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<document id="252777815C56CB531C3EDCE95EB93221" ID-CLB-Dataset="5418" ID-DOI="10.11646/zootaxa.3841.1.7" ID-GBIF-Dataset="7ca3efd1-4b06-47cf-a9cf-3985b60632f7" ID-ISSN="1175-5326" ID-Zenodo-Dep="4928200" ID-ZooBank="6FB24193-A0DB-4494-8C2C-F7591E614E9C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623432904869" checkinUser="felipe" docAuthor="Muadsub, Sopita &amp; Pinkaew, Nantasak" docDate="2014" docId="BB7D9906FFF43E2820AFCF02690CF969" docLanguage="en" docName="zt03841p134.pdf" docOrigin="Zootaxa 3841 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Helictophanes flava Muadsub and Pinkaew 2014, n. sp." docType="treatment" docVersion="7" lastPageNumber="130" masterDocId="4744E17EFFF53E2B2038CD3D6B15FFD4" masterDocTitle="Two new species of Enarmoniini (Lepidoptera: Tortricidae) from eastern Thailand" masterLastPageNumber="134" masterPageNumber="127" pageNumber="128" updateTime="1699010613421" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5585F2045E227F55174A2BB2DE0D23C2">Two new species of Enarmoniini (Lepidoptera: Tortricidae) from eastern Thailand</mods:title>
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<mods:name id="5FC63EB4904CE704BDE78F110BF2F756" type="personal">
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<mods:namePart id="893B95F454E97B8120145B8B190D552C">Muadsub, Sopita</mods:namePart>
<mods:affiliation id="3EAAB9FEC4BD4C6152A06C4957E0A0D0">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Nakhon Pathom, 73140, Thailand. E-mail: sun _ svbee @ hotmail. com</mods:affiliation>
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<mods:namePart id="433ADA28C5F1FBA0BED1C17CB8786A87">Pinkaew, Nantasak</mods:namePart>
<mods:affiliation id="D79D3847D1A580F930D501F53AE38123">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Nakhon Pathom, 73140, Thailand. E-mail: sun _ svbee @ hotmail. com &amp; Center for Advanced Studies in Tropical Natural Resources, NRU-KU, Kasetsart University, Chatuchak, Bangkok, 10900, Thailand. E-mail: agrnsp @ ku. ac. th</mods:affiliation>
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<mods:title id="797D74CCF4BBA7B429BFB8CF41F25B3E">Zootaxa</mods:title>
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<treatment id="BB7D9906FFF43E2820AFCF02690CF969" ID-DOI="http://doi.org/10.5281/zenodo.4927931" ID-GBIF-Taxon="183550744" ID-Zenodo-Dep="4927931" LSID="urn:lsid:plazi:treatment:BB7D9906FFF43E2820AFCF02690CF969" httpUri="http://treatment.plazi.org/id/BB7D9906FFF43E2820AFCF02690CF969" lastPageId="3" lastPageNumber="130" pageId="1" pageNumber="128">
<subSubSection id="7BCE7B9BFFF43E2A20AFCF026815FD8E" box="[151,768,575,602]" pageId="1" pageNumber="128" type="nomenclature">
<paragraph id="336B2810FFF43E2A20AFCF026815FD8E" blockId="1.[151,768,575,635]" box="[151,768,575,602]" pageId="1" pageNumber="128">
<heading id="68239F7CFFF43E2A20AFCF026815FD8E" bold="true" box="[151,768,575,602]" fontSize="11" level="1" pageId="1" pageNumber="128" reason="1">
<emphasis id="01A0F402FFF43E2A20AFCF026815FD8E" bold="true" box="[151,768,575,602]" pageId="1" pageNumber="128">
<taxonomicName id="F4D45393FFF43E2A20AFCF0269A4FD8E" ID-CoL="B565Z" authority="Muadsub and Pinkaew" authorityName="Muadsub and Pinkaew" authorityYear="2014" box="[151,689,575,602]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="flava" status="sp. nov.">
<emphasis id="01A0F402FFF43E2A20AFCF026A9DFD8D" bold="true" box="[151,392,575,601]" italics="true" pageId="1" pageNumber="128">Helictophanes flava</emphasis>
Muadsub and Pinkaew
</taxonomicName>
,
<taxonomicNameLabel id="1A934979FFF43E2A2285CF7D6815FD8E" box="[701,768,576,602]" pageId="1" pageNumber="128" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="7BCE7B9BFFF43E2A20AFCF5F6AFFFDAF" box="[151,490,610,635]" pageId="1" pageNumber="128" type="description">
<paragraph id="336B2810FFF43E2A20AFCF5F6AFFFDAF" blockId="1.[151,768,575,635]" box="[151,490,610,635]" pageId="1" pageNumber="128">
(
<figureCitation id="ABEF3495FFF43E2A20A7CF5F6A36FDAF" box="[159,291,610,635]" captionStart="FIGURE 1" captionStartId="2.[151,250,1942,1964]" captionTargetBox="[395,1127,195,1904]" captionTargetId="figure-70@2.[395,1127,195,1907]" captionTargetPageId="2" captionText="FIGURE 1. Adults of Helictophanes and Cyphophanes. A. Helictophanes flava, n. sp. (holotype male). B. H. flava, n. sp. (paratype female). C. Cyphophanes khitchakutensis, n. sp. (holotype male). D. C. khitchakutensis, n. sp. (paratype female) (scale bar=2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4928202" httpUri="https://zenodo.org/record/4928202/files/figure.png" pageId="1" pageNumber="128">Figs. 1AB</figureCitation>
,
<figureCitation id="ABEF3495FFF43E2A2117CF5F6A97FDAF" box="[303,386,610,635]" captionStart="FIGURE 2" captionStartId="3.[151,250,1428,1450]" captionTargetBox="[172,1426,211,1364]" captionTargetId="figure-202@3.[151,1436,193,1406]" captionTargetPageId="3" captionText="FIGURE 2. Labial palpi and wing venation. A-B. Labial palpi; A. Helictophanes flava, n. sp. B. Cyphophanes khitchakutensis, n. sp. C-D. Wing venation; C. H. flava, n. sp. D. C. khitchakutensis, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4928204" httpUri="https://zenodo.org/record/4928204/files/figure.png" pageId="1" pageNumber="128">2A, 2C</figureCitation>
,
<figureCitation id="ABEF3495FFF43E2A21B6CF5F6AA7FDAF" box="[398,434,610,635]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[208,1382,214,1871]" captionTargetId="figure-47@4.[167,1419,193,1932]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia. A. Helictophanes flava, n. sp. (holotype male). B. Cyphophanes khitchakutensis, n. sp. (holotype male)." figureDoi="http://doi.org/10.5281/zenodo.4928206" httpUri="https://zenodo.org/record/4928206/files/figure.png" pageId="1" pageNumber="128">3A</figureCitation>
,
<figureCitation id="ABEF3495FFF43E2A2186CF5F6AF7FDAF" box="[446,482,610,635]" captionStart="FIGURE 4" captionStartId="5.[151,250,1800,1822]" captionTargetBox="[160,1387,208,1730]" captionTargetId="figure-121@5.[151,1436,193,1778]" captionTargetPageId="5" captionText="FIGURE 4. Female genitalia. A. Helictophanes flava, n. sp. (paratype female). B. Cyphophanes khitchakutensis, n. sp. (paratype female)." figureDoi="http://doi.org/10.5281/zenodo.4928210" httpUri="https://zenodo.org/record/4928210/files/figure.png" pageId="1" pageNumber="128">4A</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="7BCE7B9BFFF43E2A20AFCF996FB7FC35" pageId="1" pageNumber="128" type="diagnosis">
<paragraph id="336B2810FFF43E2A20AFCF996FB7FC35" blockId="1.[151,1437,676,2033]" pageId="1" pageNumber="128">
<emphasis id="01A0F402FFF43E2A20AFCF996A04FD69" box="[151,273,676,701]" inLineHeading="true" pageId="1" pageNumber="128" reason="1">
<emphasis id="01A0F402FFF43E2A20AFCF996A1FFD69" bold="true" box="[151,266,676,701]" pageId="1" pageNumber="128">Diagnosis</emphasis>
.
</emphasis>
This species is characterized by the following features: forewing yellow with a dark triangular dorsal mark and an irregular, curved band from the middle of the costa to the termen; hindwing veins M
<subScript id="AF502A55FFF43E2A24DECFE56FFBFD32" attach="left" box="[1254,1262,728,742]" fontSize="6" pageId="1" pageNumber="128">3</subScript>
and CuA
<subScript id="AF502A55FFF43E2A2563CFE56E76FD32" attach="left" box="[1371,1379,728,742]" fontSize="6" pageId="1" pageNumber="128">1</subScript>
long stalked; male hindwing with a long ribbon-shaped anal lobe nearly reaching the anal angle; and short, nearly straight labial palpi with a small, very short terminal segment. The male genitalia are characterized by a narrow apex of the tegumen with dense scale sockets laterally; and by a valva with a distinct, curved neck and an elongate elliptical cucullus with a very short apical spine. The female genitalia are characterized by a ring-shaped sterigma, incomplete posteriorly; a very short ductus bursae; and a corpus bursae with two large triangular, bladelike signa with elongate hollow bases and a sclerotized plate posteriorly.
<emphasis id="01A0F402FFF43E2A2357CE9868C1FC68" box="[879,980,932,956]" italics="true" pageId="1" pageNumber="128">
<taxonomicName id="F4D45393FFF43E2A2357CE9868C5FC68" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[879,976,932,956]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="species" species="flava" status="sp. nov.">H. flava</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="1A934979FFF43E2A23E6CE986F06FC68" box="[990,1043,933,956]" pageId="1" pageNumber="128" rank="species">n.sp.</taxonomicNameLabel>
differs from previously described species of
<taxonomicName id="F4D45393FFF43E2A2135CEF56ABAFC34" authorityName="Meyrick" authorityYear="1881" box="[269,431,968,992]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="01A0F402FFF43E2A2135CEF56ABAFC34" box="[269,431,968,992]" italics="true" pageId="1" pageNumber="128">Helictophanes</emphasis>
</taxonomicName>
by the presence of a sclerite on the posterior area of corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="7BCE7B9BFFF43E2A20FFCED66F32F825" pageId="1" pageNumber="128" type="description">
<paragraph id="336B2810FFF43E2A20FFCED669E2FB41" blockId="1.[151,1437,676,2033]" pageId="1" pageNumber="128">
<emphasis id="01A0F402FFF43E2A20FFCED66A4DFBD0" box="[199,344,1003,1028]" inLineHeading="true" pageId="1" pageNumber="128" reason="1">
<emphasis id="01A0F402FFF43E2A20FFCED66A44FBD0" bold="true" box="[199,337,1003,1028]" pageId="1" pageNumber="128">Description</emphasis>
.
</emphasis>
Head. Ocellus large, vertex covered with golden yellow scales, upper frons golden yellow, with pale yellow, appressed scales on lower frons; labial palpus rather short (1.5X diameter of eye) (
<figureCitation id="ABEF3495FFF43E2A24EDC92D6E24FBFD" box="[1237,1329,1040,1065]" captionStart="FIGURE 2" captionStartId="3.[151,250,1428,1450]" captionTargetBox="[172,1426,211,1364]" captionTargetId="figure-202@3.[151,1436,193,1406]" captionTargetPageId="3" captionText="FIGURE 2. Labial palpi and wing venation. A-B. Labial palpi; A. Helictophanes flava, n. sp. B. Cyphophanes khitchakutensis, n. sp. C-D. Wing venation; C. H. flava, n. sp. D. C. khitchakutensis, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4928204" httpUri="https://zenodo.org/record/4928204/files/figure.png" pageId="1" pageNumber="128">Fig. 2A</figureCitation>
), porrect, yellow, with brown to dark brown scales dorsobasally on second segment, first segment short, second segment long and nearly straight, gradually widening toward apex, terminal segment short, apex acute, pointing downward; antenna to beyond middle of forewing, golden yellow.
</paragraph>
<paragraph id="336B2810FFF43E2A20FFC99D6AC5F9F0" blockId="1.[151,1437,676,2033]" pageId="1" pageNumber="128">
Thorax. Smooth, without raised scale tuft posteriorly, light brown; legs unmodified. Forewings subrectangular, length 2.9-3.0 mm in male (n = 2) (
<figureCitation id="ABEF3495FFF43E2A2213C9FE6996FB08" box="[555,643,1219,1244]" captionStart="FIGURE 1" captionStartId="2.[151,250,1942,1964]" captionTargetBox="[395,1127,195,1904]" captionTargetId="figure-70@2.[395,1127,195,1907]" captionTargetPageId="2" captionText="FIGURE 1. Adults of Helictophanes and Cyphophanes. A. Helictophanes flava, n. sp. (holotype male). B. H. flava, n. sp. (paratype female). C. Cyphophanes khitchakutensis, n. sp. (holotype male). D. C. khitchakutensis, n. sp. (paratype female) (scale bar=2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4928202" httpUri="https://zenodo.org/record/4928202/files/figure.png" pageId="1" pageNumber="128">Fig. 1A</figureCitation>
)
<quantity id="F42C85F5FFF43E2A22ACC9FE6809FB0F" box="[660,796,1219,1244]" metricMagnitude="-3" metricUnit="m" metricValue="3.8499999999999996" metricValueMax="3.9" metricValueMin="3.8" pageId="1" pageNumber="128" unit="mm" value="3.8499999999999996" valueMax="3.9" valueMin="3.8">3.83.9 mm</quantity>
in female (n = 2) (
<figureCitation id="ABEF3495FFF43E2A23CDC9FE6F58FB08" box="[1013,1101,1219,1244]" captionStart="FIGURE 1" captionStartId="2.[151,250,1942,1964]" captionTargetBox="[395,1127,195,1904]" captionTargetId="figure-70@2.[395,1127,195,1907]" captionTargetPageId="2" captionText="FIGURE 1. Adults of Helictophanes and Cyphophanes. A. Helictophanes flava, n. sp. (holotype male). B. H. flava, n. sp. (paratype female). C. Cyphophanes khitchakutensis, n. sp. (holotype male). D. C. khitchakutensis, n. sp. (paratype female) (scale bar=2 mm)." figureDoi="http://doi.org/10.5281/zenodo.4928202" httpUri="https://zenodo.org/record/4928202/files/figure.png" pageId="1" pageNumber="128">Fig. 1B</figureCitation>
); without costal fold in male, costa curved near base and before apex, apex rounded, termen oblique and nearly straight; forewing venation with all R veins separate, R
<subScript id="AF502A55FFF43E2A21A8C8246A8DFAF3" attach="left" box="[400,408,1305,1319]" fontSize="6" pageId="1" pageNumber="128">5</subScript>
reaching to termen below apex, M
<subScript id="AF502A55FFF43E2A2324C8246831FAF3" attach="left" box="[796,804,1305,1319]" fontSize="6" pageId="1" pageNumber="128">3</subScript>
and CuA
<subScript id="AF502A55FFF43E2A23B7C8246882FAF3" attach="left" box="[911,919,1305,1319]" fontSize="6" pageId="1" pageNumber="128">1</subScript>
parallel and very close in basal half (
<figureCitation id="ABEF3495FFF43E2A250EC8376E9AFAF7" box="[1334,1423,1290,1315]" captionStart="FIGURE 2" captionStartId="3.[151,250,1428,1450]" captionTargetBox="[172,1426,211,1364]" captionTargetId="figure-202@3.[151,1436,193,1406]" captionTargetPageId="3" captionText="FIGURE 2. Labial palpi and wing venation. A-B. Labial palpi; A. Helictophanes flava, n. sp. B. Cyphophanes khitchakutensis, n. sp. C-D. Wing venation; C. H. flava, n. sp. D. C. khitchakutensis, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4928204" httpUri="https://zenodo.org/record/4928204/files/figure.png" pageId="1" pageNumber="128">Fig. 2C</figureCitation>
); ground color pale yellow, with six dark brown spots along costa, three basally and the other three distally, and a long narrow dark brown band medially, curved and extending from costa to termen at M
<subScript id="AF502A55FFF43E2A24B4C85E6F81FAA5" attach="right" box="[1164,1172,1379,1393]" fontSize="6" pageId="1" pageNumber="128">1</subScript>
, brown to dark brown; dorsum with large, triangular brown mark medially, edged with dark brown scales; underside pale brown, speckled paler reflecting wing pattern. Hindwing narrow, apex pointed, with long ribbon-shaped anal lobe in male, nearly reaching anal angle; wing venation with M
<subScript id="AF502A55FFF43E2A2241C8ED6994FA0A" attach="left" box="[633,641,1488,1502]" fontSize="6" pageId="1" pageNumber="128">3</subScript>
and CuA
<subScript id="AF502A55FFF43E2A22D4C8ED69E1FA0A" attach="left" box="[748,756,1488,1502]" fontSize="6" pageId="1" pageNumber="128">1</subScript>
connate to long-stalked, without trace of CuP, 3A extending along basal half of anal margin, half as long as 1A+2A (
<figureCitation id="ABEF3495FFF43E2A2312C8DA6893F9D4" box="[810,902,1511,1536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1428,1450]" captionTargetBox="[172,1426,211,1364]" captionTargetId="figure-202@3.[151,1436,193,1406]" captionTargetPageId="3" captionText="FIGURE 2. Labial palpi and wing venation. A-B. Labial palpi; A. Helictophanes flava, n. sp. B. Cyphophanes khitchakutensis, n. sp. C-D. Wing venation; C. H. flava, n. sp. D. C. khitchakutensis, n. sp." figureDoi="http://doi.org/10.5281/zenodo.4928204" httpUri="https://zenodo.org/record/4928204/files/figure.png" pageId="1" pageNumber="128">Fig. 2C</figureCitation>
); upper side translucent, brown, paler towards base; underside pale brown.
</paragraph>
<paragraph id="336B2810FFF43E2A20FFCB136F32F825" blockId="1.[151,1437,676,2033]" pageId="1" pageNumber="128">
Abdomen. Male genitalia (
<figureCitation id="ABEF3495FFF43E2A21CBCB136958F993" box="[499,589,1582,1607]" captionStart="FIGURE 3" captionStartId="4.[151,250,1953,1975]" captionTargetBox="[208,1382,214,1871]" captionTargetId="figure-47@4.[167,1419,193,1932]" captionTargetPageId="4" captionText="FIGURE 3. Male genitalia. A. Helictophanes flava, n. sp. (holotype male). B. Cyphophanes khitchakutensis, n. sp. (holotype male)." figureDoi="http://doi.org/10.5281/zenodo.4928206" httpUri="https://zenodo.org/record/4928206/files/figure.png" pageId="1" pageNumber="128">Fig. 3A</figureCitation>
). Tegumen moderately high, weakly sclerotized, dorsolaterally with narrow lobe with dense sockets; socius elongate membranous, sparsely covered with hairs; gnathos represented by a weak sclerotized band, arising from dorsal 1/3 of tegumen; vinculum small; valva moderately sclerotized, with large basal excavation, costa curved; sacculus large, subrectangular, with moderately dense long setae apicoventrally; with deep ventral excavation; cucullus long, slender, elliptic to band-shaped, slightly widening beyond neck, rounded apically with small, single dorsoapical spine; juxta small; caulis rather long; phallus cylindrical, straight, moderately long and wide, slightly tapering to apex; with numerous lanceolate cornuti, nearly as long as phallus. Female genitalia (
<figureCitation id="ABEF3495FFF43E2A2150CA1A6AD3F894" box="[360,454,1831,1856]" captionStart="FIGURE 4" captionStartId="5.[151,250,1800,1822]" captionTargetBox="[160,1387,208,1730]" captionTargetId="figure-121@5.[151,1436,193,1778]" captionTargetPageId="5" captionText="FIGURE 4. Female genitalia. A. Helictophanes flava, n. sp. (paratype female). B. Cyphophanes khitchakutensis, n. sp. (paratype female)." figureDoi="http://doi.org/10.5281/zenodo.4928210" httpUri="https://zenodo.org/record/4928210/files/figure.png" pageId="1" pageNumber="128">Fig. 4A</figureCitation>
). Papillae anales with dense setae. Tergum VIII smooth except for lateral triangular extensions with sparse scale sockets; sternum VII weakly sclerotized, moderately scaled, sterigma beyond posterior margin of sternum VII, subcircular ridge, wider laterally, with dense microtrichia around ostium; colliculum small, irregular ring, moderately sclerotized; ductus bursae short and wide, ductus seminalis arising from lobe near bursae neck with elongate and sinuate sclerite on posterior area end of corpus bursae; corpus bursae subovate with two large, unequal signa: triangular blades with long hollow bases.
</paragraph>
</subSubSection>
<caption id="67AB7898FFF73E2920AFCAAB6A44F83E" ID-DOI="http://doi.org/10.5281/zenodo.4928202" ID-Zenodo-Dep="4928202" httpUri="https://zenodo.org/record/4928202/files/figure.png" pageId="2" pageNumber="129" startId="2.[151,250,1942,1964]" targetBox="[395,1127,195,1904]" targetPageId="2">
<paragraph id="336B2810FFF73E2920AFCAAB6A44F83E" blockId="2.[151,1436,1942,2026]" pageId="2" pageNumber="129">
<emphasis id="01A0F402FFF73E2920AFCAAB6A00F878" bold="true" box="[151,277,1942,1964]" pageId="2" pageNumber="129">FIGURE 1.</emphasis>
Adults of
<taxonomicName id="F4D45393FFF73E2921B0CAAB690FF878" authorityName="Meyrick" authorityYear="1881" box="[392,538,1942,1964]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="01A0F402FFF73E2921B0CAAB690FF878" box="[392,538,1942,1964]" italics="true" pageId="2" pageNumber="129">Helictophanes</emphasis>
</taxonomicName>
and
<taxonomicName id="F4D45393FFF73E292268CAAB69CFF878" authorityName="Meyrick" authorityYear="1937" box="[592,730,1942,1964]" class="Insecta" family="Tortricidae" genus="Cyphophanes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="genus">
<emphasis id="01A0F402FFF73E292268CAAB69CFF878" box="[592,730,1942,1964]" italics="true" pageId="2" pageNumber="129">Cyphophanes</emphasis>
</taxonomicName>
. A.
<taxonomicName id="F4D45393FFF73E292332CAAB68CDF878" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[778,984,1942,1964]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="flava" status="sp. nov.">
<emphasis id="01A0F402FFF73E292332CAAB68CDF878" box="[778,984,1942,1964]" italics="true" pageId="2" pageNumber="129">Helictophanes flava</emphasis>
</taxonomicName>
,
<emphasis id="01A0F402FFF73E2923DFCAAA6F37F878" bold="true" box="[999,1058,1943,1964]" pageId="2" pageNumber="129">
<taxonomicNameLabel id="1A934979FFF73E2923DFCAAA6F37F878" box="[999,1058,1943,1964]" pageId="2" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype male). B.
<taxonomicName id="F4D45393FFF73E2924C5CAAA6E47F878" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[1277,1362,1942,1964]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="flava" status="sp. nov.">
<emphasis id="01A0F402FFF73E2924C5CAAA6E47F878" box="[1277,1362,1942,1964]" italics="true" pageId="2" pageNumber="129">H. flava</emphasis>
</taxonomicName>
,
<emphasis id="01A0F402FFF73E292558CAAA6E89F878" bold="true" box="[1376,1436,1943,1964]" pageId="2" pageNumber="129">
<taxonomicNameLabel id="1A934979FFF73E292558CAAA6E89F878" box="[1376,1436,1943,1964]" pageId="2" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(paratype female). C.
<taxonomicName id="F4D45393FFF73E292144CA8869B9F81F" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[380,684,1973,1995]" class="Insecta" family="Tortricidae" genus="Cyphophanes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="khitchakutensis" status="sp. nov.">
<emphasis id="01A0F402FFF73E292144CA8869B9F81F" box="[380,684,1973,1995]" italics="true" pageId="2" pageNumber="129">Cyphophanes khitchakutensis</emphasis>
</taxonomicName>
,
<emphasis id="01A0F402FFF73E292283CA8B69E2F81F" bold="true" box="[699,759,1974,1995]" pageId="2" pageNumber="129">
<taxonomicNameLabel id="1A934979FFF73E292283CA8B69E2F81F" box="[699,759,1974,1995]" pageId="2" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(holotype male). D.
<taxonomicName id="F4D45393FFF73E2923EDCA886F84F81F" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[981,1169,1973,1995]" class="Insecta" family="Tortricidae" genus="Cyphophanes" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="129" phylum="Arthropoda" rank="species" species="khitchakutensis" status="sp. nov.">
<emphasis id="01A0F402FFF73E2923EDCA886F84F81F" box="[981,1169,1973,1995]" italics="true" pageId="2" pageNumber="129">C. khitchakutensis</emphasis>
</taxonomicName>
,
<emphasis id="01A0F402FFF73E292499CA8B6FC8F81F" bold="true" box="[1185,1245,1974,1995]" pageId="2" pageNumber="129">
<taxonomicNameLabel id="1A934979FFF73E292499CA8B6FC8F81F" box="[1185,1245,1974,1995]" pageId="2" pageNumber="129" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
(paratype female) (scale bar=2 mm).
</paragraph>
</caption>
<caption id="67AB7898FFF63E2820AFC8A9689FFA1C" ID-DOI="http://doi.org/10.5281/zenodo.4928204" ID-Zenodo-Dep="4928204" httpUri="https://zenodo.org/record/4928204/files/figure.png" pageId="3" pageNumber="130" startId="3.[151,250,1428,1450]" targetBox="[172,1426,211,1364]" targetPageId="3">
<paragraph id="336B2810FFF63E2820AFC8A9689FFA1C" blockId="3.[151,1435,1428,1481]" pageId="3" pageNumber="130">
<emphasis id="01A0F402FFF63E2820AFC8A96A07FA7D" bold="true" box="[151,274,1428,1450]" pageId="3" pageNumber="130">FIGURE 2.</emphasis>
Labial palpi and wing venation. A-B. Labial palpi; A.
<taxonomicName id="F4D45393FFF63E282300C8A96F16FA7E" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[824,1027,1428,1450]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="flava" status="sp. nov.">
<emphasis id="01A0F402FFF63E282300C8A96F16FA7E" box="[824,1027,1428,1450]" italics="true" pageId="3" pageNumber="130">Helictophanes flava</emphasis>
</taxonomicName>
,
<emphasis id="01A0F402FFF63E282437C8A96F5DFA7D" bold="true" box="[1039,1096,1428,1449]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="1A934979FFF63E282437C8A96F5DFA7D" box="[1039,1096,1428,1449]" pageId="3" pageNumber="130" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
B.
<taxonomicName id="F4D45393FFF63E282452C8A96E83FA7E" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[1130,1430,1428,1450]" class="Insecta" family="Tortricidae" genus="Cyphophanes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="khitchakutensis" status="sp. nov.">
<emphasis id="01A0F402FFF63E282452C8A96E83FA7E" box="[1130,1430,1428,1450]" italics="true" pageId="3" pageNumber="130">Cyphophanes khitchakutensis</emphasis>
</taxonomicName>
,
<emphasis id="01A0F402FFF63E2820AFC88E6BC5FA1C" bold="true" box="[151,208,1459,1480]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="1A934979FFF63E2820AFC88E6BC5FA1C" box="[151,208,1459,1480]" pageId="3" pageNumber="130" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
C-D. Wing venation; C.
<taxonomicName id="F4D45393FFF63E2821F6C889690AFA1D" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[462,543,1459,1481]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="flava" status="sp. nov.">
<emphasis id="01A0F402FFF63E2821F6C889690AFA1D" box="[462,543,1459,1481]" italics="true" pageId="3" pageNumber="130">H. flava</emphasis>
</taxonomicName>
,
<emphasis id="01A0F402FFF63E282214C88E6970FA1C" bold="true" box="[556,613,1459,1480]" pageId="3" pageNumber="130">
<taxonomicNameLabel id="1A934979FFF63E282214C88E6970FA1C" box="[556,613,1459,1480]" pageId="3" pageNumber="130" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
D.
<taxonomicName id="F4D45393FFF63E2822B2C88E6851FA1D" authority="Muadsub &amp; Pinkaew, 2014" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[650,836,1459,1481]" class="Insecta" family="Tortricidae" genus="Cyphophanes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="khitchakutensis" status="sp. nov.">
<emphasis id="01A0F402FFF63E2822B2C88E6851FA1D" box="[650,836,1459,1481]" italics="true" pageId="3" pageNumber="130">C. khitchakutensis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="1A934979FFF63E282369C88E689FFA1C" box="[849,906,1459,1480]" pageId="3" pageNumber="130" rank="species">n. sp.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="7BCE7B9BFFF63E2820FFC8CD6E74F9A7" pageId="3" pageNumber="130" type="materials_examined">
<paragraph id="336B2810FFF63E2820FFC8CD6887F9FF" blockId="3.[151,1436,1520,1725]" pageId="3" pageNumber="130">
<materialsCitation id="83BC224DFFF63E2820FFC8CD6887F9FF" ID-GBIF-Occurrence="3323415301" collectingDate="2012-10-12" collectionCode="R, BMNH" collectorName="N. Pinkaew &amp; Deposited" country="Thailand" elevation="30" latitude="12.395278" location="Agroforestry" longLatPrecision="21" longitude="102.67555" pageId="3" pageNumber="130" specimenCount="1" specimenCount-male="1" stateProvince="Trat" typeStatus="holotype">
<emphasis id="01A0F402FFF63E2820FFC8CD6A21F9DC" bold="true" box="[199,308,1520,1544]" pageId="3" pageNumber="130">
<typeStatus id="EC6F96B2FFF63E2820FFC8CD6A21F9DC" box="[199,308,1520,1544]" pageId="3" pageNumber="130" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="25D2E399FFF63E282104C8CC6A41F9DC" box="[316,340,1521,1544]" pageId="3" pageNumber="130" type="male"></specimenCount>
.
<collectingCountry id="4BC36880FFF63E282159C8CD6ADCF9DD" box="[353,457,1520,1545]" name="Thailand" pageId="3" pageNumber="130">Thailand</collectingCountry>
:
<collectingRegion id="F110E6F2FFF63E2821EDC8CC695CF9DC" box="[469,585,1521,1544]" country="Thailand" name="Trat" pageId="3" pageNumber="130">Trat Prov.</collectingRegion>
:
<collectingRegion id="F110E6F2FFF63E28226DC8CC6996F9DC" box="[597,643,1521,1544]" country="Thailand" name="Trat" pageId="3" pageNumber="130">Trat</collectingRegion>
<location id="360B7ECBFFF63E2822B3C8CD6809F9DD" LSID="urn:lsid:plazi:treatment:BB7D9906FFF43E2820AFCF02690CF969:360B7ECBFFF63E2822B3C8CD6809F9DD" box="[651,796,1520,1545]" country="Thailand" latitude="12.395278" longLatPrecision="21" longitude="102.67555" name="Agroforestry" pageId="3" pageNumber="130" stateProvince="Trat">Agroforestry</location>
<collectionCode id="55C5B0D5FFF63E28231CC8CD682EF9DD" box="[804,827,1520,1545]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="130">R</collectionCode>
.St.,
<geoCoordinate id="56E04ED7FFF63E282353C8CD68FAF9DD" box="[875,1007,1520,1545]" degrees="12" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="130" precision="15" seconds="43" value="12.395278">12°23'43&quot;N</geoCoordinate>
<geoCoordinate id="56E04ED7FFF63E2823CFC8CD6F9CF9DD" box="[1015,1161,1520,1545]" degrees="102" direction="east" minutes="40" orientation="longitude" pageId="3" pageNumber="130" precision="15" seconds="32" value="102.67555">102°40'32&quot;E</geoCoordinate>
, ca.
<quantity id="F42C85F5FFF63E282486C8CD6FEFF9DC" box="[1214,1274,1520,1545]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="130" unit="m" value="30.0">
<elevation id="B8F9CF23FFF63E282486C8CD6FEFF9DC" box="[1214,1274,1520,1545]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="130" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<date id="476A0ED0FFF63E282530C8CD6E8DF9DD" box="[1288,1432,1520,1545]" pageId="3" pageNumber="130" value="2012-10-12">
<collectingDate id="572EF738FFF63E282530C8CD6E8DF9DD" box="[1288,1432,1520,1545]" pageId="3" pageNumber="130" value="2012-10-12">12 Oct 2012</collectingDate>
</date>
,
<collectorName id="9E214DC6FFF63E2820AFCB296A0EF9F9" box="[151,283,1556,1581]" pageId="3" pageNumber="130">N. Pinkaew</collectorName>
, np5396 (genitalia slide NP2016).
<collectorName id="9E214DC6FFF63E282298CB296807F9F9" box="[672,786,1556,1581]" pageId="3" pageNumber="130">Deposited</collectorName>
in
<collectionCode id="55C5B0D5FFF63E28230ECB29689BF9FF" box="[822,910,1556,1579]" pageId="3" pageNumber="130">BMNH</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="336B2810FFF63E2820FFCB056E74F9A7" blockId="3.[151,1436,1520,1725]" pageId="3" pageNumber="130">
<materialsCitation id="83BC224DFFF63E2820FFCB056E48F9A7" ID-GBIF-Occurrence="3323415302" collectingDate="2012-10-12" collectionCode="R, KKIC, THNM" collectorName="N. Pinkaew &amp; Deposited" country="Thailand" elevation="30" latitude="12.395278" location="Agroforestry" longLatPrecision="21" longitude="102.67555" pageId="3" pageNumber="130" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Trat" typeStatus="paratype">
<emphasis id="01A0F402FFF63E2820FFCB056A50F984" box="[199,325,1592,1616]" inLineHeading="true" pageId="3" pageNumber="130" reason="1">
<emphasis id="01A0F402FFF63E2820FFCB056A2AF984" bold="true" box="[199,319,1592,1616]" pageId="3" pageNumber="130">
<typeStatus id="EC6F96B2FFF63E2820FFCB056A2AF984" box="[199,319,1592,1616]" pageId="3" pageNumber="130" type="paratype">Paratypes</typeStatus>
</emphasis>
.
</emphasis>
<collectingCountry id="4BC36880FFF63E282175CB056AA0F985" box="[333,437,1592,1617]" name="Thailand" pageId="3" pageNumber="130">Thailand</collectingCountry>
:
<collectingRegion id="F110E6F2FFF63E282187CB046926F984" box="[447,563,1593,1616]" country="Thailand" name="Trat" pageId="3" pageNumber="130">Trat Prov.</collectingRegion>
:
<collectingRegion id="F110E6F2FFF63E282206CB04697EF984" box="[574,619,1593,1616]" country="Thailand" name="Trat" pageId="3" pageNumber="130">Trat</collectingRegion>
<location id="360B7ECBFFF63E28224BCB056811F985" LSID="urn:lsid:plazi:treatment:BB7D9906FFF43E2820AFCF02690CF969:360B7ECBFFF63E28224BCB056811F985" box="[627,772,1592,1617]" country="Thailand" latitude="12.395278" longLatPrecision="21" longitude="102.67555" name="Agroforestry" pageId="3" pageNumber="130" stateProvince="Trat">Agroforestry</location>
<collectionCode id="55C5B0D5FFF63E282333CB056834F985" box="[779,801,1592,1617]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="3" pageNumber="130">R</collectionCode>
.St.,
<geoCoordinate id="56E04ED7FFF63E282368CB0568C0F985" box="[848,981,1592,1617]" degrees="12" direction="north" minutes="23" orientation="latitude" pageId="3" pageNumber="130" precision="15" seconds="43" value="12.395278">12°23'43&quot;N</geoCoordinate>
<geoCoordinate id="56E04ED7FFF63E2823E4CB056F78F985" box="[988,1133,1592,1617]" degrees="102" direction="east" minutes="40" orientation="longitude" pageId="3" pageNumber="130" precision="15" seconds="32" value="102.67555">102°40'32&quot;E</geoCoordinate>
, ca.
<quantity id="F42C85F5FFF63E282498CB056FCEF984" box="[1184,1243,1592,1617]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="130" unit="m" value="30.0">
<elevation id="B8F9CF23FFF63E282498CB056FCEF984" box="[1184,1243,1592,1617]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="130" unit="m" value="30.0">30 m</elevation>
</quantity>
,
<date id="476A0ED0FFF63E2824DFCB056E60F985" box="[1255,1397,1592,1617]" pageId="3" pageNumber="130" value="2012-10-12">
<collectingDate id="572EF738FFF63E2824DFCB056E60F985" box="[1255,1397,1592,1617]" pageId="3" pageNumber="130" value="2012-10-12">12 Oct 2012</collectingDate>
</date>
,
<collectorName id="9E214DC6FFF63E2825B9CB046BEDF9A1" pageId="3" pageNumber="130">N. Pinkaew</collectorName>
(
<specimenCount id="25D2E399FFF63E282130CB616A39F9A1" box="[264,300,1628,1653]" pageId="3" pageNumber="130" type="male">1♂</specimenCount>
, genitalia slide NP2014,
<specimenCount id="25D2E399FFF63E28227ECB61697FF9A1" box="[582,618,1628,1653]" pageId="3" pageNumber="130" type="female">2♀</specimenCount>
, genitalia slide NP2012, NP2017).
<collectorName id="9E214DC6FFF63E2823CFCB616F7CF9A1" box="[1015,1129,1628,1653]" pageId="3" pageNumber="130">Deposited</collectorName>
in
<collectionCode id="55C5B0D5FFF63E2824B4CB616FC5F9A1" box="[1164,1232,1628,1653]" pageId="3" pageNumber="130">KKIC</collectionCode>
and
<collectionCode id="55C5B0D5FFF63E28253FCB616E48F9A7" box="[1287,1373,1628,1651]" pageId="3" pageNumber="130">THNM</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="7BCE7B9BFFF63E2820FFCBBD6FCEF94D" box="[199,1243,1664,1689]" pageId="3" pageNumber="130" type="etymology">
<paragraph id="336B2810FFF63E2820FFCBBD6FCEF94D" blockId="3.[151,1436,1520,1725]" box="[199,1243,1664,1689]" pageId="3" pageNumber="130">
<emphasis id="01A0F402FFF63E2820FFCBBD6A58F94C" box="[199,333,1664,1688]" inLineHeading="true" pageId="3" pageNumber="130" reason="1">
<emphasis id="01A0F402FFF63E2820FFCBBD6A53F94C" bold="true" box="[199,326,1664,1688]" pageId="3" pageNumber="130">Etymology</emphasis>
.
</emphasis>
The specific epithet
<taxonomicName id="F4D45393FFF63E282200CBBD6965F94C" authorityName="Muadsub &amp; Pinkaew" authorityYear="2014" box="[568,624,1664,1688]" class="Insecta" family="Tortricidae" genus="Helictophanes" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="130" phylum="Arthropoda" rank="species" species="flava">
<emphasis id="01A0F402FFF63E282200CBBD6965F94C" box="[568,624,1664,1688]" italics="true" pageId="3" pageNumber="130">flava</emphasis>
</taxonomicName>
(latin = yellow) refers to the ground color of forewing.
</paragraph>
</subSubSection>
<subSubSection id="7BCE7B9BFFF63E2820FFCB9E690CF969" box="[199,537,1699,1725]" pageId="3" pageNumber="130" type="distribution">
<paragraph id="336B2810FFF63E2820FFCB9E690CF969" blockId="3.[151,1436,1520,1725]" box="[199,537,1699,1725]" pageId="3" pageNumber="130">
<emphasis id="01A0F402FFF63E2820FFCB9E6A75F96F" box="[199,352,1699,1724]" inLineHeading="true" pageId="3" pageNumber="130" reason="1">
<emphasis id="01A0F402FFF63E2820FFCB9E6A4CF968" bold="true" box="[199,345,1699,1724]" pageId="3" pageNumber="130">Distribution</emphasis>
.
</emphasis>
<collectingCountry id="4BC36880FFF63E28215FCB996ADEF969" box="[359,459,1700,1725]" name="Thailand" pageId="3" pageNumber="130">Thailand</collectingCountry>
(
<collectingRegion id="F110E6F2FFF63E2821E2CB99691BF969" box="[474,526,1700,1725]" country="Thailand" name="Trat" pageId="3" pageNumber="130">Trat</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>