDC93CD2B55CC771FC31DE318DBFFD54" collectingDate="2009-10-13" collectionCode="UV" collectorName="E. Florez & D. Luna" country="Colombia" county="Rio Nambi Natural Reserve" elevation="1440" location="Barbacoas" municipality="Altaquer" pageId="25" pageNumber="26" specimenCount="1" specimenCount-juvenile="1" typeStatus="paratype">
|
||
<specimenCount id="4BA7FD06B55CC771FC31DE318A8FFD88" box="[900,1011,582,606]" pageId="25" pageNumber="26" type="juvenile">1 juvenile</specimenCount>
|
||
male,
|
||
<location id="587E6054B55CC771FB88DE318DC8FD88" LSID="urn:lsid:plazi:treatment:D5088799B55CC773FF22DD3B8CE7F845:587E6054B55CC771FB88DE318DC8FD88" box="[1085,1204,582,606]" country="Colombia" county="Rio Nambi Natural Reserve" municipality="Altaquer" name="Barbacoas" pageId="25" pageNumber="26">Barbacoas</location>
|
||
,
|
||
<collectingMunicipality id="BD7AACF5B55CC771FB0ADE318C5FFD88" box="[1215,1315,582,606]" pageId="25" pageNumber="26">Altaquer</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="B47F4E03B55CC771FA98DE318810FD54" pageId="25" pageNumber="26">Río Ñambi Natural Reserve</collectingCounty>
|
||
,
|
||
<elevation id="D68CD1BCB55CC771FEC5DE1D8890FD54" box="[368,492,618,642]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="25" pageNumber="26" unit="m" value="1440.0">
|
||
<quantity id="9A599B6AB55CC771FEC5DE1D88B9FD54" box="[368,453,618,642]" metricMagnitude="3" metricUnit="m" metricValue="1.44" pageId="25" pageNumber="26" unit="m" value="1440.0">1440 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<collectionCode id="3BB0AE4AB55CC771FE4CDE1C8B5DFD54" box="[505,545,619,642]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="25" pageNumber="26">UV</collectionCode>
|
||
light,
|
||
<date id="291F104FB55CC771FDDCDE1D8A55FD54" box="[617,809,618,642]" pageId="25" pageNumber="26" value="2009-10-13">
|
||
<collectingDate id="395BE9A7B55CC771FDDCDE1D8A55FD54" box="[617,809,618,642]" pageId="25" pageNumber="26" value="2009-10-13">13 October 2009</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="F0545359B55CC771FC81DE1C8AE6FD54" box="[820,922,618,642]" pageId="25" pageNumber="26">E. Flórez</collectorName>
|
||
&
|
||
<collectorName id="F0545359B55CC771FC0BDE1C8D65FD54" box="[958,1049,619,642]" pageId="25" pageNumber="26">D. Luna</collectorName>
|
||
(ICN-As-815)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="15BB6504B55CC771FF70DEF88876FD38" pageId="25" pageNumber="26" type="etymology">
|
||
<paragraph id="5D1E368FB55CC771FF70DEF88876FD38" blockId="25.[151,1436,438,2016]" pageId="25" pageNumber="26">
|
||
<emphasis id="6FD5EA9DB55CC771FF70DEF88830FD71" bold="true" box="[197,332,655,679]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Etymology.</emphasis>
|
||
This species is named after our colleague and friend José A. Ochoa, currently at the Universidade de
|
||
<collectingRegion id="9F65F86DB55CC771FF0DDEC58855FD1C" box="[184,297,690,714]" country="Brazil" name="Sao Paulo" pageId="25" pageNumber="26">São Paulo</collectingRegion>
|
||
(
|
||
<collectingCountry id="25B6761FB55CC771FE82DEC588FDFD1C" box="[311,385,690,714]" name="Brazil" pageId="25" pageNumber="26">Brazil</collectingCountry>
|
||
), in recognition to his contributions to the knowledge of South American (primarily Peruvian) scorpions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="15BB6504B55CC773FF70DE8D8D7CF909" lastPageId="27" lastPageNumber="28" pageId="25" pageNumber="26" type="diagnosis">
|
||
<paragraph id="5D1E368FB55CC771FF70DE8D8A1FFB72" blockId="25.[151,1436,438,2016]" pageId="25" pageNumber="26">
|
||
<emphasis id="6FD5EA9DB55CC771FF70DE8D8843FCC5" bold="true" box="[197,319,762,787]" inLineHeading="true" pageId="25" pageNumber="26" reason="1">Diagnosis.</emphasis>
|
||
Anterior margin of the carapace very slightly bi-concave, without anteromedian projection (
|
||
<figureCitation id="C59A2A0AB55CC771FADBDE8D89CAFCE0" captionStart="FIGURES 36–39" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 36–39. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="25" pageNumber="26">Fig. 38</figureCitation>
|
||
); metasomal carinal formula 10:10:10:8:5, with median lateral carinae on segment III obsolete; telson vesicle densely granulose ventrally and laterally (more conspicuous in females), with three ventral longitudinal carinae armed with strong granulation, and one noticeably larger granule (tubercle-like) under the subaculear tubercle;
|
||
<emphasis id="6FD5EA9DB55CC771FA36DF108CE0FC55" box="[1411,1436,871,899]" italics="true" pageId="25" pageNumber="26">
|
||
V
|
||
<subScript id="C12534CAB55CC771FA21DF018CE0FC55" attach="left" box="[1428,1436,886,899]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
</emphasis>
|
||
and
|
||
<emphasis id="6FD5EA9DB55CC771FF73DFF989A3FC7F" box="[198,223,910,937]" italics="true" pageId="25" pageNumber="26">
|
||
V
|
||
<subScript id="C12534CAB55CC771FF62DFEB89A3FC7F" attach="left" box="[215,223,924,937]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
|
||
</emphasis>
|
||
trichobothria very slightly unaligned axially, with
|
||
<emphasis id="6FD5EA9DB55CC771FCADDFF98A4EFC7F" box="[792,818,910,937]" italics="true" pageId="25" pageNumber="26">
|
||
V
|
||
<subScript id="C12534CAB55CC771FC9FDFEB8A4EFC7F" attach="left" box="[810,818,924,937]" fontSize="6" pageId="25" pageNumber="26">2</subScript>
|
||
</emphasis>
|
||
located only slightly external to
|
||
<emphasis id="6FD5EA9DB55CC771FB17DFF98DC7FC7F" box="[1186,1211,910,937]" italics="true" pageId="25" pageNumber="26">
|
||
V
|
||
<subScript id="C12534CAB55CC771FB06DFEB8DC7FC7F" attach="left" box="[1203,1211,924,937]" fontSize="6" pageId="25" pageNumber="26">1</subScript>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="C59A2A0AB55CC771FB7FDFFA8C63FC73" box="[1226,1311,909,933]" captionStart="FIGURES 40–47" captionStartId="32.[151,263,1802,1825]" captionTargetBox="[151,1411,227,1748]" captionTargetId="figure-101@32.[151,1436,193,1781]" captionTargetPageId="32" captionText="FIGURES 40–47. Right pedipalp hand, distribution of ventral trichobothria. 40. Ananteris arcadioi Botero-Trujillo, 2008, adult female (MPUJ-SCO-383). 41. Ananteris columbiana Lourenço, 1991, adult female (IAvH-E 100778). 42. Ananteris dorae Botero-Trujillo, 2008, adult male (ICN-As-816). 43. Ananteris ehrlichi Lourenço, 1994, adult male (IAvH-E 100814). 44. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 45. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 46. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 47. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284810" httpUri="https://zenodo.org/record/5284810/files/figure.png" pageId="25" pageNumber="26">Fig. 46</figureCitation>
|
||
); fixed finger trichobothria in the order
|
||
<emphasis id="6FD5EA9DB55CC771FE5BDFC38B74FC1A" box="[494,520,948,972]" italics="true" pageId="25" pageNumber="26">eb</emphasis>
|
||
:
|
||
<emphasis id="6FD5EA9DB55CC771FDA4DFC38B4AFC1A" box="[529,566,948,972]" italics="true" pageId="25" pageNumber="26">esb</emphasis>
|
||
:
|
||
<emphasis id="6FD5EA9DB55CC771FD8BDFC28B21FC1A" box="[574,605,949,972]" italics="true" pageId="25" pageNumber="26">est</emphasis>
|
||
:
|
||
<emphasis id="6FD5EA9DB55CC771FDD1DFC38BFCFC1A" box="[612,640,948,972]" italics="true" pageId="25" pageNumber="26">db</emphasis>
|
||
:
|
||
<emphasis id="6FD5EA9DB55CC771FD3DDFC28BE0FC1A" box="[648,668,949,972]" italics="true" pageId="25" pageNumber="26">et</emphasis>
|
||
:
|
||
<emphasis id="6FD5EA9DB55CC771FD11DFC38BC6FC1A" box="[676,698,948,972]" italics="true" pageId="25" pageNumber="26">dt</emphasis>
|
||
(
|
||
<figureCitation id="C59A2A0AB55CC771FD79DFC38A5FFC1A" box="[716,803,948,972]" captionStart="FIGURES 52–55" captionStartId="34.[151,263,1859,1882]" captionTargetBox="[179,1408,232,1790]" captionTargetId="figure-71@34.[151,1436,193,1838]" captionTargetPageId="34" captionText="FIGURES 52–55. Right pedipalp chela, distribution of external and dorsal trichobothria. 52. Ananteris meridana González- Sponga, 2006, adult female (ICN-As-321). 53. Ananteris myriamae Botero-Trujillo, 2007, adult male (MPUJ-SCO-382). 54. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 55. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO- 390). Scale bars = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5284818" httpUri="https://zenodo.org/record/5284818/files/figure.png" pageId="25" pageNumber="26">Fig. 54</figureCitation>
|
||
); dorsal surface of chelicerae with dense and complete reticular pattern, and small yellowish rounded area anteriorly (
|
||
<figureCitation id="C59A2A0AB55CC771FCE3DFAF8AD6FC26" box="[854,938,984,1008]" captionStart="FIGURES 36–39" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 36–39. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="25" pageNumber="26">Fig. 38</figureCitation>
|
||
); pectines with 20–23 teeth on males (mode = 21), 22–23 on females (mode = 22); sternite VII with incomplete but strong paramedian longitudinal carinae; pedipalp hand uniformly yellow and spotless; movable and fixed fingers yellow in the basal third/quarter, the rest brownish; coxosternal region predominantly yellow and spotless, except for brownish regions on coxapophyses (and coxae on males) I and anterior tip of coxapophyses II. Hemispermatophore: capsular region with two lobes; flagellum with long
|
||
<emphasis id="6FD5EA9DB55CC771FECED8FA8897FB72" box="[379,491,1165,1188]" italics="true" pageId="25" pageNumber="26">pars recta</emphasis>
|
||
and coiled
|
||
<emphasis id="6FD5EA9DB55CC771FDDBD8FA8B8FFB72" box="[622,755,1164,1188]" italics="true" pageId="25" pageNumber="26">pars reflexa</emphasis>
|
||
(
|
||
<figureCitation id="C59A2A0AB55CC771FCB6D8FB8A2BFB72" box="[771,855,1164,1188]" captionStart="FIGURES 64–67" captionStartId="39.[151,264,1792,1815]" captionTargetBox="[199,1415,211,1744]" captionTargetId="figure-110@39.[151,1436,193,1771]" captionTargetPageId="39" captionText="FIGURES 64–67. Hemispermatophore in ectal aspect and detail of the capsular region in ventral aspect (left for Ananteris solimariae sp. nov., inverted; right for others). 64. Ananteris meridana González-Sponga, 2006 (ICN-As-435). 65. Ananteris myriamae Botero-Trujillo, 2007 (MPUJ-SCO-382). 66. Ananteris ochoai sp. nov., paratype (ICN-As-815). 67. Ananteris solimariae sp. nov., paratype (ICN-As-779). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284832" httpUri="https://zenodo.org/record/5284832/files/figure.png" pageId="25" pageNumber="26">Fig. 66</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55CC771FF70D8C788C9F836" blockId="25.[151,1436,438,2016]" pageId="25" pageNumber="26">
|
||
<emphasis id="6FD5EA9DB55CC771FF70D8C78874FB1E" bold="true" box="[197,264,1200,1224]" pageId="25" pageNumber="26">Male.</emphasis>
|
||
Description based on
|
||
<typeStatus id="821A882DB55CC771FDB0D8C78B16FB1E" box="[517,618,1200,1224]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
|
||
.
|
||
<emphasis id="6FD5EA9DB55CC771FDC1D8C68A5CFB1E" box="[628,800,1201,1224]" italics="true" pageId="25" pageNumber="26">Measurements:</emphasis>
|
||
see
|
||
<tableCitation id="10230334B55CC771FCE7D8C78AD4FB1E" box="[850,936,1200,1224]" captionStart="TABLE 2" captionStartId="5.[151,235,155,178]" captionText="TABLE 2. Meristic data for Ananteris species. Measurements in millimeters. 1Sum of prosoma, mesosoma and metasoma. 2Sum of tergites I–VII. 3Sum of metasomal segments I–V and telson. 4Sum of femur, patella and chela. 5Measured from the commissure of the junction with the movable finger to the finger tip. 6Measured along the marginal lamella. See for other measurements: Lourenço (1991) and Teruel & Roncallo (2008) for A. columbiana, Lourenço (1994) for A. ehrlichi, Botero-Trujillo (2007) for A. myriamae, Botero-Trujillo (2008a) for A. arcadioi and A. dorae." pageId="25" pageNumber="26">Table 2</tableCitation>
|
||
.
|
||
<emphasis id="6FD5EA9DB55CC771FC01D8C78D44FB1E" box="[948,1080,1200,1224]" italics="true" pageId="25" pageNumber="26">Coloration:</emphasis>
|
||
general coloration yellowish to reddish-brown with variegated pigmentation over almost the entire body and appendages (
|
||
<figureCitation id="C59A2A0AB55CC771FB20D8A38C5CFB3A" box="[1173,1312,1236,1260]" captionStart="FIGURES 24–27" captionStartId="26.[151,264,1519,1542]" captionTargetBox="[154,1433,180,1491]" captionTargetId="figure-217@26.[151,1436,174,1498]" captionTargetPageId="26" captionText="FIGURES 24–27. Ananteris ochoai sp. nov. 24–25. Holotype male, dorsal and ventral aspects (MPUJ-SCO-388). 26–27. Allotype female, dorsal and ventral aspects (MPUJ-SCO-389). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5284798" httpUri="https://zenodo.org/record/5284798/files/figure.png" pageId="25" pageNumber="26">Figs. 24–25</figureCitation>
|
||
). Carapace predominantly brown with some yellow spots and stripes; anterior and posterior margins brown; median ocular tubercle black, surrounded on its base by thin yellow lines on anterolateral and posterolateral margins. Chelicerae with yellow coxa; hand with dense and complete reticular pattern on dorsal surface, and small yellowish rounded area anteriorly (
|
||
<figureCitation id="C59A2A0AB55CC771FEF2D91388E6FAAA" box="[327,410,1380,1404]" captionStart="FIGURES 36–39" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 36–39. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="25" pageNumber="26">Fig. 38</figureCitation>
|
||
); fixed and movable fingers dark-brown, each with reddish teeth. Coxosternal region almost entirely yellow and spotless, except for brownish regions on coxapophyses I, coxae I and anterior tip of coxapophyses II; genital operculum, pectinal basal piece, pectines and sternite III entirely yellow; sternites IV–VI with lateral margins brown; sternites V–VI with inconspicuous brownish areas located posteriorly midway between the midline and the lateral margins; sternite VII with broad brown longitudinal bands between the paramedian carinae and the lateral margins; sternite V with a subtriangular posterior median hyaline area. Tergites predominantly brown; two longitudinal yellow stripes crossing tergites II–VI are only evident on the posterior half of each segment beside the midline; each side of tergites I–VI with two transverse yellow lines converging near the longitudinal lines, arrow-like; tergite VII predominantly brown, with large yellow regions dorsally and laterally; lateral margins of tergites I–VII completely yellow. Metasoma predominantly yellowish-red; dorsal intercarinal spaces of segments I–IV with a median brown design wider anteriorly, triangle-like on segments I–II but arrow-like on segments III–IV; segments I–V with variegated pigmentation on all surfaces. Telson vesicle yellowish-red, lighter than segment V and with a longitudinal red band located on each side; aculeus yellowish basally, reddish distally; subaculear tubercle yellow. Pedipalp coxa and trochanter with variegated pigmentation; femur brown on all surfaces but lighter ventrally, trichobothrial pits yellow; patella brown with variegated pigmentation on dorsal, external and ventral surfaces, trichobothrial pits yellow; hand uniformly yellow and spotless; movable and fixed fingers yellow on the basal quarter and brownish on the rest of their length. Legs with variegated pigmentation from trochanter to basitarsi, telotarsi yellow.
