treatments-xml/data/95/65/D0/9565D05FFFCFFFC5FCDA5934FD789B99.xml
2024-06-21 12:44:36 +02:00

346 lines
42 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="875EC63AF5531EDC5CFD4301AD1B4926" ID-DOI="10.26107/RBZ-2020-0052" ID-ISSN="2345-7600" ID-Zenodo-Dep="4576861" ID-ZooBank="C7D4631B-9B02-4E6E-9665-76CCFECA86A0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614780803714" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Hunyadi, András, Grego, Jozef, Reischütz, Alexander, Buczkó, Krisztina &amp; Vermeulen, Jaap J." docDate="2020" docId="9565D05FFFCFFFC5FCDA5934FD789B99" docLanguage="en" docName="RafflesBZool.68.350-368.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Clostophis incurvus Pall-Gergely &amp; Vermeulen 2020, new species" docType="treatment" docVersion="3" lastPageNumber="358" masterDocId="695CA827FFCDFFCDFF9A5A29FFDE9B6C" masterDocTitle="Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species" masterLastPageNumber="368" masterPageNumber="350" pageNumber="352" updateTime="1698878356270" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="962579FDE24A8D498666E5300E9BAB43" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3A9186245270FB22DBFB398C1689EFD9">
<mods:title id="B67D6AD62981F96B600A906D6C3180F8">Clostophis Benson, 1860, is not a monotypic diplommatinid but a speciose hypselostomatid (Gastropoda: Eupulmonata), with descriptions of six new species</mods:title>
</mods:titleInfo>
<mods:name id="C98DFE16EA36B2E5EE7B5966246D5210" type="personal">
<mods:role id="EEF3E3E785AFA400178EDAEC1ED6CA33">
<mods:roleTerm id="890E8439A683C079E48D6B107123AFC8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="159180D5572248763DEF9A02CFA1A2A6">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="0EE651517E5D77F0DA6A126A013AEA74" type="personal">
<mods:role id="2EEEA94D6E3903866187960202C06B9F">
<mods:roleTerm id="7E51D37B6A58E5FF57BA271F7A87E930">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FAFB3A319CA85A177D0D21F182765661">Hunyadi, András</mods:namePart>
</mods:name>
<mods:name id="A77FEC8C681EFC6182EAC2B49C11DEB9" type="personal">
<mods:role id="847E0D636A9BF95D60875828D7397802">
<mods:roleTerm id="4C45BF8DC85FDBFE18FD2BF5D1954571">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C84151388C9D094A792BFCDA39E0CFCB">Grego, Jozef</mods:namePart>
</mods:name>
<mods:name id="3F061E5F040535D42EC3C0CFEF9E1A5F" type="personal">
<mods:role id="C8F7E08FF7672A309D3E4776ACE4E91F">
<mods:roleTerm id="50D7E88995C7FBA700923F8465352038">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A4D50C89AE7376BDC0DE13F629B401E4">Reischütz, Alexander</mods:namePart>
</mods:name>
<mods:name id="43CDEC1C9509D972556D104B70183369" type="personal">
<mods:role id="81DD88E9AA6B2327B8C53D0B0F23DE70">
<mods:roleTerm id="368419C90F1630A13A9D70522097F82B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="65E16C16746692A76FFA20333DE2B3FD">Buczkó, Krisztina</mods:namePart>
</mods:name>
<mods:name id="3DDA1D125BA78E081CC29F1F39F66B91" type="personal">
<mods:role id="4F3373C05305FC3635828CAD9012D29A">
<mods:roleTerm id="21DEA9A3E9C5D3A383536F5F3C6A20A3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2311FF2A2651F79006F69E5698EEF7AA">Vermeulen, Jaap J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="72EF0ECE41FA8F16F39DDC0B25AB3278">text</mods:typeOfResource>
<mods:relatedItem id="46C1701A4F838B73FD17456AD97F71DE" type="host">
<mods:titleInfo id="6C18E22C3AD935FAC0268303B5163457">
<mods:title id="F85DE368418A962E3F557C22D76E7FB6">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="5793D25F9CD740E1C99125FA4063A8D8">
<mods:date id="BC5E72369779A96623476B3FC2E5918A">2020</mods:date>
<mods:detail id="6A8FF2FCB27C5E2675EE92D565F5C910" type="pubDate">
<mods:number id="74CF6CCBCDBBB22F5D17FE7498825810">2020-05-08</mods:number>
</mods:detail>
<mods:detail id="4D1D129781E535856386946429B6A4B9" type="volume">
<mods:number id="F73D400EFB1F065965E6A9B4EC65C86E">68</mods:number>
</mods:detail>
<mods:extent id="C358EBF0AC9A75679C13CC884990E460" unit="page">
<mods:start id="3655F6AC26BDB00F7D1D6068CB4B7EC4">350</mods:start>
<mods:end id="2CBF434B32BA23084D6E98B1D66CB69E">368</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="543786DB17A610AFE9BCFF825EC2284B">journal article</mods:classification>
<mods:identifier id="CEC440E89678664E4CB761CF5ED6B14B" type="DOI">10.26107/RBZ-2020-0052</mods:identifier>
<mods:identifier id="32835DC4BB7233EB6093AB7C416BDDE7" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="1F8E999A1CF04A6DE8E7F899382CAFBC" type="Zenodo-Dep">4576861</mods:identifier>
<mods:identifier id="5511D99380966C30B476E333B0599848" type="ZooBank">C7D4631B-9B02-4E6E-9665-76CCFECA86A0</mods:identifier>
</mods:mods>
<treatment id="9565D05FFFCFFFC5FCDA5934FD789B99" LSID="urn:lsid:plazi:treatment:9565D05FFFCFFFC5FCDA5934FD789B99" httpUri="http://treatment.plazi.org/id/9565D05FFFCFFFC5FCDA5934FD789B99" lastPageId="8" lastPageNumber="358" pageId="2" pageNumber="352">
<subSubSection id="55D632C2FFCFFFCFFCDA5934FB489839" pageId="2" pageNumber="352" type="nomenclature">
<paragraph id="1D736149FFCFFFCFFCDA5934FB489839" blockId="2.[832,1432,796,885]" pageId="2" pageNumber="352">
<heading id="463BD625FFCFFFCFFCDA5934FB489839" bold="true" centered="true" fontSize="10" level="2" pageId="2" pageNumber="352" reason="2">
<emphasis id="2FB8BD5BFFCFFFCFFCDA5934FB489839" bold="true" pageId="2" pageNumber="352">
<taxonomicName id="DACC1ACAFFCFFFCFFCDA5934FA839859" ID-CoL="7QQPY" authority="Pall-Gergely &amp; Vermeulen" authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2020" box="[832,1373,796,821]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="352" phylum="Mollusca" rank="species" species="incurvus" status="sp. nov.">
<emphasis id="2FB8BD5BFFCFFFCFFCDA5934FBC29859" bold="true" box="[832,1052,797,821]" italics="true" pageId="2" pageNumber="352">Clostophis incurvus</emphasis>
Páll-Gergely &amp; Vermeulen
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCFFFCFFAF05934FB489839" pageId="2" pageNumber="352" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFCFFFCFFBD95974FB4B9819" box="[1091,1173,861,885]" pageId="2" pageNumber="352" type="description">
<paragraph id="1D736149FFCFFFCFFBD95974FB4B9819" blockId="2.[832,1432,796,885]" box="[1091,1173,861,885]" pageId="2" pageNumber="352">
(
<figureCitation id="85F77DCCFFCFFFCFFBD15974FB539819" box="[1099,1165,861,885]" captionStart="Fig" captionStartId="6.[113,148,2034,2056]" captionTargetBox="[149,1415,189,2019]" captionTargetId="figure-11@6.[149,1415,189,2019]" captionTargetPageId="6" captionText="Fig. 4. Clostophis incurvus Páll-Gergely &amp; Vermeulen, new species (NHMUK 20191125, holotype). A, apertural view; B, dorsal view; C, ventral view; D, lateral view; E, protoconch sculpture; F, aperture dentition; G, sculpture of the ventral side; H, sculpture of body whorl (lateral side is shown since the part from standard apertural view has been broken off)." figureDoi="http://doi.org/10.5281/zenodo.4576869" httpUri="https://zenodo.org/record/4576869/files/figure.png" pageId="2" pageNumber="352">Fig. 4</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFCFFFCFFCB759B4FB769F39" pageId="2" pageNumber="352" type="materials_examined">
<paragraph id="1D736149FFCFFFCFFCB759B4FB769F39" blockId="2.[812,1452,925,1110]" pageId="2" pageNumber="352">
<emphasis id="2FB8BD5BFFCFFFCFFCB759B4FC0498D9" bold="true" box="[813,986,925,949]" pageId="2" pageNumber="352">Type material.</emphasis>
<materialsCitation id="ADA46B14FFCFFFCFFC7E59B4FB769F39" collectingDate="1998-10-02" collectionCode="NHMUK, JJV" collectorName="Vermeulen, J. J. &amp; Whitten, A. J." country="Vietnam" county="Halong Bay area" latitude="20.847218" location="Song Sot Cave" longLatPrecision="1" longitude="107.09445" municipality="Dao Bo Hon" pageId="2" pageNumber="352" specimenCode="NHMUK 20191125, JJV 6263" specimenCount="1" stateProvince="Quang Ninh" typeStatus="holotype">
<typeStatus id="C277DFEBFFCFFFCFFC7E59B4FB9798D9" box="[996,1097,925,949]" pageId="2" pageNumber="352" type="holotype">Holotype</typeStatus>
(SH:
<quantity id="DA34CCACFFCFFFCFFB0A59B4FB2998D9" box="[1168,1271,925,949]" metricMagnitude="-3" metricUnit="m" metricValue="1.25" pageId="2" pageNumber="352" unit="mm" value="1.25">1.25 mm</quantity>
, SW:
<quantity id="DA34CCACFFCFFFCFFAA559B4FA7A98D9" box="[1343,1444,925,949]" metricMagnitude="-3" metricUnit="m" metricValue="1.08" pageId="2" pageNumber="352" unit="mm" value="1.08">1.08 mm</quantity>
) (
<specimenCode id="4D6AC932FFCFFFCFFCAE5994FBC398B9" box="[820,1053,957,981]" collectionCode="NHMUK" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="2" pageNumber="352" type="Museum">NHMUK 20191125</specimenCode>
; original inv. number:
<specimenCode id="4D6AC932FFCFFFCFFAB45997FA4098B9" box="[1326,1438,957,981]" collectionCode="JJV" pageId="2" pageNumber="352">JJV 6263</specimenCode>
),
<collectingCountry id="65DB21D9FFCFFFCFFCB759F4FC529899" box="[813,908,989,1013]" name="Vietnam" pageId="2" pageNumber="352">Vietnam</collectingCountry>
,
<collectingRegion id="DF08AFABFFCFFFCFFC0D59F4FB589899" box="[919,1158,989,1013]" country="Vietnam" name="Quang Ninh" pageId="2" pageNumber="352">Quang Ninh Province</collectingRegion>
,
<collectingCounty id="F41219C5FFCFFFCFFB0859F4FA959899" box="[1170,1355,989,1014]" pageId="2" pageNumber="352">Halong Bay area</collectingCounty>
,
<collectingMunicipality id="FD17FB33FFCFFFCFFAC259F7FC809F79" pageId="2" pageNumber="352">Dao Bo Hon</collectingMunicipality>
,
<location id="18133792FFCFFFCFFCFD59D4FBDF9F79" LSID="urn:lsid:plazi:treatment:9565D05FFFCFFFC5FCDA5934FD789B99:18133792FFCFFFCFFCFD59D4FBDF9F79" box="[871,1025,1021,1045]" country="Vietnam" county="Halong Bay area" latitude="20.847218" longLatPrecision="1" longitude="107.09445" municipality="Dao Bo Hon" name="Song Sot Cave" pageId="2" pageNumber="352" stateProvince="Quang Ninh">Song Sot Cave</location>
,
<geoCoordinate id="78F8078EFFCFFFCFFB9059D5FB4F9F79" box="[1034,1169,1020,1045]" degrees="20" direction="north" minutes="50.833" orientation="latitude" pageId="2" pageNumber="352" precision="1" value="20.847218">20°50.833N</geoCoordinate>
,
<geoCoordinate id="78F8078EFFCFFFCFFB0359D5FAC39F79" box="[1177,1309,1020,1045]" degrees="107" direction="east" minutes="5.667" orientation="longitude" pageId="2" pageNumber="352" precision="1" value="107.09445">107°5.667E</geoCoordinate>
, drift material washed together over sinkhole in cave, coll.
<collectorName id="B039049FFFCFFFCFFA9F5E34FA759F59" box="[1285,1451,1053,1077]" pageId="2" pageNumber="352">Vermeulen, J.J.</collectorName>
&amp;
<collectorName id="B039049FFFCFFFCFFCD05E14FC029F3A" box="[842,988,1085,1110]" pageId="2" pageNumber="352">Whitten, A.J.</collectorName>
,
<date id="69724789FFCFFFCFFC735E14FB7A9F39" box="[1001,1188,1085,1109]" pageId="2" pageNumber="352" value="1998-10-02">
<collectingDate id="7936BE61FFCFFFCFFC735E14FB7A9F39" box="[1001,1188,1085,1109]" pageId="2" pageNumber="352" value="1998-10-02">02 October 1998</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFCFFFCFFCB75E54FC659FB9" pageId="2" pageNumber="352" type="diagnosis">
<paragraph id="1D736149FFCFFFCFFCB75E54FC659FB9" blockId="2.[813,1451,1149,1237]" pageId="2" pageNumber="352">
<emphasis id="2FB8BD5BFFCFFFCFFCB75E54FC789FF9" bold="true" box="[813,934,1149,1173]" pageId="2" pageNumber="352">Diagnosis.</emphasis>
A convexly conical
<taxonomicName id="DACC1ACAFFCFFFCFFB065E54FACD9FF9" authorityName="Benson" authorityYear="1860" box="[1180,1299,1149,1173]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="352" phylum="Mollusca" rank="genus">
<emphasis id="2FB8BD5BFFCFFFCFFB065E54FACD9FF9" box="[1180,1299,1149,1173]" italics="true" pageId="2" pageNumber="352">Clostophis</emphasis>
</taxonomicName>
species with strong spiral striae, a well-developed parietal and a blunt palatal tooth.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFCFFFCFFCB75ED4FA549DF9" pageId="2" pageNumber="352" type="description">
<paragraph id="1D736149FFCFFFCFFCB75ED4FA549DF9" blockId="2.[811,1453,1277,1686]" pageId="2" pageNumber="352">
<emphasis id="2FB8BD5BFFCFFFCFFCB75ED4FC649E79" bold="true" box="[813,954,1277,1301]" pageId="2" pageNumber="352">Description.</emphasis>
Shell triangular, convex, widest at its base. Whorls regularly increasing, bluntly shouldered. Protoconch consisting of ca. 1.5 whorls, no notable sculpture visible. Entire shell consisting of nearly five whorls, with fine, irregular radial growth lines and stronger, dense, equidistant spiral striae. Body whorl with ca. 17 spiral striae from apertural view. Umbilicus relatively narrow, occupies less than one third of the width. Aperture slightly oblique to shell axis, subcircular with blunt sinulus area. Parietal callus rather sharp, not smeared to penultimate whorl. Peristome relatively strongly thickened and expanded, not reflected. Parietal tooth moderately elevated, situated close to parietal callus. Parietal wall with a low, blunt, but strong tooth.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFCFFFCFFCB75C94FA9F9C39" pageId="2" pageNumber="352" type="diagnosis">
<paragraph id="1D736149FFCFFFCFFCB75C94FA9F9C39" blockId="2.[813,1452,1725,1877]" pageId="2" pageNumber="352">
<emphasis id="2FB8BD5BFFCFFFCFFCB75C94FBE69DB9" bold="true" box="[813,1080,1725,1749]" pageId="2" pageNumber="352">Differential diagnosis.</emphasis>
<taxonomicName id="DACC1ACAFFCFFFCFFBD95C94FAD79DB9" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="Pall-Gergely &amp; Jochum" baseAuthorityYear="2017" box="[1091,1289,1725,1749]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="352" phylum="Mollusca" rank="species" species="stochi">
<emphasis id="2FB8BD5BFFCFFFCFFBD95C94FAD79DB9" box="[1091,1289,1725,1749]" italics="true" pageId="2" pageNumber="352">Clostophis stochi</emphasis>
</taxonomicName>
has a weaker and denser spiral striation, a narrower umbilicus, and more rounded whorls.
<taxonomicName id="DACC1ACAFFCFFFCFFC6D5CD4FADD9C79" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[1015,1283,1789,1813]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="352" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
<emphasis id="2FB8BD5BFFCFFFCFFC6D5CD4FADD9C79" box="[1015,1283,1789,1813]" italics="true" pageId="2" pageNumber="352">Clostophis multiformis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCFFFCFFA8C5CD7FA789C79" box="[1302,1446,1789,1813]" pageId="2" pageNumber="352" rank="species">new species</taxonomicNameLabel>
, possesses no or a weaker palatal tooth and a lower parietal tooth. See also under
<taxonomicName id="DACC1ACAFFCFFFCFFB875D17FB769C39" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[1053,1192,1853,1877]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="352" phylum="Mollusca" rank="species" species="koilobasis" status="sp. nov.">
<emphasis id="2FB8BD5BFFCFFFCFFB875D17FB769C39" box="[1053,1192,1853,1877]" italics="true" pageId="2" pageNumber="352">C. koilobasis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCFFFCFFB225D17FAE39C39" box="[1208,1341,1853,1877]" pageId="2" pageNumber="352" rank="species">new species</taxonomicNameLabel>
.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFCFFFCFFCB75D54FB669CB9" pageId="2" pageNumber="352" type="etymology">
<paragraph id="1D736149FFCFFFCFFCB75D54FB669CB9" blockId="2.[813,1451,1917,2005]" pageId="2" pageNumber="352">
<emphasis id="2FB8BD5BFFCFFFCFFCB75D54FC6D9CF9" bold="true" box="[813,947,1917,1941]" pageId="2" pageNumber="352">Etymology.</emphasis>
The specific epithet means bent, curved in Latin, and refers to the last whorl, which is curved inwards in relation to the penultimate whorl.
</paragraph>
</subSubSection>
<subSubSection id="55D632C2FFCFFFC5FCB75DD4FD789B99" lastPageId="8" lastPageNumber="358" pageId="2" pageNumber="352" type="discussion">
<paragraph id="1D736149FFCFFFCFFCB75DD4FA72935A" blockId="2.[812,1452,2045,2102]" pageId="2" pageNumber="352">
<emphasis id="2FB8BD5BFFCFFFCFFCB75DD4FC459379" bold="true" box="[813,923,2045,2069]" pageId="2" pageNumber="352">Remarks.</emphasis>
Although only a single shell is available, its shell characters clearly indicate that it differs from all congeners.
</paragraph>
<caption id="49B331C1FFCEFFCEFF1D5D57FC7F932C" ID-DOI="http://doi.org/10.5281/zenodo.4576863" ID-Zenodo-Dep="4576863" httpUri="https://zenodo.org/record/4576863/files/figure.png" pageId="3" pageNumber="353" startId="3.[135,170,1918,1940]" targetBox="[190,1421,189,1902]" targetPageId="3">
<paragraph id="1D736149FFCEFFCEFF1D5D57FC7F932C" blockId="3.[135,1475,1918,2113]" pageId="3" pageNumber="353">
Fig. 1. Synoptic view of
<taxonomicName id="DACC1ACAFFCEFFCEFE1F5D57FE329CFF" authorityName="Benson" authorityYear="1860" box="[389,492,1918,1939]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="genus">
<emphasis id="2FB8BD5BFFCEFFCEFE1F5D57FE329CFF" box="[389,492,1918,1939]" italics="true" pageId="3" pageNumber="353">Clostophis</emphasis>
</taxonomicName>
species. A,
<taxonomicName id="DACC1ACAFFCEFFCEFDF05D57FCFC9CFF" authorityName="Benson" authorityYear="1860" box="[618,802,1918,1939]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="sankeyi">
<emphasis id="2FB8BD5BFFCEFFCEFDF05D57FCFC9CFF" box="[618,802,1918,1939]" italics="true" pageId="3" pageNumber="353">Clostophis sankeyi</emphasis>
</taxonomicName>
(UMZCI.103320, holotype, copyright: The Trustees of the Natural History Museum, photo: Harry Taylor); B,
<taxonomicName id="DACC1ACAFFCEFFCEFDA95DB2FD059CDC" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="Pall-Gergely &amp; Jochum" baseAuthorityYear="2017" box="[563,731,1947,1968]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="stochi">
<emphasis id="2FB8BD5BFFCEFFCEFDA95DB2FD059CDC" box="[563,731,1947,1968]" italics="true" pageId="3" pageNumber="353">Clostophis stochi</emphasis>
</taxonomicName>
(MZUF GC/49405, holotype, from the original description: Páll-Gergely &amp; Jochum in
<bibRefCitation id="795D1CB8FFCEFFCEFF695D91FE3C9CA2" author="Pall-Gergely B &amp; Jochum A &amp; Asami T" box="[243,482,1976,1998]" pageId="3" pageNumber="353" pagination="327 - 341" refId="ref8739" refString="Pall-Gergely B, Jochum A &amp; Asami T (2017) Three new species and a new genus of Hypselostomatidae (Gastropoda: Pulmonata) from Cong Troi Cave, Northern Vietnam. Acta Zoologica Academiae Scientiarum Hungaricae, 63 (3): 327 - 341." type="journal article" year="2017">Páll-Gergely et al., 2017</bibRefCitation>
); C,
<taxonomicName id="DACC1ACAFFCEFFCEFD945D91FD0E9CA1" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[526,720,1976,1997]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="infantilis" status="sp. nov.">
<emphasis id="2FB8BD5BFFCEFFCEFD945D91FD0E9CA1" box="[526,720,1976,1997]" italics="true" pageId="3" pageNumber="353">Clostophis infantilis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCEFFCEFD465D91FC919CA2" box="[732,847,1976,1998]" pageId="3" pageNumber="353" rank="species">new species</taxonomicNameLabel>
(MNHN-IM-2014-6401, holotype); D,
<taxonomicName id="DACC1ACAFFCEFFCEFB555D91FA509CA1" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[1231,1422,1976,1997]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="incurvus" status="sp. nov.">
<emphasis id="2FB8BD5BFFCEFFCEFB555D91FA509CA1" box="[1231,1422,1976,1997]" italics="true" pageId="3" pageNumber="353">Clostophis incurvus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCEFFCEFA005D91FF129C86" pageId="3" pageNumber="353" rank="species">new species</taxonomicNameLabel>
(NHMUK 20191125, holotype); E,
<taxonomicName id="DACC1ACAFFCEFFCEFDAC5DFDFCC39C85" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[566,797,2004,2025]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="platytrochus" status="sp. nov.">
<emphasis id="2FB8BD5BFFCEFFCEFDAC5DFDFCC39C85" box="[566,797,2004,2025]" italics="true" pageId="3" pageNumber="353">Clostophis platytrochus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCEFFCEFCB15DFCFC7E9C86" box="[811,928,2004,2026]" pageId="3" pageNumber="353" rank="species">new species</taxonomicNameLabel>
(HNHM 104408, holotype); F,
<taxonomicName id="DACC1ACAFFCEFFCEFB455DFDFA639C85" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[1247,1469,2004,2025]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
<emphasis id="2FB8BD5BFFCEFFCEFB455DFDFA639C85" box="[1247,1469,2004,2025]" italics="true" pageId="3" pageNumber="353">Clostophis multiformis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCEFFCEFF1D5DDBFF22936B" box="[135,252,2033,2055]" pageId="3" pageNumber="353" rank="species">new species</taxonomicNameLabel>
(NHMW 113273, holotype); G,
<taxonomicName id="DACC1ACAFFCEFFCEFDDE5DD8FD24936A" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[580,762,2033,2054]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="obtusus" status="sp. nov.">
<emphasis id="2FB8BD5BFFCEFFCEFDDE5DD8FD24936A" box="[580,762,2033,2054]" italics="true" pageId="3" pageNumber="353">Clostophis obtusus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCEFFCEFC9D5DDBFCA2936B" box="[775,892,2033,2055]" pageId="3" pageNumber="353" rank="species">new species</taxonomicNameLabel>
(HNHM 104435, holotype); H,
<taxonomicName id="DACC1ACAFFCEFFCEFB255DD8FA53936A" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[1215,1421,2033,2054]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="koilobasis" status="sp. nov.">
<emphasis id="2FB8BD5BFFCEFFCEFB255DD8FA53936A" box="[1215,1421,2033,2054]" italics="true" pageId="3" pageNumber="353">Clostophis koilobasis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCEFFCEFA005DDBFF109348" pageId="3" pageNumber="353" rank="species">new species</taxonomicNameLabel>
(NHMUK 20191126, holotype); I,
<taxonomicName id="DACC1ACAFFCEFFCEFDA85227FD23934F" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1961" box="[562,765,2062,2083]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="neglectus">
<emphasis id="2FB8BD5BFFCEFFCEFDA85227FD23934F" box="[562,765,2062,2083]" italics="true" pageId="3" pageNumber="353">Clostophis neglectus</emphasis>
</taxonomicName>
(drawing from the original description: van
<bibRefCitation id="795D1CB8FFCEFFCEFB265227FA419348" author="Benthem Jutting WSS van" box="[1212,1439,2062,2084]" pageId="3" pageNumber="353" pagination="34 - 48" refId="ref8179" refString="Benthem Jutting WSS van (1961) Additional new Species and new Localities of the Family Vertiginidae and the Genera Oophana and Opisthostoma from Malaya. Bulletin of the Raffles Museum, 26: 34 - 48. http: // lkcnhm. nus. edu. sg / rbz / biblio / 26 / 26 brm 034 - 048. pdf" type="journal article" year="1961">Benthem Jutting, 1961</bibRefCitation>
); J,
<taxonomicName id="DACC1ACAFFCEFFCEFF1D5202FEA8932C" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="Panha &amp; Burch" baseAuthorityYear="1999" box="[135,374,2091,2112]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="353" phylum="Mollusca" rank="species" species="proboscideus">
<emphasis id="2FB8BD5BFFCEFFCEFF1D5202FEA8932C" box="[135,374,2091,2112]" italics="true" pageId="3" pageNumber="353">Clostophis proboscideus</emphasis>
</taxonomicName>
(holotype from the original description). Scale = 1 mm.
</paragraph>
</caption>
<caption id="49B331C1FFC9FFC9FFEB5CDBFDED9C17" ID-DOI="http://doi.org/10.5281/zenodo.4576865" ID-Zenodo-Dep="4576865" httpUri="https://zenodo.org/record/4576865/files/figure.png" pageId="4" pageNumber="354" startId="4.[113,149,1778,1800]" targetBox="[113,1451,387,1763]" targetPageId="4">
<paragraph id="1D736149FFC9FFC9FFEB5CDBFDED9C17" blockId="4.[113,1453,1778,1915]" pageId="4" pageNumber="354">
Fig. 2. Known distribution of
<taxonomicName id="DACC1ACAFFC9FFC9FE3F5CDBFDD39C6B" authorityName="Benson" authorityYear="1860" box="[421,525,1778,1799]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="genus">
<emphasis id="2FB8BD5BFFC9FFC9FE3F5CDBFDD39C6B" box="[421,525,1778,1799]" italics="true" pageId="4" pageNumber="354">Clostophis</emphasis>
</taxonomicName>
species. Empty triangle:
<taxonomicName id="DACC1ACAFFC9FFC9FC885CDBFC639C6B" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="Pall-Gergely &amp; Jochum" baseAuthorityYear="2017" box="[786,957,1778,1799]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="stochi">
<emphasis id="2FB8BD5BFFC9FFC9FC885CDBFC639C6B" box="[786,957,1778,1799]" italics="true" pageId="4" pageNumber="354">Clostophis stochi</emphasis>
</taxonomicName>
; inverted empty triangle:
<taxonomicName id="DACC1ACAFFC9FFC9FB525CDBFAB49C6B" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="Panha &amp; Burch" baseAuthorityYear="1999" box="[1224,1386,1778,1799]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="proboscideus">
<emphasis id="2FB8BD5BFFC9FFC9FB525CDBFAB49C6B" box="[1224,1386,1778,1799]" italics="true" pageId="4" pageNumber="354">C. proboscideus</emphasis>
</taxonomicName>
; filled triangle:
<taxonomicName id="DACC1ACAFFC9FFC9FF5D5D26FEA59C48" authorityName="Benson" authorityYear="1860" box="[199,379,1807,1828]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="sankeyi">
<emphasis id="2FB8BD5BFFC9FFC9FF5D5D26FEA59C48" box="[199,379,1807,1828]" italics="true" pageId="4" pageNumber="354">Clostophis sankeyi</emphasis>
</taxonomicName>
; inverted filled triangle:
<taxonomicName id="DACC1ACAFFC9FFC9FDFD5D26FCEA9C48" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[615,820,1807,1828]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="koilobasis" status="sp. nov.">
<emphasis id="2FB8BD5BFFC9FFC9FDFD5D26FCEA9C48" box="[615,820,1807,1828]" italics="true" pageId="4" pageNumber="354">Clostophis koilobasis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC9FFC9FCDA5D26FC689C49" box="[832,950,1807,1829]" pageId="4" pageNumber="354" rank="species">new species</taxonomicNameLabel>
, and
<taxonomicName id="DACC1ACAFFC9FFC9FC7C5D26FB7B9C48" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[998,1189,1807,1828]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="incurvus" status="sp. nov.">
<emphasis id="2FB8BD5BFFC9FFC9FC7C5D26FB7B9C48" box="[998,1189,1807,1828]" italics="true" pageId="4" pageNumber="354">Clostophis incurvus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC9FFC9FB2B5D26FAF89C49" box="[1201,1318,1807,1829]" pageId="4" pageNumber="354" rank="species">new species</taxonomicNameLabel>
; filled square:
<taxonomicName id="DACC1ACAFFC9FFC9FFEB5D05FEE29C2D" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="van Benthem Jutting" baseAuthorityYear="1961" box="[113,316,1836,1857]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="neglectus">
<emphasis id="2FB8BD5BFFC9FFC9FFEB5D05FEE29C2D" box="[113,316,1836,1857]" italics="true" pageId="4" pageNumber="354">Clostophis neglectus</emphasis>
</taxonomicName>
; filled diamond:
<taxonomicName id="DACC1ACAFFC9FFC9FE7D5D05FD109C2D" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[487,718,1836,1857]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="platytrochus" status="sp. nov.">
<emphasis id="2FB8BD5BFFC9FFC9FE7D5D05FD109C2D" box="[487,718,1836,1857]" italics="true" pageId="4" pageNumber="354">Clostophis platytrochus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC9FFC9FD475D05FC8B9C2E" box="[733,853,1836,1858]" pageId="4" pageNumber="354" rank="species">new species</taxonomicNameLabel>
. Inset map shows the distribution of
<taxonomicName id="DACC1ACAFFC9FFC9FB525D05FA789C2D" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[1224,1446,1836,1857]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="multiformis" status="sp. nov.">
<emphasis id="2FB8BD5BFFC9FFC9FB525D05FA789C2D" box="[1224,1446,1836,1857]" italics="true" pageId="4" pageNumber="354">Clostophis multiformis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC9FFC9FFEB5D60FF3B9C32" box="[113,229,1864,1886]" pageId="4" pageNumber="354" rank="species">new species</taxonomicNameLabel>
(circles with labels, see explanation in the text),
<taxonomicName id="DACC1ACAFFC9FFC9FD5C5D61FCA29C31" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[710,892,1864,1885]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="obtusus" status="sp. nov.">
<emphasis id="2FB8BD5BFFC9FFC9FD5C5D61FCA29C31" box="[710,892,1864,1885]" italics="true" pageId="4" pageNumber="354">Clostophis obtusus</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC9FFC9FC135D60FC209C32" box="[905,1022,1864,1886]" pageId="4" pageNumber="354" rank="species">new species</taxonomicNameLabel>
(empty diamond), and
<taxonomicName id="DACC1ACAFFC9FFC9FB7E5D61FA7B9C31" authority="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen, 2020" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" box="[1252,1445,1864,1885]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="354" phylum="Mollusca" rank="species" species="infantilis" status="sp. nov.">
<emphasis id="2FB8BD5BFFC9FFC9FB7E5D61FA7B9C31" box="[1252,1445,1864,1885]" italics="true" pageId="4" pageNumber="354">Clostophis infantilis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFC9FFC9FFEB5D4FFF389C17" box="[113,230,1893,1915]" pageId="4" pageNumber="354" rank="species">new species</taxonomicNameLabel>
(empty square), east of Thakhek.
</paragraph>
</caption>
<caption id="49B331C1FFC8FFC8FF125D07FB5E9C0D" ID-DOI="http://doi.org/10.5281/zenodo.4576867" ID-Zenodo-Dep="4576867" httpUri="https://zenodo.org/record/4576867/files/figure.png" pageId="5" pageNumber="355" startId="5.[136,171,1838,1860]" targetBox="[136,1474,413,1823]" targetPageId="5">
<paragraph id="1D736149FFC8FFC8FF125D07FB5E9C0D" blockId="5.[136,1474,1838,1889]" pageId="5" pageNumber="355">
Fig. 3. Shells of
<taxonomicName id="DACC1ACAFFC8FFC8FEAA5D07FE499C2F" authorityName="Benson" authorityYear="1860" box="[304,407,1838,1859]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="355" phylum="Mollusca" rank="genus">
<emphasis id="2FB8BD5BFFC8FFC8FEAA5D07FE499C2F" box="[304,407,1838,1859]" italics="true" pageId="5" pageNumber="355">Clostophis</emphasis>
</taxonomicName>
species. A, holotype of
<taxonomicName id="DACC1ACAFFC8FFC8FD105D07FC9E9C2F" authorityName="Benson" authorityYear="1860" box="[650,832,1838,1859]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="355" phylum="Mollusca" rank="species" species="sankeyi">
<emphasis id="2FB8BD5BFFC8FFC8FD105D07FC9E9C2F" box="[650,832,1838,1859]" italics="true" pageId="5" pageNumber="355">Clostophis sankeyi</emphasis>
</taxonomicName>
(UMZCI.103320, copyright: The Trustees of the Natural History Museum, photo: Harry Taylor); B, holotype of
<taxonomicName id="DACC1ACAFFC8FFC8FDC65D62FC959C0C" authorityName="Páll-Gergely &amp; Hunyadi &amp; Grego &amp; Reischütz &amp; Buczkó &amp; Vermeulen" authorityYear="2020" baseAuthorityName="Panha &amp; Burch" baseAuthorityYear="1999" box="[604,843,1867,1888]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="5" pageNumber="355" phylum="Mollusca" rank="species" species="proboscideus">
<emphasis id="2FB8BD5BFFC8FFC8FDC65D62FC959C0C" box="[604,843,1867,1888]" italics="true" pageId="5" pageNumber="355">Clostophis proboscideus</emphasis>
</taxonomicName>
(from the original description).
</paragraph>
</caption>
<caption id="49B331C1FFCBFFCBFFEB5DDBFC15932D" ID-DOI="http://doi.org/10.5281/zenodo.4576869" ID-Zenodo-Dep="4576869" httpUri="https://zenodo.org/record/4576869/files/figure.png" pageId="6" pageNumber="356" startId="6.[113,148,2034,2056]" targetBox="[149,1415,189,2019]" targetPageId="6">
<paragraph id="1D736149FFCBFFCBFFEB5DDBFC15932D" blockId="6.[113,1452,2034,2113]" pageId="6" pageNumber="356">
Fig. 4.
<taxonomicName id="DACC1ACAFFCBFFCBFF2C5DDBFD5D9364" authority="Pall-Gergely &amp; Vermeulen" authorityName="Pall-Gergely &amp; Vermeulen" authorityYear="2020" box="[182,643,2034,2056]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="6" pageNumber="356" phylum="Mollusca" rank="species" species="incurvus" status="sp. nov.">
<emphasis id="2FB8BD5BFFCBFFCBFF2C5DDBFEA8936B" box="[182,374,2034,2055]" italics="true" pageId="6" pageNumber="356">Clostophis incurvus</emphasis>
Páll-Gergely &amp; Vermeulen
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCBFFCBFD175DDBFCDF9364" box="[653,769,2034,2056]" pageId="6" pageNumber="356" rank="species">new species</taxonomicNameLabel>
(NHMUK 20191125, holotype). A, apertural view; B, dorsal view; C, ventral view; D, lateral view; E, protoconch sculpture; F, aperture dentition; G, sculpture of the ventral side; H, sculpture of body whorl (lateral side is shown since the part from standard apertural view has been broken off).
</paragraph>
</caption>
<caption id="49B331C1FFCAFFCAFF125239FA1C932F" ID-DOI="http://doi.org/10.5281/zenodo.4576872" ID-Zenodo-Dep="4576872" httpUri="https://zenodo.org/record/4576872/files/figure.png" pageId="7" pageNumber="357" startId="7.[136,171,2064,2086]" targetBox="[162,1448,189,2048]" targetPageId="7">
<paragraph id="1D736149FFCAFFCAFF125239FA1C932F" blockId="7.[136,1474,2064,2115]" pageId="7" pageNumber="357">
Fig. 5.
<taxonomicName id="DACC1ACAFFCAFFCAFF4B5239FDC5934A" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2020" box="[209,539,2064,2086]" class="Gastropoda" family="Gastrocoptidae" genus="Clostophis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="357" phylum="Mollusca" rank="species" species="infantilis" status="sp. nov.">
<emphasis id="2FB8BD5BFFCAFFCAFF4B5239FE4B9349" box="[209,405,2064,2085]" italics="true" pageId="7" pageNumber="357">Clostophis infantilis</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="348B0020FFCAFFCAFDBD5239FD43934A" box="[551,669,2064,2086]" pageId="7" pageNumber="357" rank="species">new species</taxonomicNameLabel>
(MNHN-IM-2014-6401, holotype). A, dorsal view; B, ventral view; C, apertural view; D, lateral view; E, protoconch sculpture; F, sculpture of the ventral side; G, sculpture of body whorl from standard apertural view
</paragraph>
</caption>
<paragraph id="1D736149FFC5FFC5FFEB5A97FD789B99" blockId="8.[113,752,189,245]" pageId="8" pageNumber="358">However, future efforts should focus on collecting more shells in order to reveal the intraspecific variability.</paragraph>
</subSubSection>
</treatment>
</document>