303 lines
35 KiB
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303 lines
35 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1195.112112" ID-Pensoft-Pub="1313-2970-1195-157" ID-Pensoft-UUID="E26F0ADE4B4C5BCB943E2FCC63E349C4" ID-ZooBank="7CFF082DDA924ECF8E7C0A9972F6CD08" ModsDocID="1313-2970-1195-157" checkinTime="1710460428610" checkinUser="pensoft" docAuthor="Tongkerd, Piyoros, Lwin, Ngwe, Pall-Gergely, Barna, Chanabun, Ratmanee, Pholyotha, Arthit, Prasankok, Pongpun, Seesamut, Teerapong, Siriwut, Warut, Srisonchai, Ruttapon, Sutcharit, Chirasak & Panha, Somsak" docDate="2024" docId="2CDC58A571DF586896B0FF41EA56A00A" docLanguage="en" docName="ZooKeys 1195: 157-197" docOrigin="ZooKeys 1195" docPubDate="2024-03-14" docSource="http://dx.doi.org/10.3897/zookeys.1195.112112" docTitle="Bensonella taiyaiorum Tongkerd & Panha 2024, sp. nov." docType="treatment" docUuid="E83B1A5F-A724-4BBF-9A7D-BB1604ECEFEF" docUuidSource="ZooBank" docVersion="1" id="E26F0ADE4B4C5BCB943E2FCC63E349C4" lastPageNumber="157" masterDocId="E26F0ADE4B4C5BCB943E2FCC63E349C4" masterDocTitle="Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species" masterLastPageNumber="197" masterPageNumber="157" pageNumber="157" updateTime="1710460428610" updateUser="pensoft">
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<mods:title id="707BF6E3A10486F79868ADE9F0144DFA">Contributions of a small collection of terrestrial microsnails (Pupilloidea, Hypselostomatidae) from Myanmar with description of three new species</mods:title>
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<mods:namePart id="F1F6C0C84E16249640AF70545C1DFF97">Tongkerd, Piyoros</mods:namePart>
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<mods:affiliation id="9860BF063087F01C12D6A67D5B023C18">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="74A77568F35809A304B31C394C79742A">Lwin, Ngwe</mods:namePart>
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<mods:affiliation id="161DDBF321A4621E00E27BE4C427B2DD">Fauna and Flora International, No. 35, 3 rd Floor, Shan Gone Condo, Myay Ni Gone Market Street, Sanchaung Township, Yangon, Myanmar</mods:affiliation>
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<mods:namePart id="3707F6DDEB1256FBCA768E250CE9671B">Pall-Gergely, Barna</mods:namePart>
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<mods:affiliation id="DB5D33123B80C3DFACE2B7D0A657F356">Plant Protection Institute, Centre for Agricultural Research, ELKH, Herman Otto ut 15, 1022 Budapest, Hungary</mods:affiliation>
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<mods:affiliation id="033D3CF5CE66235C2314127E92B77994">Program in Animal Science, Faculty of Agricultural Technology, Sakon Nakhon Rajabhat University, Sakon Nakhon 47000, Thailand</mods:affiliation>
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<mods:namePart id="BA944F4DC6EAAF4598E7E1EB279E9C73">Pholyotha, Arthit</mods:namePart>
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<mods:affiliation id="D9E82F9BFC5CB2F07765B747A6E7DA57">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:namePart id="55509E93DF048F73F5E5357033F0BA76">Prasankok, Pongpun</mods:namePart>
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<mods:affiliation id="2D9B404603AF3A4306D9060BA1791BA3">Biodiversity and Utilization Research Unit, Center of Excellence in Modern Agriculture, Sakon Nakhon Rajabhat University, Sakon Nakhon 47000, Thailand</mods:affiliation>
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<mods:namePart id="DEA9B2CC135E11832AA61AC308C6843D">Seesamut, Teerapong</mods:namePart>
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<mods:affiliation id="0C0CE3EAE68BB9EE07A528689FAC374B">School of Biology, Institute of Science, Suranaree University of Technology, Nakhon Ratchasima 30000, Thailand</mods:affiliation>
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<mods:namePart id="BA9787A0818E57F363CEDF5E519A1A95">Siriwut, Warut</mods:namePart>
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<mods:affiliation id="4F2D352A0A9340D5FADA11CCCF63B675">Department of Biology, Faculty of Science, Rangsit University, Pathum Thani 12000, Thailand</mods:affiliation>
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<mods:namePart id="7ED812D6817B872BBD96C34380ED1797">Srisonchai, Ruttapon</mods:namePart>
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<mods:affiliation id="9C63E6EBAB7C3237D2C3C154A068950D">Animal Systematics and Molecular Ecology Laboratory, Department of Biology, Faculty of Science, Mahidol University, Bangkok 10400, Thailand</mods:affiliation>
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<mods:namePart id="0AB53FA01E52CF06744317181558F513">Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation id="B90F6A823BB47B6CE0E2DC883FF1DC2C">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier id="89BAB54132ACCF0E5A2138E1C5AF3404" type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="391CF5167DDE6AA2843283A3F593866B">Panha, Somsak</mods:namePart>
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<mods:affiliation id="D1793B1FCCB7774DC45195BB26B90072">Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Department of Biology, Faculty of Science, Khon Kaen University, Khon Kaen 40002, Thailand</mods:affiliation>
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<mods:nameIdentifier id="7F76DBE6ACDA866AC3C780C414EA4E8E" type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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<mods:title id="BF7B2A94E32E85ADA338322FD7706168">ZooKeys</mods:title>
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<mods:date id="EEE95A676B6772C1D13F79FD71D11DB7">2024</mods:date>
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<mods:number id="3E1DB20697C794285FDF5189B326E7CE">2024-03-14</mods:number>
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<mods:number id="F496F74F36B77F935B58CFC1E045CEA2">1195</mods:number>
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<mods:identifier id="0B2CC9575371C5F90975EB0AC9B4FCE0" type="DOI">http://dx.doi.org/10.3897/zookeys.1195.112112</mods:identifier>
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<treatment id="2CDC58A571DF586896B0FF41EA56A00A" LSID="urn:lsid:zoobank.org:act:E83B1A5F-A724-4BBF-9A7D-BB1604ECEFEF" httpUri="http://treatment.plazi.org/id/2CDC58A571DF586896B0FF41EA56A00A" lastPageNumber="157" pageId="0" pageNumber="157">
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<subSubSection id="4D8F120F9A0290EA4A155C1891398594" pageId="0" pageNumber="157" type="nomenclature">
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<paragraph id="0995C048E3A45028B6A32A7F2341053D" pageId="0" pageNumber="157">
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<taxonomicName id="DA610C2BD6336DE3B6DB33A85C9C6CCE" LSID="https://zoobank.org/E83B1A5F-A724-4BBF-9A7D-BB1604ECEFEF" authority="Tongkerd & Panha" authorityName="Tongkerd & Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella taiyaiorum" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="taiyaiorum" status="sp. nov.">Bensonella taiyaiorum Tongkerd & Panha</taxonomicName>
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<taxonomicNameLabel id="E9D5DC6939C27EED74D6B12EBCA60521" pageId="0" pageNumber="157">sp. nov.</taxonomicNameLabel>
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</paragraph>
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<subSubSection id="9C8444A13061F4FE0CD8FF4FA253CDF2" pageId="0" pageNumber="157" type="description">
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<paragraph id="6470D4A53C983C10EF07B2AB8FC87550" pageId="0" pageNumber="157">
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<figureCitation id="0D5584AD44A7189CDCAA6458D194D6EC" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Bensonella anguloobtusa, specimen CUMZ 14398 from Taunggyi District, Shan State B Bensonella taiyaiorum sp. nov., holotype CUMZ 14380 from Taunggyi District, Shan State C Bensonella lophiodera sp. nov., holotype CUMZ 14378 from Taunggyi District, Shan State. Abbreviation: ip: infraparietal lamella." figureDoi="10.3897/zookeys.1195.112112.figure7" httpUri="https://binary.pensoft.net/fig/1002056" pageId="0" pageNumber="157">Figs 7B</figureCitation>
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<figureCitation id="24D55FA09168DFE27B8C9476E153B81F" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bensonella taiyaiorum sp. nov., from Taunggyi District, Shan State A-C paratypes CUMZ 14403. The white arrow indicates constriction. The yellow arrow indicates a cervical crest. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, sbc: subcolumellar lamella, spc: supracolumellar lamella, sppl: suprapalatal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure8" httpUri="https://binary.pensoft.net/fig/1002057" pageId="0" pageNumber="157">, 8</figureCitation>
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<figureCitation id="D09B54D7C3E43459F0DC58209419A27A" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">, 13H</figureCitation>
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</paragraph>
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||
</subSubSection>
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<subSubSection id="F9408A84442B343A27990A6DF0AE3039" pageId="0" pageNumber="157" type="type locality">
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<paragraph id="E7C5EC39032EE79120566E889B785BC7" pageId="0" pageNumber="157">Type locality.</paragraph>
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||
<paragraph id="83B134C9A86FCDE0469076C49F1E69C3" pageId="0" pageNumber="157">
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||
Dragon Rock, Pindaya Township, Taunggyi District, Shan State, Myanmar (locality code SH6;
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<geoCoordinate id="CFBC272DD959FF7BF206F951936CD4D8" degrees="20" direction="north" minutes="55" orientation="latitude" precision="1" seconds="31.5" value="20.925417">20°55'31.5"N</geoCoordinate>
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,
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<geoCoordinate id="0E2D24412F38C2605771E567544E5019" degrees="96" direction="east" minutes="39" orientation="longitude" precision="1" seconds="01.2" value="96.65034">96°39'01.2"E</geoCoordinate>
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||
; 1300 m a.s.l.).
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||
</paragraph>
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||
</subSubSection>
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<subSubSection id="AA7E3985A822B59876C887E9771D8B40" pageId="0" pageNumber="157" type="type material">
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<paragraph id="736A105E05AD46D2AF15E295AA06B9AC" pageId="0" pageNumber="157">Type material.</paragraph>
|
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<paragraph id="8A6BF3284843FF29610444C471318245" pageId="0" pageNumber="157">
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||
<emphasis id="A1CD99C6D6657A007738009439AF22D7" bold="true" italics="true" pageId="0" pageNumber="157">Holotype</emphasis>
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CUMZ 14380 (height 2.8 mm, width 2.1 mm; Fig.
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<figureCitation id="09DE9704D82E341F6413C16AD26D1894" captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. A Bensonella anguloobtusa, specimen CUMZ 14398 from Taunggyi District, Shan State B Bensonella taiyaiorum sp. nov., holotype CUMZ 14380 from Taunggyi District, Shan State C Bensonella lophiodera sp. nov., holotype CUMZ 14378 from Taunggyi District, Shan State. Abbreviation: ip: infraparietal lamella." figureDoi="10.3897/zookeys.1195.112112.figure7" httpUri="https://binary.pensoft.net/fig/1002056" pageId="0" pageNumber="157">7B</figureCitation>
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).
|
||
<emphasis id="7D4C52D2E9DA2012E58EAF93C7C1D07C" bold="true" italics="true" pageId="0" pageNumber="157">Paratypes</emphasis>
|
||
CUMZ 14381 (41 shells); CUMZ 14402 (12 shells; measured); CUMZ 14403 (3 shells; Figs
|
||
<figureCitation id="E8C4E14CE7484406EA547AA1A037F9CA" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Bensonella taiyaiorum sp. nov., from Taunggyi District, Shan State A-C paratypes CUMZ 14403. The white arrow indicates constriction. The yellow arrow indicates a cervical crest. Abbreviations: a: angular lamella, b: basal plica, c: columellar lamella, ip: infraparietal lamella, itpl: interpalatal plica, lpl: lower palatal plica, p: parietal lamella, pt: palatal tubercle, sbc: subcolumellar lamella, spc: supracolumellar lamella, sppl: suprapalatal lamella, upl: upper palatal plica." figureDoi="10.3897/zookeys.1195.112112.figure8" httpUri="https://binary.pensoft.net/fig/1002057" pageId="0" pageNumber="157">8</figureCitation>
|
||
,
|
||
<figureCitation id="27BFE8B6F2266AA87384865231EF149D" captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Synoptic view of representatives of hypselostomatid snails recorded from Myanmar A Acinolaemus dayanum, specimen HA Collection B Acinolaemus cryptidentatus, specimen CUMZ 14413 C Anauchen eotvosi, holotype NHMUK 1903.7.1.1227.1 (after Pall-Gergely 2023 a: fig. 1 c) D Angustopila occidentalis, holotype HNHM 103483 (after Pall-Gergely et al. 2023: fig. 70 a) E Angustopila elevata, specimen HNHM 103484 (after Pall-Gergely et al. 2023: fig. 13 a) F Bensonella salwiniana, specimen CUMZ 14393 G Bensonella anguloobtusa, specimen CUMZ 14400 H Bensonella taiyaiorum sp. nov., paratype CUMZ 14403 I Bensonella lophiodera sp. nov., paratype CUMZ 14405 J Clostophis sankeyi, holotype UMZC I. 103320 (after Preece et al. 2022: fig. 70 e) K Clostophis thinbowguensis, holotype MNHN-IM- 2000 - 38057 (after Pall-Gergely et al. 2022: fig. 9 c) L Gyliotrachela bensonianum, syntype NHMUK 20191141 (after Gojsina et al. 2022: fig. 4 a) M Gyliotrachela hungerfordiana, specimen NHMUK collection from Perak, Malaysia N Gyliotrachela tianxingqiaoensis, holotype IZCAS TM 025075 (after Gojsina et al. 2022: fig. 3 a) O Gyliotrachela muangon, specimen CUMZ 14410 P Gyliotrachela aunglini sp. nov., paratype CUMZ 14412 Q Hypselostoma tubiferum, specimen NHMUK 1888.12.4.17 - 22 ex. Theobald collection from Tonduong, Burma." figureDoi="10.3897/zookeys.1195.112112.figure13" httpUri="https://binary.pensoft.net/fig/1002062" pageId="0" pageNumber="157">13H</figureCitation>
|
||
), NHMUK 20230591 (3 shells), SMF 373019 (3 shells) all from the type locality.
|
||
</paragraph>
|
||
<caption id="F268262D7CE1D20E8D55674127338035" doi="10.3897/zookeys.1195.112112.figure8" httpUri="https://binary.pensoft.net/fig/1002057" pageId="0" pageNumber="157" start="Figure 8" startId="F8">
|
||
<paragraph id="82BE957F861814EB4A61CCF5A5435A7B" pageId="0" pageNumber="157">
|
||
<emphasis id="0E068F26A24BE332066672D0FBFD31F4" bold="true" pageId="0" pageNumber="157">Figure 8.</emphasis>
|
||
<taxonomicName id="B8E4151775FC27E7F678BD0317344792" authorityName="Tongkerd & Panha" authorityYear="2024" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella taiyaiorum" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="species" species="taiyaiorum">
|
||
<emphasis id="110606F6A3657EAABA9FD06BBF72B1C9" italics="true" pageId="0" pageNumber="157">Bensonella taiyaiorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., from Taunggyi District, Shan State
|
||
<emphasis id="AA61F38BBDFA72169493AD1569FF58C5" bold="true" pageId="0" pageNumber="157">A-C</emphasis>
|
||
paratypes CUMZ 14403. The white arrow indicates constriction. The yellow arrow indicates a cervical crest. Abbreviations:
|
||
<emphasis id="20388349AFCACFF7FBCF10E0DBE7F354" bold="true" pageId="0" pageNumber="157">a</emphasis>
|
||
: angular lamella,
|
||
<emphasis id="2640940CAB8CB45CD3CA3C6A675B68F7" bold="true" pageId="0" pageNumber="157">b</emphasis>
|
||
: basal plica,
|
||
<emphasis id="8C33F3C580B6F296F950D59FE91F1119" bold="true" pageId="0" pageNumber="157">c</emphasis>
|
||
: columellar lamella,
|
||
<emphasis id="078E28DA7D23422EB485036944565CA4" bold="true" pageId="0" pageNumber="157">ip</emphasis>
|
||
: infraparietal lamella,
|
||
<emphasis id="DB1D5A19E13390B5769C7269B54AA994" bold="true" pageId="0" pageNumber="157">itpl</emphasis>
|
||
: interpalatal plica,
|
||
<emphasis id="C74AD2420B990B30E19B7DA5134187DC" bold="true" pageId="0" pageNumber="157">lpl</emphasis>
|
||
: lower palatal plica,
|
||
<emphasis id="F511B4F8D34C8636CFA55C25344C26FC" bold="true" pageId="0" pageNumber="157">p</emphasis>
|
||
: parietal lamella,
|
||
<emphasis id="2EB5A3312930EE2D8489A0A73D7289AA" bold="true" pageId="0" pageNumber="157">pt</emphasis>
|
||
: palatal tubercle,
|
||
<emphasis id="4283FD75E680733F118D0CE8E8B48B6A" bold="true" pageId="0" pageNumber="157">sbc</emphasis>
|
||
: subcolumellar lamella,
|
||
<emphasis id="DD532A70A00436EC38AEB623B1BABA0A" bold="true" pageId="0" pageNumber="157">spc</emphasis>
|
||
: supracolumellar lamella,
|
||
<emphasis id="8388C8C771905B5FC9D0BCCC692E8342" bold="true" pageId="0" pageNumber="157">sppl</emphasis>
|
||
: suprapalatal lamella,
|
||
<emphasis id="90C0FB1EF87476E3899C65AAE6163D03" bold="true" pageId="0" pageNumber="157">upl</emphasis>
|
||
: upper palatal plica.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
<subSubSection id="4F7A0DC31EBF981A2EFD36C66BAD1D36" pageId="0" pageNumber="157" type="diagnosis">
|
||
<paragraph id="4B5A8D72EC8439D8E980D6E6E1904BE0" pageId="0" pageNumber="157">Diagnosis.</paragraph>
|
||
<paragraph id="F9E3871F4F8EBFD0300AD2802B43D8FD" pageId="0" pageNumber="157">Shell elongate-conical and yellowish brown in colour. Cervical crest situated far from peristome. Aperture with several hookless barriers (i.e., parietal, infraparietal, angular, columellar, supra- and sub-columellae, basal, and upper, inter and lower palatals), and interpalatal and lower palatal plicae forming u-shaped plica.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B6B910BEAD29575B4C9C1558C3568590" pageId="0" pageNumber="157" type="etymology">
|
||
<paragraph id="D9CC6B781CE662163732DC3D3DB8FD9F" pageId="0" pageNumber="157">Etymology.</paragraph>
|
||
<paragraph id="5F0707BB5E92E22AD6D21E967C03572D" pageId="0" pageNumber="157">
|
||
The specific name
|
||
<emphasis id="3054741FF8B5BA63AD72F3B62DC7686B" italics="true" pageId="0" pageNumber="157">taiyaiorum</emphasis>
|
||
is in recognition of the 'Tai
|
||
<normalizedToken id="B9D685D6A5D8DFCF67EBE842685467FC" originalValue="Yai’">Yai'</normalizedToken>
|
||
or
|
||
<normalizedToken id="9E0BDCB044B76C66A16A6A7D962E00B6" originalValue="‘Shan’">'Shan'</normalizedToken>
|
||
, the largest ethnic group in Shan State, which is the type locality of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A41FC811705EF67A48474416AE20CDA0" pageId="0" pageNumber="157" type="description">
|
||
<paragraph id="2B9E6BA63605BA308B932B086F1E4595" pageId="0" pageNumber="157">Description.</paragraph>
|
||
<paragraph id="7D89362AB6CDE821161BC972EAD36173" pageId="0" pageNumber="157">
|
||
Shell conical, yellowish brown to reddish brown and with
|
||
<normalizedToken id="47A424107DB63E99A4603F6D667D1320" originalValue="5¼">51/4</normalizedToken>
|
||
-6 widely convex whorls. Shell height 2.5-2.9 mm and shell width 2.0-2.2 mm. Apex blunt; protoconch ~
|
||
<normalizedToken id="58134D36C7A105BC5F4E731A524D86ED" originalValue="1½">11/2</normalizedToken>
|
||
whorls with wrinkles and very weak spiral ridges. Teleoconch with strong and irregular growth lines, sometimes white growth lines are conspicuous; suture impressed and deep. Last whorl large and rounded. Peristome thickened and slightly expanded; constriction weak; lip reddish brown. Cervical crest sharp and situated far from peristome. Aperture subrectangular and with many apertural dentitions. Parietal lamella strongly developed and consisting of two parts separated by an incision: outer part large, strong, with rectangular shape and reaching peristome; inner part blunt. Infraparietal lamella small and blunt. Angular lamella lower than parietal lamella: outer part short and reaching peristome; inner part more strongly elevated. Palatal tubercle strongly developed with triangular shape; two upper palatal plicae with strong folding; inter- and lower palatal large, equal in size and connected to form u-shaped plica. Basal plica small, low and tubercle-like. Columellar lamella large, strong and distinct in its horizontal alignment; one small supracolumellar lamella and one small subcolumellar lamella present. Umbilicus narrowly perforate, ~ 1/5 of shell width, rounded and deep.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C314BD31FC7F0B6F85C9EE9D81026371" pageId="0" pageNumber="157" type="distribution">
|
||
<paragraph id="6E45564869E091A45C30E83ABFC78DB9" pageId="0" pageNumber="157">Distribution.</paragraph>
|
||
<paragraph id="6A01BD906C69EC729EE8FF1A54B71157" pageId="0" pageNumber="157">This new species is known only from the type locality in Shan State. The snails live on limestone walls and can be found under leaf litter within rock crevices.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="456C8BDB6F6BC52D1508A2424B48432A" pageId="0" pageNumber="157" type="differential diagnosis">
|
||
<paragraph id="D577A717F818062B302A06544C8FBB25" pageId="0" pageNumber="157">Differential diagnosis.</paragraph>
|
||
<paragraph id="88A8A01F9523205E44BA0B7B4B8F8855" pageId="0" pageNumber="157">
|
||
The new species can be distinguished from
|
||
<taxonomicName id="A576E8FA9F99C40E03F3F449DE42D800" authorityName="Pilsbry & Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
|
||
<emphasis id="5291813D37CD9DD7B28D904A68B5B270" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
reported from Thailand and Laos in having inter- and lower- palatal plicae connected, forming a u-shaped plica, and with prominent cervical crest. In contrast, the three species from Thailand differ by:
|
||
<taxonomicName id="CE0F0F05AA7B5B2CB8B0E108586E0ABF" lsidName="B. nabhitabhatai" pageId="0" pageNumber="157" rank="species" species="nabhitabhatai">
|
||
<emphasis id="464A2925532DB0EB98131D6565320EA6" italics="true" pageId="0" pageNumber="157">B. nabhitabhatai</emphasis>
|
||
</taxonomicName>
|
||
has one upper palatal plica and no supracolumellar and infraparietal lamellae;
|
||
<taxonomicName id="D7F9F02A738140F1DB8C6E11BF239071" lsidName="B. tamphathai" pageId="0" pageNumber="157" rank="species" species="tamphathai">
|
||
<emphasis id="100E14163980BDF4C83166C9746ECE58" italics="true" pageId="0" pageNumber="157">B. tamphathai</emphasis>
|
||
</taxonomicName>
|
||
possesses fine spiral striae on the teleoconch, one upper palatal plica, and bears no supracolumellar lamellae;
|
||
<taxonomicName id="3B2688653DD03C3E299F9F1B0DCF7B0A" lsidName="B. pangmapaensis" pageId="0" pageNumber="157" rank="species" species="pangmapaensis">
|
||
<emphasis id="D012B42A9E02FD28A9C51E335590A854" italics="true" pageId="0" pageNumber="157">B. pangmapaensis</emphasis>
|
||
</taxonomicName>
|
||
has fine spiral striae on the teleoconch, and hooked palatal and basal plicae (
|
||
<bibRefCitation id="12032F665FEC26DD00399D5686F9107F" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" pagination="107 - 112" refId="B59" refString="Panha, S, Burch, JB, 2000. A new species of Paraboysidia from Thailand (Pulmonata: Pupillidae). Walkerana 11 (25): 107 - 112" title="A new species of Paraboysidia from Thailand (Pulmonata: Pupillidae)." volume="11" year="2000">Panha and Burch 2000</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="A35FF079EC7064A1E6E9788F1FEE9965" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B61" refString="Panha, S, Burch, JB, 2002b. The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12[2001](28): 77-94. [Date of publication follow Jirapatrasilp et al. (2023)]" title="The pupillid genus Paraboysidia in Thailand (Pulmonata: Stylommatophora). Walkerana 12 [2001] (28): 77 - 94. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2002 b">2002b</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="19E5A6111579C25973042B9CD80D4719" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B63" refString="Panha, S, Burch, JB, 2005. An introduction to the microsnails of Thailand. Malacological Review 37/38(2004-2005): 1-155. [Date of publication follow Jirapatrasilp et al. (2023)]" title="An introduction to the microsnails of Thailand. Malacological Review 37 / 38 (2004 - 2005): 1 - 155. [Date of publication follow Jirapatrasilp et al. (2023)]" year="2005">2005</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="B3224CDA2FDD66C82769C3DD38F3641A" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B64" refString="Panha, S, Tongkerd, P, Sutcharit, C, Tumpeesuwan, S, Vongsombath, C, 2003. A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana 13[2002](29/30): 123-128." title="A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana 13 [2002] (29 / 30): 123 - 128." year="2003">Panha et al. 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="5880B1EF6DAE36B28EAE3784E3AD7680" author="Inkhavilay, K" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="157" pagination="213 - 232" refId="B28" refString="Inkhavilay, K, Sutcharit, C, Tongkerd, P, Panha, S, 2016. New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology 42 (4): 213 - 232" title="New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae)." volume="42" year="2016">Inkhavilay et al. 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="83F881C633A1EF0503210E34FD4650B2" pageId="0" pageNumber="157">
|
||
For further comparison, the three species from Laos can be distinguished by:
|
||
<taxonomicName id="EA6024122C77EA20527538902D1A0281" lsidName="B. paralella" pageId="0" pageNumber="157" rank="species" species="paralella">
|
||
<emphasis id="3D8E4F5022D5E0F4984ABBB19A8ECB4B" italics="true" pageId="0" pageNumber="157">B. paralella</emphasis>
|
||
</taxonomicName>
|
||
has one upper palatal plica, and no interpalatal plicae, supra- and sub- columellar lamellae;
|
||
<taxonomicName id="9E43DB1FE2BFC433CE2C504FC65B0EC7" lsidName="B. anguloobtusa" pageId="0" pageNumber="157" rank="species" species="anguloobtusa">
|
||
<emphasis id="BC500F2958CBA2F1089EBD9E6818E8A2" italics="true" pageId="0" pageNumber="157">B. anguloobtusa</emphasis>
|
||
</taxonomicName>
|
||
has one upper palatal plica and no basal and supracolumellar lamellae;
|
||
<taxonomicName id="72B0E416517628F9C6A108B1315C91B3" lsidName="B. wangviangensis" pageId="0" pageNumber="157" rank="species" species="wangviangensis">
|
||
<emphasis id="9A52D429FD0D83FD90704A7EFFACCC84" italics="true" pageId="0" pageNumber="157">B. wangviangensis</emphasis>
|
||
</taxonomicName>
|
||
has a unique shape, smaller shell size (height 1-2 mm), four whorls, shouldered last whorl, and bears no basal plicae and columellar lamellae (
|
||
<bibRefCitation id="56C6147BA1995624C6B32F7598306D9E" author="Panha, S" journalOrPublisher="Walkerana" pageId="0" pageNumber="157" refId="B64" refString="Panha, S, Tongkerd, P, Sutcharit, C, Tumpeesuwan, S, Vongsombath, C, 2003. A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana 13[2002](29/30): 123-128." title="A new species of Paraboysidia (Pupillidae: Gastrocoptinae) from Laos. Walkerana 13 [2002] (29 / 30): 123 - 128." year="2003">Panha et al. 2003</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="9F92840AF50C28BA4CCF67F8E7B581CD" author="Inkhavilay, K" journalOrPublisher="Journal of Conchology" pageId="0" pageNumber="157" pagination="213 - 232" refId="B28" refString="Inkhavilay, K, Sutcharit, C, Tongkerd, P, Panha, S, 2016. New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology 42 (4): 213 - 232" title="New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae)." volume="42" year="2016">Inkhavilay et al. 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A65B1C51A009264186F50E45DFD5FB80" pageId="0" pageNumber="157" type="remarks">
|
||
<paragraph id="5A96D014FBA167905EAFCF7834F3D9D9" pageId="0" pageNumber="157">Remarks.</paragraph>
|
||
<paragraph id="1FC0BDBBED23D2F7644FC8A88577CD27" pageId="0" pageNumber="157">
|
||
The two new species,
|
||
<taxonomicName id="D9E09658DA7EE06FABF912EEFBEA5FAE" lsidName="B. taiyaiorum" pageId="0" pageNumber="157" rank="species" species="taiyaiorum">
|
||
<emphasis id="05F72B1973488373634D044C2C312ADD" italics="true" pageId="0" pageNumber="157">B. taiyaiorum</emphasis>
|
||
</taxonomicName>
|
||
sp. nov. and
|
||
<taxonomicName id="24CD391D885B9CF4C20BC74932C25EC3" lsidName="B. lophiodera" pageId="0" pageNumber="157" rank="species" species="lophiodera">
|
||
<emphasis id="DA253D8363F6D533F00A0BE474247E71" italics="true" pageId="0" pageNumber="157">B. lophiodera</emphasis>
|
||
</taxonomicName>
|
||
sp. nov., are the first
|
||
<taxonomicName id="84178C9DAC3EF42D09879A6392DCB3DC" authorityName="Pilsbry & Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
|
||
<emphasis id="62BBCDB7B68F9B83130A4E423DA9CD58" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
species to be described from Myanmar. The strongly developed and separated parietal and angular lamellae, and lack of a tuba are the main characters underscoring the assignment of these species to
|
||
<taxonomicName id="4CC5C76A34E393C4621AC65CD2F56A2D" authorityName="Pilsbry & Vanatta" authorityYear="1900" class="Gastropoda" family="Hypselostomatidae" genus="Bensonella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Bensonella" order="Stylommatophora" pageId="0" pageNumber="157" phylum="Mollusca" rank="genus">
|
||
<emphasis id="DEC9FF0190223D8298D559AD37E062E6" italics="true" pageId="0" pageNumber="157">Bensonella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F10A46CB55FBACE1C0DAD54C679A8CFA" pageId="0" pageNumber="157">Though the spiral striae on the protoconch are obscured under the light microscope, they are very faint but detectable under the SEM microscope. Of all the type series examined, no hooked-shaped dentitions were observed, while the u-shaped plicae are present in all specimens. A constriction on the palatal wall near the aperture and the cervical crest is possibly an important trait in response to resistance to desiccation.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |