217 lines
13 KiB
XML
217 lines
13 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.601.9040" ID-GBIF-Dataset="b393a68f-54c2-4334-b33e-f57e5a6ce8f0" ID-PMC="PMC4978076" ID-Pensoft-Pub="1313-2970-601-1" ID-PubMed="27551184" ID-ZBK="BAF2B0C0FB8540E8B78AB847EF195BE3" ModsDocAuthor="" ModsDocDate="2016" ModsDocID="1313-2970-601-1" ModsDocOrigin="ZooKeys 601" ModsDocTitle="Three new species of western California springsnails previously confused with Pyrgulopsisstearnsiana (Caenogastropoda, Hydrobiidae)" checkinTime="1467213884928" checkinUser="pensoft" docAuthor="Hershler, Robert, Liu, Hsiu-Ping, Babbitt, Caitlin, Kellogg, Michael G. & Howard, Jeanette K." docDate="2016" docId="86445AD80237F0FDCF8097C7687C2B4B" docLanguage="en" docName="ZooKeys 601: 1-19" docOrigin="ZooKeys 601" docSource="http://dx.doi.org/10.3897/zookeys.601.9040" docTitle="Pyrgulopsis torrida Hershler, Liu, Babbitt, Kellogg & Howard, sp. n." docType="treatment" docUuid="2FBB4B8B-32C2-4308-AB78-C8454A1B8ED1" docUuidSource="ZooBank" docVersion="5" lastPageNumber="11" masterDocId="FFAA08334E20A2343D62F64E9074FFFB" masterDocTitle="Three new species of western California springsnails previously confused with Pyrgulopsisstearnsiana (Caenogastropoda, Hydrobiidae)" masterLastPageNumber="19" masterPageNumber="1" pageNumber="9" updateTime="1668163257691" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Three new species of western California springsnails previously confused with Pyrgulopsisstearnsiana (Caenogastropoda, Hydrobiidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hershler, Robert</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Liu, Hsiu-Ping</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Babbitt, Caitlin</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Kellogg, Michael G.</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Howard, Jeanette K.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2016</mods:date>
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<mods:detail type="volume">
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<mods:number>601</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>19</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.601.9040</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.601.9040</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-601-1</mods:identifier>
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<mods:identifier type="ZBK">BAF2B0C0FB8540E8B78AB847EF195BE3</mods:identifier>
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<mods:identifier type="ZooBank">BAF2B0C0FB8540E8B78AB847EF195BE3</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="127901437" LSID="urn:lsid:zoobank.org:act:2FBB4B8B-32C2-4308-AB78-C8454A1B8ED1" httpUri="http://treatment.plazi.org/id/86445AD80237F0FDCF8097C7687C2B4B" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9">
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<subSubSection pageId="8" pageNumber="9" type="multiple">
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<paragraph pageId="8" pageNumber="9">Taxon classification Animalia Littorinimorpha Hydrobiidae</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="nomenclature">
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName LSID="http://zoobank.org/2FBB4B8B-32C2-4308-AB78-C8454A1B8ED1" authority="Hershler, Liu, Babbitt, Kellogg & Howard" class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgulopsis torrida" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="torrida">Pyrgulopsis torrida Hershler, Liu, Babbitt, Kellogg & Howard</taxonomicName>
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<taxonomicNameLabel pageId="8" pageNumber="9">sp. n.</taxonomicNameLabel>
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Figs 3C, 6
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="reference_group">
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<paragraph pageId="8" pageNumber="9">
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<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgulopsis stearnsiana" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="stearnsiana">Pyrgulopsis stearnsiana</taxonomicName>
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.-
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<bibRefCitation author="Hershler, R" journalOrPublisher="Zootaxa" pageId="13" pageNumber="14" pagination="1 - 17" title="Two new, possibly threatened species of Pyrgulopsis (Gastropoda: Hydrobiidae) from southwestern California." volume="2343" year="2010">Hershler and Liu 2010</bibRefCitation>
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(in part).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="types">
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<paragraph pageId="8" pageNumber="9">Types.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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Holotype, SBMNH 74238, Little Sycamore Canyon, creek 3.2 km up flow from Hwy 1, Ventura County, California,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="34.07509">34.07509°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-118.95508">118.95508°W</geoCoordinate>
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, 11/11/1961, W. B. Miller. Paratypes, SBMNH 460492 (ca. 200 dried shells), from same lot.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="materials_examined">
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<paragraph pageId="8" pageNumber="9">Referred material.</paragraph>
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<paragraph pageId="8" pageNumber="9">California. Ventura County: SBMNH 74236, * USNM 1120443, ibid, 9/9/1956, 10/21/2008, respectively.</paragraph>
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</subSubSection>
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<subSubSection pageId="8" pageNumber="9" type="diagnosis">
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<paragraph pageId="8" pageNumber="9">Diagnosis.</paragraph>
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<paragraph pageId="8" pageNumber="9">
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A medium-sized congener (maximum shell height, 2.8 mm) having an ovate-conic shell. Distinguished from
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<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgulopsis stearnsiana" order="Littorinimorpha" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="stearnsiana">Pyrgulopsis stearnsiana</taxonomicName>
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by its shorter penial filament and larger terminal gland.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="description">
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<paragraph pageId="8" pageNumber="9">Description.</paragraph>
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<paragraph lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
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Shell (Fig. 6
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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, Table 5) ovate-conic, whorls 4.00. Teleoconch whorls medium convex, narrowly shouldered. Aperture ovate, slightly angled above; parietal lip complete, nearly straight, narrowly adnate adapically or slightly disjunct,
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<pageBreakToken pageId="9" pageNumber="10" start="start">thin</pageBreakToken>
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or slightly thickened; umbilicus small. Outer lip thin, orthocline. Teleoconch smooth or sculptured with weak spiral striae.
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</paragraph>
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<caption pageId="9" pageNumber="10">
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<paragraph pageId="9" pageNumber="10">
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Figure 6. Shells, opercula and radula,
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<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgulopsis torrida" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="torrida">Pyrgulopsis torrida</taxonomicName>
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sp. n. A Holotype, SBMNH 74238 B Shell, SBMNH 460492 C, D Opercula (outer, inner sides), SBMNH 460492 E Portion of radular ribbon, SBMNH 460492 F Central teeth, SBMNH 460492 G Lateral teeth, SBMNH 460492. Scale bars:
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<normalizedToken originalValue="A–B">A-B</normalizedToken>
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=1.0 mm;
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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=100
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<normalizedToken originalValue="µm">µm</normalizedToken>
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;
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<normalizedToken originalValue="E–G">E-G</normalizedToken>
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=10
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<normalizedToken originalValue="µm">µm</normalizedToken>
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.
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</paragraph>
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</caption>
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<caption pageId="9" pageNumber="10">
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<paragraph pageId="9" pageNumber="10">
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Table 5. Shell parameters for
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<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgulopsis torrida" order="Littorinimorpha" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="torrida">Pyrgulopsis torrida</taxonomicName>
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. Measurements are in mm.
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</paragraph>
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</caption>
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<paragraph pageId="9" pageNumber="10">
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<table pageId="9" pageNumber="10">
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<tr pageId="9" pageNumber="10">
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">WH</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">SH</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">SW</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">HBW</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">WBW</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">AH</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">AW</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">SW</abbrev>
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/
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<abbrev pageId="9" pageNumber="10">SH</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">HBW</abbrev>
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/
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<abbrev pageId="9" pageNumber="10">SH</abbrev>
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</th>
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<th colspan="1" pageId="9" pageNumber="10" rowspan="1">
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<abbrev pageId="9" pageNumber="10">AH</abbrev>
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/
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<abbrev pageId="9" pageNumber="10">SH</abbrev>
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</th>
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</tr>
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<tr pageId="9" pageNumber="10">
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<td colspan="11" pageId="9" pageNumber="10" rowspan="1">SBMNH</td>
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</tr>
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<tr pageId="9" pageNumber="10">
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<td colspan="11" pageId="9" pageNumber="10" rowspan="1">SBMNH</td>
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</tr>
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</table>
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</paragraph>
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<paragraph pageId="9" pageNumber="10">
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Operculum (Fig. 6
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<normalizedToken originalValue="C–D">C-D</normalizedToken>
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) as for genus; portion of attachment scar margin slightly thickened on inner side. Radula (Fig. 6
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<normalizedToken originalValue="E–G">E-G</normalizedToken>
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) as for genus; dorsal edge of central teeth concave, lateral cusps
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<normalizedToken originalValue="three–six">three-six</normalizedToken>
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, basal cusps one to (rarely) two. Lateral teeth having
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<normalizedToken originalValue="two–four">two-four</normalizedToken>
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cusps on inner and
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<normalizedToken originalValue="three–five">three-five</normalizedToken>
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cusps on outer side. Inner marginal teeth with 19-24 cusps, outer marginal teeth with 21-27 cusps. Radula data are from SBMNH 460492.
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</paragraph>
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<paragraph pageId="10" pageNumber="11">
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<pageBreakToken pageId="10" pageNumber="11" start="start">Penis</pageBreakToken>
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(Fig. 3C) small, filament weakly pigmented or pale, filament short, narrow, horizontal, weakly tapering; lobe small, rectangular, horizontal; terminal gland fairly large, narrow, overlapping dorsal and ventral edges of lobe. Penial data are from USNM 1120443 (2 specimens).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="etymology">
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<paragraph pageId="10" pageNumber="11">Etymology.</paragraph>
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<paragraph pageId="10" pageNumber="11">The species name is an adjective derived from the New Latin torridus, meaning dry or parched, and refers to the recent desiccation of the stream in Little Sycamore Canyon. We propose "Little Sycamore pyrg" as the common name for this species.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="distribution">
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<paragraph pageId="10" pageNumber="11">Distribution.</paragraph>
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<paragraph pageId="10" pageNumber="11">Endemic to the type locality, a small, shallow stream which runs for about 1.6 km. Snails were collected from the mud bottoms of a series of small puddle-like pools along the middle section of the stream.</paragraph>
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</subSubSection>
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<subSubSection pageId="10" pageNumber="11" type="conservation status">
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<paragraph pageId="10" pageNumber="11">Conservation status.</paragraph>
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<paragraph pageId="10" pageNumber="11">
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<taxonomicName class="Gastropoda" family="Hydrobiidae" genus="Pyrgulopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pyrgulopsis torrida" order="Littorinimorpha" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="torrida">Pyrgulopsis torrida</taxonomicName>
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was found only rarely in the Little Sycamore Canyon creek in 2000. The entirely length of the canyon was dry when re-visited in 2015, suggesting that this population may now be extirpated.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |