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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.220.2178" ID-GBIF-Dataset="07cc435f-a623-4a17-b510-a38bf0569844" ID-PMC="PMC3459032" ID-Pensoft-Pub="1313-2970-220-1" ID-PubMed="23077429" ModsDocAuthor="" ModsDocDate="2012" ModsDocID="1313-2970-220-1" ModsDocOrigin="ZooKeys 220" ModsDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" checkinTime="1451248758882" checkinUser="pensoft" docAuthor="Henry, Thomas J." docDate="2012" docId="1F4B0239A5762E0A64753B1998011304" docLanguage="en" docName="ZooKeys 220: 1-114" docOrigin="ZooKeys 220" docSource="http://dx.doi.org/10.3897/zookeys.220.2178" docTitle="Tytthus pallidus Henry, sp. n." docType="treatment" docVersion="4" lastPageNumber="47" masterDocId="FFA2D657D204FFEEFF8DF82F5143FF91" masterDocTitle="Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)" masterLastPageNumber="114" masterPageNumber="1" pageNumber="47" updateTime="1668154436991" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the Plant Bug Genus Tytthus (Hemiptera, Heteroptera, Miridae, Phylinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Henry, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="volume">
<mods:number>220</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>114</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.220.2178</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.220.2178</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-220-1</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="152036998" LSID="urn:lsid:zoobank.org:act:E2C46E49-A43D-4C0B-9742-FBAFF41F3C2D" httpUri="http://treatment.plazi.org/id/1F4B0239A5762E0A64753B1998011304" lastPageNumber="47" pageId="46" pageNumber="47">
<subSubSection pageId="46" pageNumber="47" type="nomenclature">
<paragraph pageId="46" pageNumber="47">
<taxonomicName LSID="urn:lsid:zoobank.org:act:E2C46E49-A43D-4C0B-9742-FBAFF41F3C2D" authority="Henry" class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus pallidus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pallidus">
<pageBreakToken pageId="46" pageNumber="47" start="start">Tytthus</pageBreakToken>
pallidus Henry
</taxonomicName>
<taxonomicNameLabel pageId="46" pageNumber="47">sp. n.</taxonomicNameLabel>
Figs 28, 29160-163
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="diagnosis">
<paragraph pageId="46" pageNumber="47">Diagnosis.</paragraph>
<paragraph pageId="46" pageNumber="47">This species is recognized by the combination of the dark brown head, pronotum, and scutellum; the pale antennal segment I, with only the base fuscous, the brown antennal segment II; and the uniformly pale yellow legs. All known specimens are macropterous.</paragraph>
<paragraph pageId="46" pageNumber="47">
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus pallidus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="pallidus">Tytthus pallidus</taxonomicName>
keys out with
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus piceus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="piceus">Tytthus piceus</taxonomicName>
based on the pale antennal segment I and the pale hemelytra with smoky-brown shading. It can be distinguished from
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus piceus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="piceus">Tytthus piceus</taxonomicName>
by the broader head, the longer antennal segment I that is longer than the interocular width, and the less prominent calli lacking a glaucous sheen.
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="description">
<paragraph pageId="46" pageNumber="47">Description.</paragraph>
<paragraph pageId="46" pageNumber="47">Holotype male (Fig. 28): Length to apex of hemelytron 2.88 mm, length to base of cuneus 2.21 mm, width across hemelytra 0.88 mm. Head: Length 0.35 mm, width across eyes 0.61 mm, interocular width 0.29 mm. Labium: Length 1.31 mm. Antenna: Segment I length 0.35 mm, II 1.22 mm, III and IV missing. Pronotum: Length 0.35 mm, basal width 0.75 mm.</paragraph>
<paragraph pageId="46" pageNumber="47">
Coloration: Head: Uniformly dark brown; pale interocular spot found in all other species apparently absent; fuscous to dark reddish brown. Labium: Pale brownish yellow. Antenna: Segment I uniformly pale yellow, with a dark brown or fuscous ring at base; segments
<normalizedToken originalValue="IIIV">II-IV</normalizedToken>
dark brown. Pronotum: Uniformly dark brown. Mesoscutum: Uniformly dark brown. Scutellum: Uniformly dark brown. Hemelyton: Uniformly pale translucent brown. Ostiolar evaporative area: Dark brown to fuscous. Ventral surface: Thorax dark brown to dark reddish brown; abdomen dark reddish brown on segment II, III, and genital capsule, slightly paler in between. Legs: Coxa pale yellow, reddish brown at bases; femora, tibiae, tarsi, and claws uniformly pale yellow.
</paragraph>
<paragraph pageId="46" pageNumber="47">Structure, texture, and vestiture: Antenna: Segment I set with a few, scattered, recumbent setae and two erect, subapical, bristlelike setae. Labium: Extending beyond metacoxae to abdominal segment II or III. Pronotum: Shiny, impunctate; anterior angles rounded; lateral margins weakly concave, flaring at posterior angles; posterior margin distinctly sinuate; calli weakly swollen; set with relatively long, semierect setae. Mesoscutum: Impunctate, broadly exposed; with a few scattered semierect setae. Scutellum: Weakly shining, impunctate; equilateral; set with scattered, semierect setae. Hemelytra: Macropterous, cuneus and membrane fully developed, extending well beyond apex of abdomen.</paragraph>
<paragraph pageId="46" pageNumber="47">Male genitalia: Left paramere (Fig. 160): Mitt-shaped; right arm long, broad, apically blunt; left arm short, apically acute. Right paramere (Fig. 161): Round. Endosoma: Teneral and damaged; not drawn. Phallotheca (Fig. 162): slender, apically acute.</paragraph>
<paragraph pageId="46" pageNumber="47">Female (n = 4) (Fig. 29): Length to apex of hemelytron 2.69-3.33 mm, length to base of cuneus 2.05-2.40 mm, width across hemelytra 0.82-1.02 mm. Head: Length 0.32-0.38 mm, width across eyes 0.56-0.64 mm, interocular width 0.29-0.30 mm. Labium: Length 1.12-1.44 mm. Antenna: Segment I length 0.26-0.34 mm, II 0.85-1.23 mm, III 0.61-0.78 mm, IV 0.51-0.56 mm. Pronotum: Length 0.29-0.37 mm, basal width 0.69-0.85 mm.</paragraph>
<caption pageId="46" pageNumber="47">
<paragraph pageId="46" pageNumber="47">
Figures 163-170. Male genitalia 163-166
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus parviceps" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="parviceps">Tytthus parviceps</taxonomicName>
163 left paramere 164 right paramere 165 endosoma 166 phallotheca 167-170
<taxonomicName class="Insecta" family="Miridae" genus="Tytthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Tytthus piceus" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="piceus">Tytthus piceus</taxonomicName>
167 left paramere 168 right paramere 169 endosoma 170 phallotheca.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="etymology">
<paragraph pageId="46" pageNumber="47">Etymology.</paragraph>
<paragraph pageId="46" pageNumber="47">
The specific epithet
<normalizedToken originalValue="“pallidus”">&quot;pallidus&quot;</normalizedToken>
refers to the characteristic pale yellow first antennal segment.
</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="host">
<paragraph pageId="46" pageNumber="47">Host.</paragraph>
<paragraph pageId="46" pageNumber="47">Unknown.</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="distribution">
<paragraph pageId="46" pageNumber="47">Distribution.</paragraph>
<paragraph pageId="46" pageNumber="47">Known from Brazil and Panama.</paragraph>
</subSubSection>
<subSubSection pageId="46" pageNumber="47" type="type material">
<paragraph pageId="46" pageNumber="47">Type material.</paragraph>
<paragraph pageId="46" pageNumber="47">Holotype ♂ (00162199) (USNM): PANAMA:Darien: [Pinogana District], El Real, 19 Mar. 1953, F. S. Blanton, 1 ♂ (USNM). El Real, 08 Aug 1952, F. S. Blanton, 2 ♀♀ (00161392, 00161393) (USNM). Paratypes:BRAZIL: Amazonas: Reserva Ducke, 25 km NNE of Manaus, 120 m, 26 Jul 1973, R.T. Schuh, 2 ♀♀ (00165829) (AMNH).</paragraph>
</subSubSection>
</treatment>
</document>