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<document id="B90229EED7F87D20A9D154123557BFF7" ID-DOI="10.11646/palaeoentomology.6.3.9" ID-ISSN="2624-2834" ID-Zenodo-Dep="8073120" ID-ZooBank="02D62C96-11B3-4ECF-B08D-C239EB442981" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1687503520151" checkinUser="plazi" docAuthor="JATTIOT, ROMAIN, TRINCAL, VINCENT, GARROUSTE, ROMAIN &amp; NEL, ANDRÉ" docDate="2023" docId="7B6E7700A401FFB9C48BFE5BFAEA3D35" docLanguage="en" docName="palaeoentomology.6.3.9.pdf" docOrigin="Palaeoentomology 6 (3)" docSource="http://dx.doi.org/10.11646/palaeoentomology.6.3.9" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.4:Palaeoentomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeoentomology.2018-.journal_article" docStyleVersion="4" docTitle="Ilyocoris andancensis JATTIOT &amp; TRINCAL &amp; GARROUSTE &amp; NEL 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="261" masterDocId="87570F78A400FFB8C403F861FFF43B16" masterDocTitle="A new fossil species of the saucer bug genus Ilyocoris (Heteroptera, Naucoridae) from the Upper Miocene maar paleolake of la Montagne dAndance (France)" masterLastPageNumber="267" masterPageNumber="260" pageNumber="261" updateTime="1687503915283" updateUser="ExternalLinkService">
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<mods:title id="16ACBBEEB964D8E6B35A7186402E3EDA">A new fossil species of the saucer bug genus Ilyocoris (Heteroptera, Naucoridae) from the Upper Miocene maar paleolake of la Montagne dAndance (France)</mods:title>
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<mods:namePart id="C4A592BB4120DEEDAAFF1B21175242F6">JATTIOT, ROMAIN</mods:namePart>
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<mods:namePart id="7EB93EB4B9E22B18C169C8931D7DE5D6">TRINCAL, VINCENT</mods:namePart>
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<mods:namePart id="7D803B15BEE2E482C4F5E6EFE97ACF09">GARROUSTE, ROMAIN</mods:namePart>
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<mods:namePart id="F623C685C6FEDFA5B7CB19719CAADC6F">NEL, ANDRÉ</mods:namePart>
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<mods:title id="1E70CABA8422470FC30BB1C396895C8C">Palaeoentomology</mods:title>
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<mods:date id="27A2A0ECA112F6149178C8D45BB72277">2023</mods:date>
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<heading id="A830717AA401FFB9C48BFE5BFE103D43" bold="true" box="[136,484,1594,1621]" fontSize="10" level="2" pageId="1" pageNumber="261" reason="3">
<emphasis id="C1B31A04A401FFB9C48BFE5BFE103D43" bold="true" box="[136,484,1594,1621]" pageId="1" pageNumber="261">
<taxonomicName id="34C7BD95A401FFB9C48BFE5BFE773D43" authority="JATTIOT &amp; TRINCAL &amp; GARROUSTE &amp; NEL, 2023" authorityName="JATTIOT &amp; TRINCAL &amp; GARROUSTE &amp; NEL" authorityYear="2023" box="[136,387,1594,1621]" class="Insecta" family="Naucoridae" genus="Ilyocoris" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="261" phylum="Arthropoda" rank="species" species="andancensis" status="sp. nov.">
<emphasis id="C1B31A04A401FFB9C48BFE5BFE773D43" bold="true" box="[136,387,1594,1621]" italics="true" pageId="1" pageNumber="261">Ilyocoris andancensis</emphasis>
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<taxonomicNameLabel id="DA80A77FA401FFB9C589FE5AFE103D43" box="[394,484,1595,1621]" pageId="1" pageNumber="261" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="F378C616A401FFB9C48BFE3EFEA63DD4" blockId="1.[136,777,1594,1730]" pageId="1" pageNumber="261">
(
<figureCitation id="6BFCDA93A401FFB9C493FE3EFF073D6F" box="[144,243,1631,1657]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="2.[136,235,1951,1975]" captionStartId-1="3.[136,235,1482,1506]" captionStartId-2="4.[136,235,1482,1506]" captionStartId-3="5.[136,235,1114,1138]" captionTargetBox-0="[263,1324,194,1923]" captionTargetBox-1="[280,1308,191,1459]" captionTargetBox-2="[280,1308,190,1459]" captionTargetBox-3="[196,1391,166,1090]" captionTargetId-0="figure-19@2.[258,1329,189,1927]" captionTargetId-1="figure-241@3.[280,1308,189,1459]" captionTargetId-2="figure-225@4.[280,1308,189,1459]" captionTargetId-3="figure-437@5.[196,1391,166,1090]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="FIGURE 1. Ilyocoris andancensis sp. nov., holotype MNHN-F.A71364. Photographs of general habitus. A, Dry. B, Under alcohol. Scale bars = 2 mm." captionText-1="FIGURE 2. Ilyocoris andancensis sp. nov., holotype MNHN-F.A71364. Photographs of apex of abdomen. A, Under normal light, dry. B, Under UV light showing ovipositor, dry. Scale bar = 2 mm." captionText-2="FIGURE 3. Ilyocoris andancensis sp. nov., paratype MNHN-F.A71365. Photographs. A, Habitus, dry. B, Head and right fore leg, dry. Scale bars = 2 mm." captionText-3="FIGURE 4. Ilyocoris andancensis sp. nov., paratype MNHN-F.A71366. Photograph of habitus, dry. Scale bar = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8073124" figureDoi-1="http://doi.org/10.5281/zenodo.8073126" figureDoi-2="http://doi.org/10.5281/zenodo.8073128" figureDoi-3="http://doi.org/10.5281/zenodo.8073130" httpUri-0="https://zenodo.org/record/8073124/files/figure.png" httpUri-1="https://zenodo.org/record/8073126/files/figure.png" httpUri-2="https://zenodo.org/record/8073128/files/figure.png" httpUri-3="https://zenodo.org/record/8073130/files/figure.png" pageId="1" pageNumber="261">Figs 14</figureCitation>
)
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urn:lsid:zoobank.org:act:
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<subSubSection id="BBDD959DA401FFB9C48BFE91FD153C6E" pageId="1" pageNumber="261" type="materials_examined">
<paragraph id="F378C616A401FFB9C48BFE91FD153C6E" blockId="1.[136,778,1776,2093]" pageId="1" pageNumber="261">
<emphasis id="C1B31A04A401FFB9C48BFE91FF033C1C" bold="true" box="[136,247,1776,1802]" pageId="1" pageNumber="261">Material.</emphasis>
<materialsCitation id="43AFCC4BA401FFB9C506FE91FD153C6E" collectorName="Romain Jattiot" country="France" location="Museum National d'Histoire Naturelle" municipality="Paris" pageId="1" pageNumber="261" specimenCount="2" typeStatus="holotype">
<typeStatus id="2C7C78B4A401FFB9C506FE91FE993C1C" box="[261,365,1776,1802]" pageId="1" pageNumber="261">Holotype</typeStatus>
MNHN-F.A71364 (
<collectorName id="5E32A3C0A401FFB9C65EFE91FCFD3C1C" box="[605,777,1776,1802]" pageId="1" pageNumber="261">Romain Jattiot</collectorName>
coll.),
<typeStatus id="2C7C78B4A401FFB9C4EBFF74FEA73C39" box="[232,339,1813,1839]" pageId="1" pageNumber="261">paratypes</typeStatus>
MNHN-F.A71365, and MNHNF.A71366 (all compressed mummies), stored at the
<location id="F61890CDA401FFB9C48BFF3CFDC83C61" LSID="urn:lsid:plazi:treatment:7B6E7700A401FFB9C48BFE5BFAEA3D35:F61890CDA401FFB9C48BFF3CFDC83C61" box="[136,572,1885,1911]" country="France" municipality="Paris" name="Museum National d'Histoire Naturelle" pageId="1" pageNumber="261">Muséum National dHistoire Naturelle</location>
,
<collectingMunicipality id="131C5C6CA401FFB9C644FF3CFD763C61" box="[583,642,1885,1911]" pageId="1" pageNumber="261">Paris</collectingMunicipality>
,
<collectingCountry id="8BD08686A401FFB9C68EFF3FFD293C6E" box="[653,733,1886,1912]" name="France" pageId="1" pageNumber="261">France</collectingCountry>
.
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</paragraph>
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<subSubSection id="BBDD959DA401FFB9C4BEFFE0FD16333A" pageId="1" pageNumber="261" type="diagnosis">
<paragraph id="F378C616A401FFB9C4BEFFE0FD16333A" blockId="1.[136,778,1776,2093]" pageId="1" pageNumber="261">
<emphasis id="C1B31A04A401FFB9C4BEFFE0FECC3C8D" bold="true" box="[189,312,1921,1947]" pageId="1" pageNumber="261">Diagnosis.</emphasis>
Adult characters only. Color patterns identical on the three fossils. Connexivum brownish along posterior part and yellowish along anterior part; green-yellow anterior part of pronotum clearly narrower than black posterior part; outer margin of embolium black.
</paragraph>
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<subSubSection id="BBDD959DA401FFB9C75CF8FFFBB73A31" pageId="1" pageNumber="261" type="materials_examined">
<paragraph id="F378C616A401FFB9C75CF8FFFBB73A31" blockId="1.[810,1452,158,1572]" pageId="1" pageNumber="261">
<materialsCitation id="43AFCC4BA401FFB9C75CF8FFFBCB3A31" collectorName="La Montagne d'Andance'" country="France" location="Saint-Bauzile village" pageId="1" pageNumber="261" specimenCount="1" typeStatus="holotype">
<emphasis id="C1B31A04A401FFB9C75CF8FFFB6C3BAE" bold="true" box="[863,1176,158,184]" pageId="1" pageNumber="261">
<typeStatus id="2C7C78B4A401FFB9C75CF8FFFC6E3BAE" box="[863,922,158,184]" pageId="1" pageNumber="261">Type</typeStatus>
locality and horizon.
</emphasis>
Lower limit of Turolian to Tortonian-Messinian boundary, 7.30 ± 0.15 Ma (
<bibRefCitation id="9756BBE7A401FFB9C166F8A2FC413A14" author="Pastre, J. - F. &amp; Singer, B. S. &amp; Guillou, H. &amp; Pupin, J. - P. &amp; Riou, B." pageId="1" pageNumber="261" pagination="3 - 10" refId="ref4013" refString="Pastre, J. - F., Singer, B. S., Guillou, H., Pupin, J. - P. &amp; Riou, B. (2004) Chronostratigraphy of the key Upper Miocene (Lower Turolian) sequence of la Montagne d'Andance (Ardeche, France). Implications of new 40 Ar / 39 Ar laser fusion and unspiked K-Ar dating of trachytic tephra and basalts. Bulletin de la Societe Geologique de France, 175, 3 - 10. https: // doi. org / 10.2113 / 175.1.3" type="journal article" year="2004">
Pastre
<emphasis id="C1B31A04A401FFB9C729F888FCAB3A14" box="[810,863,232,258]" italics="true" pageId="1" pageNumber="261">et al</emphasis>
., 2004
</bibRefCitation>
),
<collectorName id="5E32A3C0A401FFB9C7D2F889FAF73A14" box="[977,1283,232,258]" pageId="1" pageNumber="261">La Montagne dAndance</collectorName>
,
<location id="F61890CDA401FFB9C110F889FC8D3A31" LSID="urn:lsid:plazi:treatment:7B6E7700A401FFB9C48BFE5BFAEA3D35:F61890CDA401FFB9C110F889FC8D3A31" country="France" name="Saint-Bauzile village" pageId="1" pageNumber="261">Saint-Bauzile village</location>
, Ardèche,
<collectingCountry id="8BD08686A401FFB9C7ECF96CFBCB3A31" box="[1007,1087,269,295]" name="France" pageId="1" pageNumber="261">France</collectingCountry>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="BBDD959DA401FFB9C75CF951FAEA3D35" pageId="1" pageNumber="261" type="description">
<paragraph id="F378C616A401FFB9C75CF951FB803FF3" blockId="1.[810,1452,158,1572]" pageId="1" pageNumber="261">
<emphasis id="C1B31A04A401FFB9C75CF951FC1D3A5C" bold="true" box="[863,1001,304,330]" pageId="1" pageNumber="261">Description</emphasis>
(based on all specimens). Head transverse; ocelli absent; eyes sessile; labium short and thick, arising near apex of head, conical, obviously segmented, without transverse sulcation; labium; apex of head without a laminated extension; vertex less than two times and half broader than transversal diameter of compound eye; head, pronotum, and scutellum shiny; ground color black and green-yellow; brown dots on head, some of which form two longitudinal, parallel, median lines, each of them widening posteriorly; anterior margin of pronotum straight behind interocular space; lateral pronotal margins limited inside by a strong sunken line; anterior part of pronotum green-yellow, and posterior part black; scutellum black; no meso- and metasternal midventral carinae (observed on
<typeStatus id="2C7C78B4A401FFB9C0C4FB40FADD382D" box="[1223,1321,801,827]" pageId="1" pageNumber="261">holotype</typeStatus>
by partly removing the mummy); macropterous, hemelytra olive brown, opaque, very finely and densely punctate, clavus and margin paler, embolium with a less dense punctuation than the rest of hemelytra but with outer margin black; membrane without venation; fore legs raptorial, fore femur only moderately incrassate with inner antero-basal angle not produced, forming a continuous curve with trochanter, tarsus without a fringe of setae; hind legs flattened; tibiae with strong brownish spines and long swimming setae. Legs pale yellow; apex of abdomen without respiratory processes; connexivum brownish along posterior part and yellowish along anterior part.
</paragraph>
<paragraph id="F378C616A401FFB9C75CFC8CFAEA3D35" blockId="1.[810,1452,158,1572]" pageId="1" pageNumber="261">
<emphasis id="C1B31A04A401FFB9C75CFC8CFC053E11" bold="true" box="[863,1009,1261,1287]" pageId="1" pageNumber="261">Dimensions.</emphasis>
Body
<quantity id="343F6BF3A401FFB9C049FC8FFB413E1E" box="[1098,1205,1262,1288]" metricMagnitude="-2" metricUnit="m" metricValue="1.47" pageId="1" pageNumber="261" unit="mm" value="14.7">14.7 mm</quantity>
long,
<quantity id="343F6BF3A401FFB9C109FC8FFA933E1E" box="[1290,1383,1262,1288]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="1" pageNumber="261" unit="mm" value="6.4">6.4 mm</quantity>
wide (MNHN-F.A71364), 12.0 mm long, 6.0 mm wide (MNHN-F.A71365),
<quantity id="343F6BF3A401FFB9C025FD55FB6D3E59" box="[1062,1177,1332,1359]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="1" pageNumber="261" unit="mm" value="13.5">13.5 mm</quantity>
long, 6.0 mm wide (MNHN-F.A71366); head
<quantity id="343F6BF3A401FFB9C063FD39FB4F3E65" box="[1120,1211,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="1" pageNumber="261" unit="mm" value="1.3">1.3 mm</quantity>
long,
<quantity id="343F6BF3A401FFB9C10EFD39FA9D3E65" box="[1293,1385,1368,1395]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="1" pageNumber="261" unit="mm" value="3.4">3.4 mm</quantity>
wide (MNHN-F.A71364); pronotum 2.0 mm long,
<quantity id="343F6BF3A401FFB9C14DFD1DFA583E80" box="[1358,1452,1404,1430]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="261" unit="mm" value="4.1">4.1 mm</quantity>
wide (MNHN-F.A71364); abdomen
<quantity id="343F6BF3A401FFB9C0DBFDFEFAC13EAC" box="[1240,1333,1439,1466]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="1" pageNumber="261" unit="mm" value="9.7">9.7 mm</quantity>
long,
<quantity id="343F6BF3A401FFB9C18BFDFEFCA23ECB" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="1" pageNumber="261" unit="mm" value="6.5">6.5 mm</quantity>
wide (MNHN-F.A71364); fore tibia
<quantity id="343F6BF3A401FFB9C0F3FDA2FAB03ECB" box="[1264,1348,1475,1501]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="1" pageNumber="261" unit="mm" value="2.1">2.1 mm</quantity>
long,
<quantity id="343F6BF3A401FFB9C18BFDA2FCA23D17" metricMagnitude="-4" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="261" unit="mm" value="0.9">0.9 mm</quantity>
wide; hind tibia 3.0 mm long, tarsus
<quantity id="343F6BF3A401FFB9C113FD87FA9D3D17" box="[1296,1385,1510,1537]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="261" unit="mm" value="4.1">4.1 mm</quantity>
long, hemelytra
<quantity id="343F6BF3A401FFB9C7A2FE68FC033D32" box="[929,1015,1545,1572]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" pageId="1" pageNumber="261" unit="mm" value="8.6">8.6 mm</quantity>
long (MNHN-F.A71364).
</paragraph>
</subSubSection>
</treatment>
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