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<document ID-DOI="http://dx.doi.org/10.3897/mycokeys.32.23277" ID-GBIF-Dataset="2708d4e6-390d-4625-8bbd-089abd4c73bb" ID-PMC="PMC5904377" ID-Pensoft-Pub="1314-4049-32-49" ID-PubMed="29681737" ModsDocAuthor="" ModsDocDate="2018" ModsDocID="1314-4049-32-49" ModsDocOrigin="MycoKeys 32" ModsDocTitle="A new species of Stamnaria (Leotiomycetes, Helotiales) from Western Siberia" checkinTime="1521626544417" checkinUser="pensoft" docAuthor="Haelewaters, Danny, Filippova, Nina V. &amp; Baral, Hans-Otto" docDate="2018" docId="D2CA6CB8185D00C4384C7D0F2A509542" docLanguage="en" docName="MycoKeys 32: 49-63" docOrigin="MycoKeys 32" docSource="http://dx.doi.org/10.3897/mycokeys.32.23277" docTitle="Stamnaria yugrana Filippova, Haelew. &amp; Baral, sp. nov." docType="treatment" docVersion="4" lastPageNumber="52" masterDocId="0D79FFE0FFAF2117FFC4FFEAD25BF73D" masterDocTitle="A new species of Stamnaria (Leotiomycetes, Helotiales) from Western Siberia" masterLastPageNumber="63" masterPageNumber="49" pageNumber="52" updateTime="1668135967939" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new species of Stamnaria (Leotiomycetes, Helotiales) from Western Siberia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Haelewaters, Danny</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Filippova, Nina V.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Baral, Hans-Otto</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>MycoKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2018</mods:date>
<mods:detail type="volume">
<mods:number>32</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>49</mods:start>
<mods:end>63</mods:end>
</mods:extent>
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<mods:location>
<mods:url>http://dx.doi.org/10.3897/mycokeys.32.23277</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/mycokeys.32.23277</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-4049-32-49</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="142462580" LSID="urn:lsid:plazi:treatment:D2CA6CB8185D00C4384C7D0F2A509542" httpUri="http://treatment.plazi.org/id/D2CA6CB8185D00C4384C7D0F2A509542" lastPageNumber="52" pageId="3" pageNumber="52">
<subSubSection pageId="3" pageNumber="52" type="nomenclature">
<paragraph pageId="3" pageNumber="52">
<taxonomicName LSID="823742" authority="Filippova, Haelew. &amp; Baral" class="Leotiomycetes" family="Helotiaceae" genus="Stamnaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Stamnaria yugrana" order="Helotiales" pageId="3" pageNumber="52" phylum="Ascomycota" rank="species" species="yugrana">Stamnaria yugrana Filippova, Haelew. &amp; Baral</taxonomicName>
<taxonomicNameLabel pageId="3" pageNumber="52">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="52" type="diagnosis">
<paragraph pageId="3" pageNumber="52">Diagnosis.</paragraph>
<paragraph pageId="3" pageNumber="52">
Characterised by the presence of both lanceolate and cylindrical paraphyses, fusoid-clavate ascospores with a length/width ratio of predominantly&gt;4 and free-ending hyphae at the inner excipulum of the tube-shaped, even collar. Saprophytic on dead branches of
<taxonomicName lsidName="E. sylvaticum" pageId="3" pageNumber="52" rank="species" species="sylvaticum">E. sylvaticum</taxonomicName>
.
</paragraph>
<caption pageId="3" pageNumber="52">
<paragraph pageId="3" pageNumber="52">
Figure 1. Study site of
<taxonomicName class="Leotiomycetes" family="Calloriaceae" genus="Stamnaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Stamnaria yugrana" order="Rhytismatales" pageId="3" pageNumber="52" phylum="Ascomycota" rank="species" species="yugrana">Stamnaria yugrana</taxonomicName>
growing on litter of
<taxonomicName class="Polypodiopsida" family="Equisetaceae" genus="Equisetum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Equisetum sylvaticum" order="Equisetales" pageId="3" pageNumber="52" phylum="Tracheophyta" rank="species" species="sylvaticum">Equisetum sylvaticum</taxonomicName>
in Western Siberia near the Khanty-Mansiysk town.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="52" type="types">
<paragraph pageId="3" pageNumber="52">Types.</paragraph>
<paragraph pageId="3" pageNumber="52">
Holotype: Russia, Western Siberia, Khanty-Mansi Autonomous Okrug - Yugra, 25 km ENE of Khanty-Mansiysk town, 1.2 km SSE of Shapsha village, 61.07929&quot;N, 69.46925&quot;E, alt. 40 m, 9 Jun 2012, leg. N.V. Filippova, on fallen side branches of
<taxonomicName class="Polypodiopsida" family="Equisetaceae" genus="Equisetum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Equisetum sylvaticum" order="Equisetales" pageId="3" pageNumber="52" phylum="Tracheophyta" rank="species" species="sylvaticum">Equisetum sylvaticum</taxonomicName>
L. lying amongst other forest litter in a mixed coniferous-deciduous forest; Biological Collection of Yugra State University (YSU-F-03519). Isotypes: LE-295215; FH 01146308. Paratypes: ibid., 16 Jun 2008 (YSU-F-00097, material lost; LE-295060); ibid., 22 Feb 2014, substrate collected from under snow and grown in a moist chamber (YSU-F-04933); ibid., 25 Feb 2015 (YSU-F-06579; LE-296061).
</paragraph>
<caption pageId="3" pageNumber="52">
<paragraph pageId="3" pageNumber="52">
Figure 2. Apothecia of
<taxonomicName class="Leotiomycetes" family="Calloriaceae" genus="Stamnaria" higherTaxonomySource="CoL" kingdom="Fungi" lsidName="Stamnaria yugrana" order="Rhytismatales" pageId="3" pageNumber="52" phylum="Ascomycota" rank="species" species="yugrana">Stamnaria yugrana</taxonomicName>
on side branches of
<taxonomicName class="Polypodiopsida" family="Equisetaceae" genus="Equisetum" higherTaxonomySource="CoL" kingdom="Plantae" lsidName="Equisetum sylvaticum" order="Equisetales" pageId="3" pageNumber="52" phylum="Tracheophyta" rank="species" species="sylvaticum">Equisetum sylvaticum</taxonomicName>
: A Apothecia grown in situ under well-lit conditions B Apothecia grown in shady conditions after incubation in a moist chamber C Median section through an apothecium after incubation in a moist chamber (dead, in CR). A from YSU-F-03519, B from YSU-F-04933, C from YSU-F00097. Scale bars: A 1.0 mm, B, C 0.5 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="52" type="description">
<paragraph pageId="3" pageNumber="52">Description.</paragraph>
<paragraph pageId="3" pageNumber="52">
Apothecia urn-shaped, stipitate, 0.25-0.6 mm in diameter when mature, 0.5-1 mm high, varying depending on light conditions, being stouter with shorter stipe when substrate exposed to sunlight; receptacle light yellow(-ochraceous) when fresh, with even, whitish collar ~80-120
<normalizedToken originalValue="µm">µm</normalizedToken>
high, stipe pale yellowish-translucent, 100-380
<normalizedToken originalValue="×">x</normalizedToken>
130-300
<normalizedToken originalValue="µm">µm</normalizedToken>
, receptacle becoming light brown on drying; scattered to moderately gregarious, often abundant on the branches. Ectal excipulum outer layer *40-45
<normalizedToken originalValue="µm">µm</normalizedToken>
thick at middle flanks and margin, made up of strongly gelatinous tissue of loose, parallel to wavy hyphae 2-3
<normalizedToken originalValue="μm">μm</normalizedToken>
broad, septate, embedded in abundant gelatinous matrix (textura oblita); inner layer *~45
<normalizedToken originalValue="µm">µm</normalizedToken>
thick at middle flanks, made up of textura prismatica-porrecta running parallel to outside, cells at middle flanks *17-44
<normalizedToken originalValue="×">x</normalizedToken>
7-12
<normalizedToken originalValue="μm">μm</normalizedToken>
, slightly narrower in the collar region, much narrower in stipe (*20-45
<normalizedToken originalValue="×">x</normalizedToken>
3-5
<normalizedToken originalValue="µm">µm</normalizedToken>
); the inner layer of the collar composed of narrow hyphae, free upper part of these hyphae internally covered by lateral cellular outgrowths *2-5
<normalizedToken originalValue="×">x</normalizedToken>
1.5-2
<normalizedToken originalValue="µm">µm</normalizedToken>
. Medullary excipulum well developed, of dense, parallel, septate hyphae (textura porrecta) without gel, cells *65-90
<normalizedToken originalValue="×">x</normalizedToken>
2.5-6 (-7.7)
<normalizedToken originalValue="μm">μm</normalizedToken>
; subhymenium well developed (*20-30 thick), of intricate hyphae *2
<normalizedToken originalValue="μm">μm</normalizedToken>
broad. Asci cylindrical, developing from croziers which are difficult to see in mature asci, with apical thickening enclosing a hemiamyloid ring of Calycina-type (rb: dirty red at high, blue at low concentration), *146
<normalizedToken originalValue="×">x</normalizedToken>
12.5 [123-159 (-206)
<normalizedToken originalValue="×">x</normalizedToken>
11.7-13.5]
<normalizedToken originalValue="μm">μm</normalizedToken>
, †98
<normalizedToken originalValue="×">x</normalizedToken>
9 (90-110
<normalizedToken originalValue="×">x</normalizedToken>
8-10.5)
<normalizedToken originalValue="μm">μm</normalizedToken>
, 8-spored, spores *obliquely biseriate. Paraphyses of two types: (1) lanceolate, exceeding asci for *12-20
<normalizedToken originalValue="µm">µm</normalizedToken>
when young and *30-40
<normalizedToken originalValue="μm">μm</normalizedToken>
when fully developed, septate in lower part, non-septate in broad upper part, with quite acute tip, in young paraphyses with granular (multiguttulate) vacuolar content of moderate refractivity (VBs), later replaced by larger non-refractive vacuoles, *5-7 (†3-6)
<normalizedToken originalValue="μm">μm</normalizedToken>
broad in upper part; (2) cylindrical, more abundant, not exceeding the asci, *2.3-3
<normalizedToken originalValue="μm">μm</normalizedToken>
broad above, septate, with obtuse tip, rarely branched below and scarcely enlarged in upper segment, without VBs, with pale yellow-orange pigment in middle and lower part. Ascospores fusoid-clavate, slightly to distinctly heteropolar, with rounded to obtuse ends, usually without any gel around, filled with granular oil content in both halves, leaving a central zone for the single nucleus, variable in length, *19.8
<normalizedToken originalValue="×">x</normalizedToken>
4.8 (16.5-24.5
<normalizedToken originalValue="×">x</normalizedToken>
4.2-5.6), n=18, Q=4.1 (YSU-F-04933); †20.5
<normalizedToken originalValue="×">x</normalizedToken>
4.0 (17.2-24.2
<normalizedToken originalValue="×">x</normalizedToken>
3.6-4.6)
<normalizedToken originalValue="μm">μm</normalizedToken>
, n=37, Q=5.1 (YSU-F-03519, YSU-F-04933).
</paragraph>
<caption pageId="3" pageNumber="52">
<paragraph pageId="3" pageNumber="52">
Figure 3. A1 Young living paraphysis with multiguttulate, medium refractive vacuolar content (VBs), A2 Mature living paraphysis with large non-refractive vacuoles A3 Dead paraphysis with small oil drops (LBs), vacuoles disappeared B Paraphyses of cylindrical and lanceolate type C Mature living asci D Mature ejected ascospores with granular lipid content of minute LBs (confluent in dead state) E Ascus bases with croziers F Apices of mature and immature ascus (stained in IKI). All * and F from YSU-F-04933, all other † from YSU-F-00097. Scale bars: A, D, E, F 10
<normalizedToken originalValue="μm">μm</normalizedToken>
, B, C 20
<normalizedToken originalValue="μm">μm</normalizedToken>
.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="3" pageNumber="52" type="etymology">
<paragraph pageId="3" pageNumber="52">Etymology.</paragraph>
<paragraph pageId="3" pageNumber="52">Referring to Yugra, the historical name of the region (currently &quot;Khanty-Mansi Autonomous Okrug - Yugra&quot;).</paragraph>
</subSubSection>
<subSubSection pageId="3" pageNumber="52" type="distribution">
<paragraph pageId="3" pageNumber="52">Distribution.</paragraph>
<paragraph pageId="3" pageNumber="52">Known only from the type locality.</paragraph>
</subSubSection>
</treatment>
</document>