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<document id="F4B6FCB3BDB454479EFF092BF51ACB6A" ID-DOI="10.5281/zenodo.4645613" ID-ISSN="1942-1354" ID-Zenodo-Dep="4645613" ID-ZooBank="112A1F0B-1A82-4672-842B-A79A21F251D6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617040325248" checkinUser="felipe" docAuthor="Gordon, Robert D. &amp; Hanley, Guy A." docDate="2014" docId="03943535FFF74D08E6CAE9DEFA36FA5D" docLanguage="en" docName="InsectaMundi.0333.1-91.pdf" docOrigin="Insecta Mundi 2014 (333)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Glaresis donaldi Gordon and Hanley 2014, new species" docType="treatment" docVersion="3" lastPageNumber="48" masterDocId="FFAD4D4DFFC74D39E60AEF1DFF96FF86" masterDocTitle="Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea)" masterLastPageNumber="91" masterPageNumber="1" pageNumber="47" updateTime="1698895390200" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="ED2891C89F3927D83E75D556B4580296" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9C756439513B896EA910C02A7D0108DD">
<mods:title id="1A261B44B12A3B7DC16CFF13216B3E70">Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea)</mods:title>
</mods:titleInfo>
<mods:name id="910E81B37B4590E1587288CF79311242" type="personal">
<mods:role id="6F0D16F07C683865B913899207C0275A">
<mods:roleTerm id="6232338A9EF96A2D0861B2443DD080E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8401731B69F28DC9DD69F2FD405226EF">Gordon, Robert D.</mods:namePart>
</mods:name>
<mods:name id="A2EE93D6CA03B810B77B532C34894220" type="personal">
<mods:role id="D14E02422638701464CCC3F38E41809F">
<mods:roleTerm id="4053D5E4D2BD02B3FE9BFBBAC26FB327">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F4CE19CFED8B4CAADB39A3F90D9DF502">Hanley, Guy A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="12D3DA6C922D4B31B9B164B7B2F55E33">text</mods:typeOfResource>
<mods:relatedItem id="547A97F41AE6F36D75EEA34C35CE333F" type="host">
<mods:titleInfo id="BF4E6B5CD6D7C960C7821C5B9501C24A">
<mods:title id="D33761B636076F07E890DB137C87A16D">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="3CDC5051ED5DECDD85637220B706CE88">
<mods:date id="8CD7AA0EFDE2C4F800BCC4FBE65944F9">2014</mods:date>
<mods:detail id="943FACB1DBFDFA5FEBE62263AF43F9E4" type="pubDate">
<mods:number id="CE4793392C5E166C8D0DEB3411CCD412">2014-01-10</mods:number>
</mods:detail>
<mods:detail id="AE12586AADDBDE47717A82F89D03207F" type="volume">
<mods:number id="BEF5FB29F070DACD549BD1CF8A15391D">2014</mods:number>
</mods:detail>
<mods:detail id="44AE24526B230D7B1806319B90048F76" type="issue">
<mods:number id="706F84175B832D2123E53A866E611121">333</mods:number>
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<mods:classification id="FF98F7A338827537728A24D2BF316747">journal article</mods:classification>
<mods:identifier id="797B8D19E27CB4DF54125A4975474305" type="DOI">10.5281/zenodo.4645613</mods:identifier>
<mods:identifier id="9A38A745E4F3C5F16AB9058A93753D01" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="72196D8EA2945D24DB39A05A1D83E8BF" type="Zenodo-Dep">4645613</mods:identifier>
<mods:identifier id="2A27480D7F95889FDC59B6E791D51EE6" type="ZooBank">112A1F0B-1A82-4672-842B-A79A21F251D6</mods:identifier>
</mods:mods>
<treatment id="03943535FFF74D08E6CAE9DEFA36FA5D" ID-DOI="http://doi.org/10.5281/zenodo.4645778" ID-Zenodo-Dep="4645778" LSID="urn:lsid:plazi:treatment:03943535FFF74D08E6CAE9DEFA36FA5D" httpUri="http://treatment.plazi.org/id/03943535FFF74D08E6CAE9DEFA36FA5D" lastPageId="49" lastPageNumber="48" pageId="48" pageNumber="47">
<subSubSection id="C327D7A8FFF74D09E6CAE9DEFCCAF95D" box="[192,860,1731,1755]" pageId="48" pageNumber="47" type="nomenclature">
<paragraph id="8B828423FFF74D09E6CAE9DEFCCAF95D" blockId="48.[192,860,1731,1755]" box="[192,860,1731,1755]" pageId="48" pageNumber="47">
<heading id="D0CA334FFFF74D09E6CAE9DEFCCAF95D" bold="true" box="[192,860,1731,1755]" fontSize="10" level="2" pageId="48" pageNumber="47" reason="3">
<emphasis id="B9495831FFF74D09E6CAE9DEFCCAF95D" bold="true" box="[192,860,1731,1755]" pageId="48" pageNumber="47">
<taxonomicName id="4C3DFFA0FFF74D09E6CAE9DEFD27F95D" ID-CoL="3G5YC" authority="Gordon and Hanley" authorityName="Gordon and Hanley" authorityYear="2014" box="[192,689,1731,1755]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="47" phylum="Arthropoda" rank="species" species="donaldi" status="sp. nov.">
<emphasis id="B9495831FFF74D09E6CAE9DEFE37F95D" bold="true" box="[192,417,1731,1755]" italics="true" pageId="48" pageNumber="47">Glaresis donaldi</emphasis>
Gordon and Hanley
</taxonomicName>
,
<taxonomicNameLabel id="A27AE54AFFF74D09E4B6E9DEFCCAF95D" box="[700,860,1731,1755]" pageId="48" pageNumber="47" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF74D08E6CAE81EFD95FBDD" lastPageId="49" lastPageNumber="48" pageId="48" pageNumber="47" type="description">
<paragraph id="8B828423FFF74D08E6CAE81EFCA7FC7D" blockId="48.[192,1440,1795,1915]" lastBlockId="49.[192,1441,195,1051]" lastPageId="49" lastPageNumber="48" pageId="48" pageNumber="47">
<emphasis id="B9495831FFF74D09E6CAE81EFEF5F89D" bold="true" box="[192,355,1795,1819]" pageId="48" pageNumber="47">Description</emphasis>
.
<emphasis id="B9495831FFF74D09E779E81EFE41F89D" bold="true" box="[371,471,1795,1819]" pageId="48" pageNumber="47">Female</emphasis>
. Length
<quantity id="4CC529C6FFF74D09E44CE81EFD33F89D" box="[582,677,1795,1819]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="48" pageNumber="47" unit="mm" value="2.8">2.8 mm</quantity>
, width
<quantity id="4CC529C6FFF74D09E50BE81EFCF7F89D" box="[769,865,1795,1819]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="48" pageNumber="47" unit="mm" value="1.5">1.5 mm</quantity>
; body form short, wide, strongly widened from elytral base to apical 1/3 (
<figureCitation id="130698A6FFF74D09E7FEE83EFDF7F8BD" box="[500,609,1827,1851]" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="48" pageNumber="47">Fig. 32A</figureCitation>
). Color dark brown. Head with clypeal surface and anteromedian 1/3 of frons slightly shiny, with finely reticulate sculpture and small, sparse tubercles, posterior portion of frons and vertex feebly shiny, with coarse reticulate sculpture; setae short, indistinct. Clypeal apex truncate, with small, evenly spaced tubercles, appearing somewhat serrate, lateral angles oblique, angulate (
<figureCitation id="130698A6FFF64D08E6C3EFFEFEA4FF7D" box="[201,306,227,251]" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="49" pageNumber="48">Fig. 32B</figureCitation>
). Mandible pair symmetrical; mesal tooth strong; lateral prominence strong, pronounced; outer margin slightly abruptly rounded. Pronotum with deep, long, transverse fovea in anterior 1/4 extended completely across pronotum, small slightly transverse fovea on each side of middle in anterior 1/2, wide, deep fovea on each side medially near lateral margin, deep central furrow extended from base to transverse furrow; surface densely rugose, surface with sparse, short, slightly irregular, setae-bearing carinae, setae about 1/2 length of carinae (
<figureCitation id="130698A6FFF64D08E497EE9EFC9FFE1D" box="[669,777,387,411]" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="49" pageNumber="48">Fig. 32C</figureCitation>
); anterior and lateral pronotal margins crenulate, not bordered, lateral margin slightly crenulate. Elytra with surface shiny, finely microreticulate; all striae smooth, strongly carinate, carinae appearing continuous but narrowly interrupted adjacent to space between interval punctures, each carinal segment bearing an apical seta about 1/2 length of segment; intervals with deep, round punctures (
<figureCitation id="130698A6FFF64D08E481ED1EFD62FD9D" box="[651,756,515,539]" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="49" pageNumber="48">Fig. 32E</figureCitation>
). Metasternum long, dull, finely, densely microreticulate, median surface not tuberculate medially, most of remaining surface with short, dense, irregular carinae forming reticulate pattern, median area flat, with median carina extended 2/3 distance to mesocoxae; metasternal groove narrow, shallow, inner margin rounded, outer margin with weak ridge (
<figureCitation id="130698A6FFF64D08E32FED7EFA02FDFD" box="[1317,1428,611,635]" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="49" pageNumber="48">Fig. 32D</figureCitation>
). Lateral protibial teeth not evenly spaced, 2nd and 3rd teeth close together. Mesotibia with 4 short, closely spaced spines from base to apex of posterolateral emargination, tibia strongly projecting at apex, projection with acute apex (
<figureCitation id="130698A6FFF64D08E401EDDEFDE1FD5D" box="[523,631,707,731]" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="49" pageNumber="48">Fig. 32D</figureCitation>
). Posterior metatrochanteral margin smooth, with indistinct row of tubercles on surface of posterior flange; posterosuperior surface of metatrochanter with single large tooth, tooth not visible directly in ventral view. Metafemoral surface densely, coarsely microreticulate, with sparse, small, round, setae-bearing tubercles in anterior 1/2; width to length ratio 1.0:1.6, with wide flange on anterior margin, posterior margin without teeth; posterosuperior margin without teeth (
<figureCitation id="130698A6FFF64D08E37AEC5EFF65FCFD" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="49" pageNumber="48">Fig. 32F</figureCitation>
). Metatibia narrow at base, abruptly widened before apex, surface entirely, strongly microreticulate, lateral margin with narrow row of small teeth from near base to lateral projection, with distinct posteromedian lateral projection, medially with uniform row of large, widely spaced, tubercles, tubercles posterior to posteromedian lateral projection transversely oblique, inner margin smooth, pubescent (
<figureCitation id="130698A6FFF64D08E37AECDEFF62FC7D" captionStart="Figures 32" captionStartId="87.[192,286,1344,1366]" captionTargetBox="[207,1413,211,1311]" captionTargetId="figure-15@87.[192,1440,192,1346]" captionTargetPageId="87" captionText="Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.4645717" httpUri="https://zenodo.org/record/4645717/files/figure.png" pageId="49" pageNumber="48">Fig. 32F</figureCitation>
). Apex of 5th abdominal ventrite not examined.
</paragraph>
<paragraph id="8B828423FFF64D08E6FAEB1EFE52FB9D" blockId="49.[192,1441,195,1051]" box="[240,452,1027,1051]" pageId="49" pageNumber="48">
<emphasis id="B9495831FFF64D08E6FAEB1EFEA5FB9D" bold="true" box="[240,307,1027,1051]" pageId="49" pageNumber="48">Male</emphasis>
. Unknown.
</paragraph>
<paragraph id="8B828423FFF64D08E6CAEB5EFD95FBDD" blockId="49.[192,515,1091,1115]" box="[192,515,1091,1115]" pageId="49" pageNumber="48">
<emphasis id="B9495831FFF64D08E6CAEB5EFED4FBDD" bold="true" box="[192,322,1091,1115]" pageId="49" pageNumber="48">Variation</emphasis>
.
<typeStatus id="54863A81FFF64D08E758EB5EFE57FBDD" box="[338,449,1091,1115]" pageId="49" pageNumber="48" type="holotype">Holotype</typeStatus>
only.
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF64D08E6CAEB9EFDF9FB3D" pageId="49" pageNumber="48" type="materials_examined">
<paragraph id="8B828423FFF64D08E6CAEB9EFDF9FB3D" blockId="49.[192,1440,1155,1211]" pageId="49" pageNumber="48">
<emphasis id="B9495831FFF64D08E6CAEB9EFE17FB1D" bold="true" box="[192,385,1155,1179]" pageId="49" pageNumber="48">Type material</emphasis>
.
<materialsCitation id="3B558E7EFFF64D08E785EB9EFDF9FB3D" collectionCode="TAMU" collectorName="Mi. SW &amp; Mastro &amp; Schaffner" country="Mexico" elevation="9" location="Tehuacan" municipality="July" pageId="49" pageNumber="48" specimenCount="1" stateProvince="Mexico" typeStatus="holotype">
<typeStatus id="54863A81FFF64D08E785EB9EFE68FB1D" box="[399,510,1155,1179]" pageId="49" pageNumber="48" type="holotype">Holotype</typeStatus>
female:
<collectingRegion id="49F94AC1FFF64D08E46FEB9EFD29FB1D" box="[613,703,1155,1179]" country="Mexico" name="Mexico" pageId="49" pageNumber="48">Mexico</collectingRegion>
.
<emphasis id="B9495831FFF64D08E4C3EB9EFCBCFB1D" bold="true" box="[713,810,1155,1179]" pageId="49" pageNumber="48">
<collectingRegion id="49F94AC1FFF64D08E4C3EB9EFCBCFB1D" box="[713,810,1155,1179]" country="Mexico" name="Puebla" pageId="49" pageNumber="48">Puebla</collectingRegion>
</emphasis>
:
<collectingCountry id="F32AC4B3FFF64D08E533EB9EFC39FB1D" box="[825,943,1155,1179]" name="Mexico" pageId="49" pageNumber="48">MEXICO</collectingCountry>
:
<collectingRegion id="49F94AC1FFF64D08E5B2EB9EFB86FB1D" box="[952,1040,1155,1179]" country="Mexico" name="Puebla" pageId="49" pageNumber="48">Puebla</collectingRegion>
, 6
<collectorName id="26C8E1F5FFF64D08E23BEB9EFB1EFB1D" box="[1073,1160,1155,1179]" pageId="49" pageNumber="48">Mi. SW</collectorName>
.
<location id="8EE2D2F8FFF64D08E298EB9EFA9BFB1D" LSID="urn:lsid:plazi:treatment:03943535FFF74D08E6CAE9DEFA36FA5D:8EE2D2F8FFF64D08E298EB9EFA9BFB1D" box="[1170,1293,1155,1179]" country="Mexico" municipality="July" name="Tehuacan" pageId="49" pageNumber="48" stateProvince="Mexico">Tehuacan</location>
,
<collectingMunicipality id="6BE61E59FFF64D08E312EB9EFADBFB1D" box="[1304,1357,1155,1179]" pageId="49" pageNumber="48">July</collectingMunicipality>
<quantity id="4CC529C6FFF64D08E35EEB9EFA36FB1D" box="[1364,1440,1155,1179]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="49" pageNumber="48" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">
<elevation id="00106310FFF64D08E35EEB9EFA36FB1D" box="[1364,1440,1155,1179]" metricMagnitude="0" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="49" pageNumber="48" unit="m" value="9.0" valueMax="10.0" valueMin="8.0">8-10m</elevation>
</quantity>
1973,
<collectorName id="26C8E1F5FFF64D08E703EBB9FEF6FB3D" box="[265,352,1188,1211]" pageId="49" pageNumber="48">Mastro</collectorName>
&amp;
<collectorName id="26C8E1F5FFF64D08E789EBBEFE6CFB3D" box="[387,506,1187,1211]" pageId="49" pageNumber="48">Schaffner</collectorName>
(
<collectionCode id="ED2C1CE6FFF64D08E402EBBEFDF4FB3D" box="[520,610,1187,1211]" pageId="49" pageNumber="48">TAMU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF64D08E6CAEBFEFBE0FAFD" pageId="49" pageNumber="48" type="discussion">
<paragraph id="8B828423FFF64D08E6CAEBFEFB1CFABD" blockId="49.[192,1440,1251,1403]" pageId="49" pageNumber="48">
<emphasis id="B9495831FFF64D08E6CAEBFEFEACFB7D" bold="true" box="[192,314,1251,1275]" pageId="49" pageNumber="48">Remarks</emphasis>
. The unique female
<typeStatus id="54863A81FFF64D08E421EBFEFD04FB7D" box="[555,658,1251,1275]" pageId="49" pageNumber="48" type="holotype">holotype</typeStatus>
is distinguished from other known species in this subgroup by the metatibia having a distinct posteromedian lateral projection. It is also smaller in size and has a ventral surface somewhat more strongly, densely microreticulate than the other species.
</paragraph>
<paragraph id="8B828423FFF64D08E6FAEA5EFBE0FAFD" blockId="49.[192,1440,1251,1403]" pageId="49" pageNumber="48">
This specimen represents the first published record of a
<taxonomicName id="4C3DFFA0FFF64D08E591EA5EFB96FADD" authorityName="Erichson" authorityYear="1848" box="[923,1024,1347,1371]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="48" phylum="Arthropoda" rank="genus">
<emphasis id="B9495831FFF64D08E591EA5EFB96FADD" box="[923,1024,1347,1371]" italics="true" pageId="49" pageNumber="48">Glaresis</emphasis>
</taxonomicName>
species from southern
<collectingCountry id="F32AC4B3FFF64D08E313EA5EFAE5FADD" box="[1305,1395,1347,1371]" name="Mexico" pageId="49" pageNumber="48">Mexico</collectingCountry>
. All other Mexican species are recorded from northern
<collectingCountry id="F32AC4B3FFF64D08E52CEA7EFCEAFAFD" box="[806,892,1379,1403]" name="Mexico" pageId="49" pageNumber="48">Mexico</collectingCountry>
and
<collectingRegion id="49F94AC1FFF64D08E5BDEA7EFBE5FAFD" box="[951,1139,1379,1403]" country="Mexico" name="Baja California" pageId="49" pageNumber="48">Baja California</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C327D7A8FFF64D08E6CAEABEFA36FA5D" pageId="49" pageNumber="48" type="etymology">
<paragraph id="8B828423FFF64D08E6CAEABEFA36FA5D" blockId="49.[192,1440,1443,1499]" pageId="49" pageNumber="48">
<emphasis id="B9495831FFF64D08E6CAEABEFEC5FA3D" bold="true" box="[192,339,1443,1467]" pageId="49" pageNumber="48">Etymology</emphasis>
. This species is named in honor of the late Donald Whitehead, coleopterist, old
<collectingCountry id="F32AC4B3FFF64D08E30AEABEFAC0FA3D" box="[1280,1366,1443,1467]" name="Mexico" pageId="49" pageNumber="48">Mexico</collectingCountry>
hand, and former colleague of one of us (RDG). A good friend, excellent scientist, and good man to travel with.
</paragraph>
</subSubSection>
</treatment>
</document>
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