|
||
</paragraph>
|
||
<caption id="09DE6607B55FC772FF22D9988AA8F9F2" ID-DOI="http://doi.org/10.5281/zenodo.5284798" ID-Zenodo-Dep="5284798" httpUri="https://zenodo.org/record/5284798/files/figure.png" pageId="26" pageNumber="27" startId="26.[151,264,1519,1542]" targetBox="[154,1433,180,1491]" targetPageId="26">
|
||
<paragraph id="5D1E368FB55FC772FF22D9988AA8F9F2" blockId="26.[151,1436,1519,1572]" pageId="26" pageNumber="27">
|
||
<emphasis id="6FD5EA9DB55FC772FF22D998882BF9D0" bold="true" box="[151,343,1519,1542]" pageId="26" pageNumber="27">FIGURES 24–27.</emphasis>
|
||
<taxonomicName id="9AA14D0CB55FC772FED4D9878B6EF9D0" authority="Botero-Trujillo & Flórez, 2011" authorityName="Botero-Trujillo & Flórez" authorityYear="2011" box="[353,530,1520,1542]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="ochoai" status="sp. nov.">
|
||
<emphasis id="6FD5EA9DB55FC772FED4D9878B6EF9D0" box="[353,530,1520,1542]" italics="true" pageId="26" pageNumber="27">Ananteris ochoai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6FD5EA9DB55FC772FDA9D9878BBCF9D0" bold="true" box="[540,704,1519,1542]" pageId="26" pageNumber="27">
|
||
<taxonomicNameLabel id="74E657E6B55FC772FDA9D9878B0CF9D3" box="[540,624,1520,1541]" pageId="26" pageNumber="27" rank="species">sp. nov.</taxonomicNameLabel>
|
||
24–25.
|
||
</emphasis>
|
||
Holotype male, dorsal and ventral aspects (MPUJ-SCO-388).
|
||
<emphasis id="6FD5EA9DB55FC772FAE3D9988CE0F9D0" bold="true" box="[1366,1436,1519,1542]" pageId="26" pageNumber="27">26–27.</emphasis>
|
||
Allotype female, dorsal and ventral aspects (MPUJ-SCO-389). Scale bars = 1 cm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="5D1E368FB55FC772FF70DA2689A9F8CB" blockId="26.[151,1437,1617,2037]" pageId="26" pageNumber="27">
|
||
<emphasis id="6FD5EA9DB55FC772FF70DA268840F9BF" box="[197,316,1617,1641]" italics="true" pageId="26" pageNumber="27">Carapace:</emphasis>
|
||
densely covered with rounded granules especially on the brown areas; lateral margins converging anteriorly; anterior margin very slightly bi-concave, without anteromedian projection (
|
||
<figureCitation id="C59A2A0AB55FC772FB3DDA028D9CF95B" box="[1160,1248,1653,1677]" captionStart="FIGURES 36–39" captionStartId="30.[151,263,1801,1824]" captionTargetBox="[265,1323,740,1771]" captionTargetId="figure-337@30.[264,1323,733,1781]" captionTargetPageId="30" captionText="FIGURES 36–39. Anterior margin of carapace and chelicerae. 36. Ananteris meridana González-Sponga, 2006, adult female (ICN-As-321). 37. Ananteris myriamae Botero-Trujillo, 2007, adult male (ICN-As-385). 38. Ananteris ochoai sp. nov., holotype male (MPUJ-SCO-388). 39. Ananteris solimariae sp. nov., holotype male (MPUJ-SCO-390)." figureDoi="http://doi.org/10.5281/zenodo.5284808" httpUri="https://zenodo.org/record/5284808/files/figure.png" pageId="26" pageNumber="27">Fig. 38</figureCitation>
|
||
); ocular carinae strong and granulose, others inconspicuous; with a median depression anterior to the ocular tubercle; postocular furrow evident; median ocular tubercle low, located on the posterior half of the anterior third of carapace; lateral ocular tubercles each with four ocelli, being the posteriormost very reduced and unaligned with the remaining three pairs.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55FC772FF70DB5E8B53F823" blockId="26.[151,1437,1617,2037]" pageId="26" pageNumber="27">
|
||
<emphasis id="6FD5EA9DB55FC772FF70DB5E8835F897" box="[197,329,1833,1857]" italics="true" pageId="26" pageNumber="27">Chelicerae:</emphasis>
|
||
with abundant fine white setae on the internal and ventral surfaces; cheliceral dentition characteristic of the family
|
||
<taxonomicName id="9AA14D0CB55FC772FEDFDB3A8BFDF8B3" authority="(Vachon 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[362,641,1869,1893]" class="Arachnida" family="Buthidae" kingdom="Animalia" order="Scorpiones" pageId="26" pageNumber="27" phylum="Arthropoda" rank="family">
|
||
Buthidae (
|
||
<bibRefCitation id="39304B7EB55FC772FE6ADB3A8B05F8B3" author="Vachon" box="[479,633,1869,1893]" firstAuthor="Vachon" pageId="26" pageNumber="27" pagination="161 - 166" refId="ref33387" refString="Vachon, M. (1963) De l'utilite', en systematique, d'une nomenclature des dents des cheliceres chez les scorpions. Bulletin du Museum national de Histoire naturelle Paris (2 e ser.), 35 (2), 161 - 166." type="journal article" year="1963">Vachon 1963</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Movable finger dorsally with two small basal teeth, one median pronounced, one subdistal slightly shorter than the median, and one distal tooth. Ventrally with three strong teeth, one basal, one median, and one distal tooth larger than its dorsal counterpart. Fixed finger dorsally with one basal and one median tooth forming a bicuspid, one subdistal, and one distal tooth. Ventrally with only one subdistal tooth slightly basal to the dorsal subdistal.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DCEF8A37FF21" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DCEF88DAFF79" box="[197,422,152,176]" italics="true" pageId="27" pageNumber="28">Coxosternal region:</emphasis>
|
||
sternum subtriangular, with deep median depression and two anterolateral furrows; all the components of this region with few setae, almost completely smooth except for few granulations on the anterior margin of coxae; coxapophyses I–II densely setose anteriorly.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DD738CFDFEB2" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DD738B4AFECA" box="[197,566,260,284]" italics="true" pageId="27" pageNumber="28">Genital operculum and pectines:</emphasis>
|
||
genital operculum completely divided longitudinally; pectinal basal piece as long as wide, deeply notched anteriorly; pectines long, with distal tip of marginal lamella reaching posterior end of coxae IV; count of pieces on the pectines: marginal lamellae 3:3, middle lamellae 8:9, teeth 21:21, fulcra absent.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DD078A5FFE25" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DD078853FE5E" box="[197,303,368,392]" italics="true" pageId="27" pageNumber="28">Sternites:</emphasis>
|
||
III completely smooth, lateral margins of IV–VI and posterior margin of VI with few granulations, VII densely granulose and with incomplete but strong paramedian longitudinal carinae, lateral carinae absent; V with subtriangular posterior median smooth area; spiracles elongated, approximately twice longer than wide in sternites III–IV and three times longer than wide in V–VI.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DE778D57FD89" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DE77885AFDC1" box="[197,294,512,535]" italics="true" pageId="27" pageNumber="28">Tergites:</emphasis>
|
||
with similar granulation to that of carapace; axial carina only evident on the posterior half of tergites I–VI; dorsolateral carinae absent on tergites I–VI; tergite VII tetracarinate, with incomplete paired dorsolateral and lateral carinae, and granulose median elevation on the position of the axial carina.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DE1A8B8FFC8A" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DE1A883EFD52" box="[197,322,621,644]" italics="true" pageId="27" pageNumber="28">Metasoma:</emphasis>
|
||
with few setae; segments I–III with ten carinae (paired ventrosubmedian, ventrolateral, median lateral, dorsolateral and dorsosubmedian carinae); segment IV with eight (median lateral carinae absent); segment V with five (ventromedian, paired ventrolateral and dorsolateral carinae); ventrolateral and median lateral carinae converge distally on segments I–II; ventrosubmedian and ventrolateral carinae are connected anteriorly by a transverse row of granules on segments II–IV; median lateral carinae on segment III obsolete; all carinae serrulate; dorsosubmedian carinae on segments II–IV with the distal granules well-defined and larger than the preceding; intercarinal spaces with minute dispersed granulation.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DF1F8A1CFC11" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DF1F886AFC56" box="[197,278,872,896]" italics="true" pageId="27" pageNumber="28">Telson:</emphasis>
|
||
vesicle densely granulose laterally and (primarily) ventrally, with three ventral longitudinal carinae armed with strong granulation, and one noticeably larger granule (tubercle-like) under the subaculear tubercle; subaculear tubercle strong and spine-like; aculeus long and curved.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DFA38CEBFBAA" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DFA38842FC3A" box="[197,318,980,1004]" italics="true" pageId="27" pageNumber="28">Pedipalps:</emphasis>
|
||
with fine granulation on dorsal and internal surfaces of femur; femur with five longitudinal carinae (dorsoexternal, dorsointernal, ventroexternal, ventrointernal and internal median carinae); patella without distinct carinae but with few spine-like tubercles on the position of the dorsointernal and ventrointernal carinae; chela acarinate; fixed finger with six almost linear rows of granules, being the basal the longest; movable finger with seven rows (including the short apical row). Trichobothriotaxy
|
||
<typeStatus id="821A882DB55EC773FCA0D8138A39FBAA" box="[789,837,1124,1148]" pageId="27" pageNumber="28">type</typeStatus>
|
||
A, femur with β configuration (
|
||
<bibRefCitation id="39304B7EB55EC773FB1ED8138C3FFBAA" author="Vachon" box="[1195,1347,1124,1148]" firstAuthor="Vachon" pageId="27" pageNumber="28" pagination="857 - 958" refId="ref33434" refString="Vachon, M. (1974) Etude des caracteres utilises pour classer les familles et les genres de scorpions (Arachnides). 1. La trichobothriotaxie en arachnologie. Sigles trichobothriaux et types de trichobothriotaxie chez les scorpions. Bulletin du Museum national de Histoire naturelle, Paris (3 e ser.), 104, 857 - 958." type="journal article" year="1974">Vachon 1974</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="39304B7EB55EC773FAFBD8138CF6FBAA" author="Vachon" box="[1358,1418,1124,1148]" firstAuthor="Vachon" pageId="27" pageNumber="28" pagination="1597 - 1599" refId="ref33497" refString="Vachon, M. (1975) Sur l'utilisation de la trichobothriotaxie du bras des pedipalpes des scorpions (Arachnides) dans le classement des genres de la famille des Buthidae Simon. Comptes Rendus des seances de l'Academie des Sciences, Paris (ser. D), 281, 1597 - 1599." type="journal article" year="1975">1975</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70D8FE8DDCFB12" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70D8FE887FFB76" box="[197,259,1161,1184]" italics="true" pageId="27" pageNumber="28">Legs:</emphasis>
|
||
tibia, basitarsus and telotarsus with numerous ventral setae; tibial spur present on legs III–IV; prolateral pedal spur single on legs I–II, bifid on legs III–IV; retrolateral pedal spur present on all legs.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70D8A78BA4FAA1" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70D8A788CFFB3E" box="[197,435,1232,1256]" italics="true" pageId="27" pageNumber="28">Hemispermatophore:</emphasis>
|
||
Description based on
|
||
<specimenCount id="4BA7FD06B55EC773FD1ED8A78A3FFB31" box="[683,835,1231,1255]" pageId="27" pageNumber="28" type="generic">one specimen</specimenCount>
|
||
from ICN-As-815: Flagelliform, thin and sclerotized. Foot narrow and flat. Pedal flexure inconspicuous but movable. Body long, slightly wider on basal third. Capsular region with a large, markedly sclerotized basal lobe, and an internal lobe. Flagellum with long
|
||
<emphasis id="6FD5EA9DB55EC773FB09D96E8C51FAE6" box="[1212,1325,1305,1328]" italics="true" pageId="27" pageNumber="28">pars recta</emphasis>
|
||
and long, coiled
|
||
<emphasis id="6FD5EA9DB55EC773FF57D94A881BFA82" box="[226,359,1340,1364]" italics="true" pageId="27" pageNumber="28">pars reflexa</emphasis>
|
||
(
|
||
<figureCitation id="C59A2A0AB55EC773FEC3D94B88B6FA82" box="[374,458,1340,1364]" captionStart="FIGURES 64–67" captionStartId="39.[151,264,1792,1815]" captionTargetBox="[199,1415,211,1744]" captionTargetId="figure-110@39.[151,1436,193,1771]" captionTargetPageId="39" captionText="FIGURES 64–67. Hemispermatophore in ectal aspect and detail of the capsular region in ventral aspect (left for Ananteris solimariae sp. nov., inverted; right for others). 64. Ananteris meridana González-Sponga, 2006 (ICN-As-435). 65. Ananteris myriamae Botero-Trujillo, 2007 (MPUJ-SCO-382). 66. Ananteris ochoai sp. nov., paratype (ICN-As-815). 67. Ananteris solimariae sp. nov., paratype (ICN-As-779). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5284832" httpUri="https://zenodo.org/record/5284832/files/figure.png" pageId="27" pageNumber="28">Fig. 66</figureCitation>
|
||
). Carapace length to hemispermatophore body length ratio = 1:1.28. There was no variation between both hemispermatophores examined.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70D9F48835F941" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70D9F488FBFA4A" bold="true" box="[197,391,1411,1436]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">
|
||
<typeStatus id="821A882DB55EC773FF70D9F48856FA4A" box="[197,298,1411,1436]" pageId="27" pageNumber="28" type="allotype">Allotype</typeStatus>
|
||
female.
|
||
</emphasis>
|
||
<emphasis id="6FD5EA9DB55EC773FE38D9F28B45FA4A" box="[397,569,1413,1436]" italics="true" pageId="27" pageNumber="28">Measurements:</emphasis>
|
||
see
|
||
<tableCitation id="10230334B55EC773FDDCD9F38BC3FA4A" box="[617,703,1412,1436]" captionStart="TABLE 2" captionStartId="5.[151,235,155,178]" captionText="TABLE 2. Meristic data for Ananteris species. Measurements in millimeters. 1Sum of prosoma, mesosoma and metasoma. 2Sum of tergites I–VII. 3Sum of metasomal segments I–V and telson. 4Sum of femur, patella and chela. 5Measured from the commissure of the junction with the movable finger to the finger tip. 6Measured along the marginal lamella. See for other measurements: Lourenço (1991) and Teruel & Roncallo (2008) for A. columbiana, Lourenço (1994) for A. ehrlichi, Botero-Trujillo (2007) for A. myriamae, Botero-Trujillo (2008a) for A. arcadioi and A. dorae." pageId="27" pageNumber="28">Table 2</tableCitation>
|
||
.
|
||
<figureCitation id="C59A2A0AB55EC773FD7CD9F38A2DFA4A" box="[713,849,1412,1436]" captionStart="FIGURES 24–27" captionStartId="26.[151,264,1519,1542]" captionTargetBox="[154,1433,180,1491]" captionTargetId="figure-217@26.[151,1436,174,1498]" captionTargetPageId="26" captionText="FIGURES 24–27. Ananteris ochoai sp. nov. 24–25. Holotype male, dorsal and ventral aspects (MPUJ-SCO-388). 26–27. Allotype female, dorsal and ventral aspects (MPUJ-SCO-389). Scale bars = 1 cm." figureDoi="http://doi.org/10.5281/zenodo.5284798" httpUri="https://zenodo.org/record/5284798/files/figure.png" pageId="27" pageNumber="28">Figs. 26–27</figureCitation>
|
||
. Similar to
|
||
<typeStatus id="821A882DB55EC773FC7AD9F38D4DFA4A" box="[975,1073,1412,1436]" pageId="27" pageNumber="28" type="holotype">holotype</typeStatus>
|
||
male but with the following differences deemed to be due to sexual dimorphism: body size larger; posterior median hyaline and smooth area of sternite V smaller and less evident; pectines markedly smaller, not reaching posterior end of coxae IV; telson not elongated. Aside from these primarily sexual differences, others include: general coloration darker; coxosternal region with minute brownish spots restricted to anterior half of coxapophyses I and anterior tip of coxapophyses II; anterior margin of coxae without any granulations; count of pieces on the pectines: marginal lamellae 3:3, middle lamellae 8:10, teeth 22:23, vestigial fulcra present; sternite IV with the same inconspicuous brownish areas that are found on V–VI.
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DAD48D7CF909" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DAD48836F96A" bold="true" box="[197,330,1699,1724]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Variability.</emphasis>
|
||
<emphasis id="6FD5EA9DB55EC773FEE7DAD38B4FF96A" box="[338,563,1700,1724]" italics="true" pageId="27" pageNumber="28">Pectinal teeth count</emphasis>
|
||
: 22 to 23 on females (n = 6; mode = 22); 20 to 23 on males (n = 16; mode = 21).
|
||
<emphasis id="6FD5EA9DB55EC773FF7FDABF8B27F936" box="[202,603,1736,1760]" italics="true" pageId="27" pageNumber="28">Total body length (including telson)</emphasis>
|
||
: male
|
||
<quantity id="9A599B6AB55EC773FD12DABF8A5DF909" box="[679,801,1736,1760]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" metricValueMax="3.2" metricValueMin="2.9" pageId="27" pageNumber="28" unit="mm" value="30.5" valueMax="32.0" valueMin="29.0">29–32 mm</quantity>
|
||
, female
|
||
<quantity id="9A599B6AB55EC773FC35DABF8A80F909" box="[896,1020,1736,1760]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.3" metricValueMin="4.1" pageId="27" pageNumber="28" unit="mm" value="42.0" valueMax="43.0" valueMin="41.0">41–43 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="15BB6504B55EC773FF70DA9C8CE7F845" pageId="27" pageNumber="28" type="distribution">
|
||
<paragraph id="5D1E368FB55EC773FF70DA9C8AA1F8FE" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DA9C8822F8D2" bold="true" box="[197,350,1771,1796]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="821A882DB55EC773FDC5DA9B8BDCF8D2" box="[624,672,1772,1796]" pageId="27" pageNumber="28">type</typeStatus>
|
||
locality in Barbacoas municipality (
|
||
<collectingRegion id="9F65F86DB55EC773FBFDDA9B8C50F8D2" box="[1096,1324,1772,1796]" country="Colombia" name="Narino" pageId="27" pageNumber="28">Nariño Department</collectingRegion>
|
||
), extreme southwestern
|
||
<collectingCountry id="25B6761FB55EC773FE87DB6788DCF8FE" box="[306,416,1808,1832]" name="Colombia" pageId="27" pageNumber="28">Colombia</collectingCountry>
|
||
in the western slope of the Andes (
|
||
<figureCitation id="C59A2A0AB55EC773FC9EDB678A05F8FE" box="[811,889,1808,1832]" captionStart="FIGURE 1" captionStartId="6.[151,250,1822,1845]" captionTargetBox="[177,1415,197,1775]" captionTargetId="figure-117@6.[174,1418,195,1778]" captionTargetPageId="6" captionText="FIGURE 1. Known distribution of Colombian Ananteris. () Ananteris arcadioi Botero-Trujillo, 2008. () Ananteris columbiana Lourenço, 1991. () Ananteris dorae Botero-Trujillo, 2008. () A. dorae and Ananteris ochoai sp. nov. ()Ananteris ehrlichi Lourenço, 1994. () Ananteris faguasi Botero-Trujillo, 2009. () Ananteris gorgonae Lourenço & Flórez, 1989. () Ananteris leilae Lourenço, 1999. () Ananteris meridana González-Sponga, 2006 (also type locality in Venezuela). () Ananteris myriamae Botero-Trujillo, 2007. () Ananteris solimariae sp. nov. ()Ananteris tolimana Teruel & García, 2007. () Ananteris volschenki Botero-Trujillo, 2009." figureDoi="http://doi.org/10.5281/zenodo.5284778" httpUri="https://zenodo.org/record/5284778/files/figure.png" pageId="27" pageNumber="28">Figs. 1</figureCitation>
|
||
,
|
||
<figureCitation id="C59A2A0AB55EC773FC32DB678AADF8FE" box="[903,977,1808,1832]" captionStart="FIGURES 56–57" captionStartId="35.[151,263,1091,1114]" captionTargetBox="[152,1435,198,1061]" captionTargetId="figure-698@35.[151,1436,193,1070]" captionTargetPageId="35" captionText="FIGURES 56–57. Two views of the collection locality of Ananteris ochoai sp. nov. and Ananteris dorae Botero-Trujillo, 2008 in Río Ñambi Natural Reserve, Nariño Department. Photographs by José A. Ochoa." figureDoi="http://doi.org/10.5281/zenodo.5284822" httpUri="https://zenodo.org/record/5284822/files/figure.png" pageId="27" pageNumber="28">56–57</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="5D1E368FB55EC773FF70DB448CE7F845" blockId="27.[151,1437,151,1940]" pageId="27" pageNumber="28">
|
||
<emphasis id="6FD5EA9DB55EC773FF70DB4488F2F89A" bold="true" box="[197,398,1843,1868]" inLineHeading="true" pageId="27" pageNumber="28" reason="1">Ecological notes.</emphasis>
|
||
All specimens of
|
||
<taxonomicName id="9AA14D0CB55EC773FDD1DB438BAEF89A" authority="Botero-Trujillo & Flórez, 2011" authorityName="Botero-Trujillo & Flórez" authorityYear="2011" box="[612,722,1844,1868]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ochoai" status="sp. nov.">
|
||
<emphasis id="6FD5EA9DB55EC773FDD1DB438BAEF89A" box="[612,722,1844,1868]" italics="true" pageId="27" pageNumber="28">A. ochoai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6FD5EA9DB55EC773FD69DB438A4BF89A" bold="true" box="[732,823,1844,1868]" pageId="27" pageNumber="28">
|
||
<taxonomicNameLabel id="74E657E6B55EC773FD69DB438A4BF89A" box="[732,823,1844,1868]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
were hand-captured at night after detection with UV light. Some individuals were detected when actively moving in the leaf litter, whereas others were found under decomposing logs.
|
||
<taxonomicName id="9AA14D0CB55EC773FEC5DB0B8B4CF842" authority="Botero-Trujillo & Flórez, 2011" authorityName="Botero-Trujillo & Flórez" authorityYear="2011" box="[368,560,1916,1940]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="ochoai" status="sp. nov.">
|
||
<emphasis id="6FD5EA9DB55EC773FEC5DB0B8B4CF842" box="[368,560,1916,1940]" italics="true" pageId="27" pageNumber="28">Ananteris ochoai</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="6FD5EA9DB55EC773FD82DB0B8BECF842" bold="true" box="[567,656,1916,1940]" pageId="27" pageNumber="28">
|
||
<taxonomicNameLabel id="74E657E6B55EC773FD82DB0B8BECF842" box="[567,656,1916,1940]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
was found in sympatry with
|
||
<taxonomicName id="9AA14D0CB55EC773FC62DB0B8D4BF842" authorityName="Botero-Trujillo" authorityYear="2008" box="[983,1079,1916,1940]" class="Arachnida" family="Buthidae" genus="Ananteris" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="dorae">
|
||
<emphasis id="6FD5EA9DB55EC773FC62DB0B8D4BF842" box="[983,1079,1916,1940]" italics="true" pageId="27" pageNumber="28">A. dorae</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="9AA14D0CB55EC773FBF0DB0A8DD3F845" box="[1093,1199,1916,1940]" class="Arachnida" family="Buthidae" genus="Tityus" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="6FD5EA9DB55EC773FBF0DB0A8DF4F842" box="[1093,1160,1917,1940]" italics="true" pageId="27" pageNumber="28">Tityus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="9AA14D0CB55EC773FB50DB0B8CE7F845" box="[1253,1435,1916,1940]" class="Arachnida" family="Chactidae" genus="Teuthraustes" kingdom="Animalia" order="Scorpiones" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="6FD5EA9DB55EC773FB50DB0B8C0FF842" box="[1253,1395,1916,1940]" italics="true" pageId="27" pageNumber="28">Teuthraustes</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |