251 lines
21 KiB
XML
251 lines
21 KiB
XML
<document ID-DOI="10.3897/zookeys.39.424" ID-GBIF-Dataset="4e1adc72-60c9-4846-af21-ed080ed81401" ID-ISSN="1313–2970" ID-Zenodo-Dep="576637" ID-ZooBank="66F92D10-9BCF-4B7A-AAD9-AF6042E8CEA0" approvalRequired="2" approvalRequired_for_treatments="2" checkinTime="1587975101049" checkinUser="plazi" docAuthor="Ferris, Clifford & Lafontaine, Donald" docDate="2010" docId="0A3ACB5EB87D851F4CC49C29FE8F0697" docLanguage="en" docName="ZK_article_2172.pdf" docOrigin="ZooKeys 39 (39)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6a77953c" docStyleName="zookeys.2008.journal_article" docTitle="Pseudomarimatha Ferris & Lafontaine 2010, gen. n." docType="treatment" docUuid="E5E1E7DD-E742-4003-9B0E-91167C8277C3" docUuidSource="ZooBank" docVersion="1" lastPageNumber="133" masterDocId="F603B326B870850F4C489919FF970720" masterDocTitle="Review of the North American species of Marimatha Walker with descriptions of three new species (Lepidoptera, Noctuidae, Eustrotiinae) and the description of Pseudomarimatha flava (Noctuinae, Elaphriini), a new genus and species confused with Marimatha" masterLastPageNumber="135" masterPageNumber="117" pageNumber="130" updateTime="1669282270975" updateUser="tatiana" zenodo-license-document="CC-BY-3.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Review of the North American species of Marimatha Walker with descriptions of three new species (Lepidoptera, Noctuidae, Eustrotiinae) and the description of Pseudomarimatha flava (Noctuinae, Elaphriini), a new genus and species confused with Marimatha</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Ferris, Clifford</mods:namePart>
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<mods:affiliation>, Laramie, United States of America</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Lafontaine, Donald</mods:namePart>
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<mods:affiliation>Agriculture and Agri-Food Canada, Ottawa, Canada</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2010</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2010-03-18</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>39</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>39</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>117</mods:start>
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<mods:end>135</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.3897/zookeys.39.424</mods:identifier>
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<mods:identifier type="GBIF-Dataset">4e1adc72-60c9-4846-af21-ed080ed81401</mods:identifier>
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<mods:identifier type="ISSN">1313–2970</mods:identifier>
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<mods:identifier type="Zenodo-Dep">576637</mods:identifier>
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<mods:identifier type="ZooBank">66F92D10-9BCF-4B7A-AAD9-AF6042E8CEA0</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788624" ID-GBIF-Taxon="163749567" ID-Zenodo-Dep="3788624" LSID="urn:lsid:zoobank.org:act:E5E1E7DD-E742-4003-9B0E-91167C8277C3" httpUri="http://treatment.plazi.org/id/0A3ACB5EB87D851F4CC49C29FE8F0697" lastPageId="16" lastPageNumber="133" pageId="13" pageNumber="130">
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<subSubSection box="[140,697,1328,1355]" pageId="13" pageNumber="130" type="nomenclature">
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<paragraph blockId="13.[140,971,1328,1390]" box="[140,697,1328,1355]" pageId="13" pageNumber="130">
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<heading box="[140,697,1328,1355]" fontSize="11" level="2" pageId="13" pageNumber="130" reason="3">
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<emphasis bold="true" box="[140,697,1328,1355]" pageId="13" pageNumber="130">
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<taxonomicName authority="Ferris & Lafontaine" authorityName="Ferris & Lafontaine" authorityYear="2010" box="[140,604,1328,1355]" class="Insecta" family="Noctuidae" genus="Pseudomarimatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="genus" status="gen. n.">
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<emphasis bold="true" box="[140,358,1328,1355]" italics="true" pageId="13" pageNumber="130">Pseudomarimatha</emphasis>
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Ferris & Lafontaine
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</taxonomicName>
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,
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<taxonomicNameLabel box="[616,697,1328,1354]" pageId="13" pageNumber="130" rank="genus">gen. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,971,1363,1390]" pageId="13" pageNumber="130" type="reference_group">
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<paragraph blockId="13.[140,971,1328,1390]" box="[140,971,1363,1390]" pageId="13" pageNumber="130">
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<uri box="[140,971,1363,1390]" pageId="13" pageNumber="130">
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urn:lsid:zoobank.org:act:
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<uuid box="[423,971,1363,1390]" pageId="13" pageNumber="130">E5E1E7DD-E742-4003-9B0E-91167C8277C3</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection box="[140,864,1433,1460]" pageId="13" pageNumber="130" type="type_taxon">
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<paragraph blockId="13.[140,1108,1433,1672]" box="[140,864,1433,1460]" pageId="13" pageNumber="130">
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<heading box="[140,864,1433,1460]" fontSize="11" level="2" pageId="13" pageNumber="130" reason="3">
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<emphasis bold="true" box="[140,295,1433,1460]" pageId="13" pageNumber="130">
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<emphasis box="[140,198,1434,1460]" inLineHeading="true" pageId="13" pageNumber="130" reason="1">
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<typeStatus box="[140,198,1434,1460]" pageId="13" pageNumber="130">Type</typeStatus>
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</emphasis>
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species.
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</emphasis>
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<taxonomicName authority="Ferris & Lafontaine" authorityName="Ferris & Lafontaine" authorityYear="2010" box="[302,792,1434,1460]" class="Insecta" family="Noctuidae" genus="Pseudomarimatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="species" species="flava" status="sp. n.">
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<emphasis box="[302,558,1434,1460]" italics="true" pageId="13" pageNumber="130">Pseudomarimatha flava</emphasis>
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Ferris & Lafontaine
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</taxonomicName>
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,
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<taxonomicNameLabel box="[803,864,1434,1460]" pageId="13" pageNumber="130" rank="species">sp. n.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="13" pageNumber="130" type="etymology">
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<paragraph blockId="13.[140,1108,1433,1672]" pageId="13" pageNumber="130">
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<emphasis bold="true" box="[185,320,1469,1495]" pageId="13" pageNumber="130">
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<emphasis box="[185,315,1469,1495]" inLineHeading="true" pageId="13" pageNumber="130" reason="1">Etymology</emphasis>
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.
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</emphasis>
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The generic name
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<taxonomicName authorityName="Ferris & Lafontaine" authorityYear="2010" box="[547,743,1469,1495]" class="Insecta" family="Noctuidae" genus="Pseudomarimatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis box="[547,743,1469,1495]" italics="true" pageId="13" pageNumber="130">Pseudomarimatha</emphasis>
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</taxonomicName>
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is taken from the Latin
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<taxonomicName authorityName="Berge & Vader" authorityYear="2001" box="[1034,1108,1469,1495]" class="Malacostraca" family="Stegocephalidae" genus="Pseudo" kingdom="Animalia" order="Amphipoda" pageId="13" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis box="[1034,1108,1469,1495]" italics="true" pageId="13" pageNumber="130">Pseudo</emphasis>
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</taxonomicName>
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(false) and
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<taxonomicName box="[264,390,1504,1530]" class="Insecta" family="Noctuidae" genus="Marimatha" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis box="[264,390,1504,1530]" italics="true" pageId="13" pageNumber="130">Marimatha</emphasis>
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</taxonomicName>
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(the noctuid genus with which
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<taxonomicName authorityName="Ferris & Lafontaine" authorityYear="2010" box="[750,946,1504,1530]" class="Insecta" family="Noctuidae" genus="Pseudomarimatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="13" pageNumber="130" phylum="Arthropoda" rank="genus">
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<emphasis box="[750,946,1504,1530]" italics="true" pageId="13" pageNumber="130">Pseudomarimatha</emphasis>
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</taxonomicName>
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has been confused and is likely to continue to be confused.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="15" lastPageNumber="132" pageId="13" pageNumber="130" type="diagnosis">
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<paragraph blockId="13.[140,1108,1433,1672]" lastBlockId="15.[140,1108,167,1672]" lastPageId="15" lastPageNumber="132" pageId="13" pageNumber="130">
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<emphasis bold="true" box="[185,312,1574,1600]" pageId="13" pageNumber="130">
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<emphasis box="[185,307,1574,1600]" inLineHeading="true" pageId="13" pageNumber="130" reason="1">Diagnosis</emphasis>
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.
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</emphasis>
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Head with rounded slightly bulging frons; hindwing with vein M
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<subScript attach="left" box="[1099,1108,1593,1607]" fontSize="6" pageId="13" pageNumber="130">2</subScript>
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reduced to fold near middle of cell, veins M2 and CuA1on common stalk; tympanic membrane with central sclerite raised and nodular. Male genitalia: tegumen broad, enlarged into posterior lobe ventrally; valva with triangular sacculus with apex separated from remainder of sacculus by membranous band; valva narrow at apex of sacculus, gradually widening to broadly rounded apex, without corona; clasper on costal margin of valva, about 1/4 × valva length; vesica slightly longer than aedeagus with subbasal diverticulum and large apical cornutus 3/4 × as long as aedeagus. Female genitalia: corpus bursae globular posterior, abruptly tapered anteriorly; posterior 1/3 of ductus bursae heavily sclerotized; apophyses long and narrow with triangular plate in posterior apophysis near middle; anal papillae sclerotized.
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</paragraph>
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<caption pageId="14" pageNumber="131" startId="14.[140,222,1426,1448]" targetBox="[177,1071,250,1384]" targetPageId="14">
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<paragraph blockId="14.[140,1105,1422,1480]" pageId="14" pageNumber="131">
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<emphasis bold="true" box="[140,261,1426,1448]" pageId="14" pageNumber="131">Figures 23</emphasis>
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<emphasis bold="true" box="[261,276,1422,1449]" pageId="14" pageNumber="131">–</emphasis>
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<emphasis bold="true" box="[276,309,1426,1448]" pageId="14" pageNumber="131">26.</emphasis>
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<taxonomicName authorityName="Ferris & Lafontaine" authorityYear="2010" box="[321,536,1426,1447]" class="Insecta" family="Noctuidae" genus="Pseudomarimatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="14" pageNumber="131" phylum="Arthropoda" rank="species" species="flava">
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<emphasis box="[321,536,1426,1447]" italics="true" pageId="14" pageNumber="131">Pseudomarimatha flava</emphasis>
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</taxonomicName>
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adults and genitalia.
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<emphasis bold="true" box="[757,783,1426,1448]" pageId="14" pageNumber="131">23</emphasis>
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♂, Portal, Cochise Co,
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<collectingRegion box="[1033,1105,1426,1448]" country="United States of America" name="Arizona" pageId="14" pageNumber="131">Arizona</collectingRegion>
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<specimenCount box="[140,184,1458,1480]" pageId="14" pageNumber="131" type="female">
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<emphasis bold="true" box="[140,166,1458,1480]" pageId="14" pageNumber="131">24</emphasis>
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♀
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</specimenCount>
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, Onion Saddle, Chiricahua Mts, Cochise Co.,
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<collectingRegion box="[621,693,1458,1480]" country="United States of America" name="Arizona" pageId="14" pageNumber="131">Arizona</collectingRegion>
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<specimenCount box="[705,781,1458,1480]" pageId="14" pageNumber="131" type="male">
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<emphasis bold="true" box="[705,731,1458,1480]" pageId="14" pageNumber="131">25</emphasis>
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male
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</specimenCount>
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genitalia
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<specimenCount box="[871,964,1458,1480]" pageId="14" pageNumber="131" type="female">
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<emphasis bold="true" box="[871,897,1458,1480]" pageId="14" pageNumber="131">26</emphasis>
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female
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</specimenCount>
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genitalia.
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</paragraph>
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</caption>
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<paragraph blockId="15.[140,1108,167,1672]" pageId="15" pageNumber="132">
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<taxonomicName authorityName="Ferris & Lafontaine" authorityYear="2010" box="[185,381,307,333]" class="Insecta" family="Noctuidae" genus="Pseudomarimatha" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="132" phylum="Arthropoda" rank="genus">
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<emphasis box="[185,381,307,333]" italics="true" pageId="15" pageNumber="132">Pseudomarimatha</emphasis>
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</taxonomicName>
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appears to be most closely related to
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<taxonomicName authority="Hubner" authorityName="Hubner" box="[792,983,307,333]" class="Insecta" family="Noctuidae" genus="Elaphria" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="132" phylum="Arthropoda" rank="genus">
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<emphasis box="[792,887,307,333]" italics="true" pageId="15" pageNumber="132">Elaphria</emphasis>
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Hübner
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</taxonomicName>
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, but differs in that the costa of the male valva is continuously sclerotized almost to the apex of the valva; the clasper is a simple tapered sinuate structure arising from a swollen base; the isolated apical part of the sacculus is lightly sclerotized, like the rest of the sacculus. In
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<taxonomicName box="[140,235,448,474]" class="Insecta" family="Noctuidae" genus="Elaphria" kingdom="Animalia" order="Lepidoptera" pageId="15" pageNumber="132" phylum="Arthropoda" rank="genus">
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<emphasis box="[140,235,448,474]" italics="true" pageId="15" pageNumber="132">Elaphria</emphasis>
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</taxonomicName>
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there is a gap in the sclerotization of the costal margin of the valva near the end of the clasper, so in dissection the valva tends to fold back on itself at this point; the base of the clasper is forked with a ventrally directed process; and the isolated apical part of the sacculus is much more heavily sclerotized than the basal part.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="15" pageNumber="132" type="description">
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<paragraph blockId="15.[140,1108,167,1672]" pageId="15" pageNumber="132">
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<emphasis bold="true" box="[185,419,589,615]" pageId="15" pageNumber="132">
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<emphasis box="[185,330,589,615]" inLineHeading="true" pageId="15" pageNumber="132" reason="1">Description</emphasis>
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. Adult:
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</emphasis>
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<emphasis box="[428,487,589,615]" italics="true" pageId="15" pageNumber="132">Head</emphasis>
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<emphasis bold="true" box="[495,511,589,615]" pageId="15" pageNumber="132">–</emphasis>
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Frons rounded, slightly bulging; labial palpus curving upward with third segment slightly porrect, middle segment 3 × as long as basal and apical segments; male antenna filiform, densely setose ventrally, female similar but sparsely setose ventrally; eye rounded; ocellus present; vestiture of head and palpi of broad flat scales with slightly raised tuft of scales projecting forward between bases of antennae.
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<emphasis bold="true" box="[292,401,765,791]" pageId="15" pageNumber="132">Thorax –</emphasis>
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Patagia clothed with broad, yellow scales; tegulae and posterior part of thorax covered with blackish-gray scales with white on sides and tip and amount of dark shading more extensive on anterior and middle part of thorax than farther posteriorly.
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<emphasis box="[359,404,871,896]" italics="true" pageId="15" pageNumber="132">Legs</emphasis>
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: spiniform setae confined to tarsi and setae minute; foretibia slightly shorter (0.8 ×) than femur and tarsus.
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<emphasis box="[652,719,907,932]" italics="true" pageId="15" pageNumber="132">Wings</emphasis>
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: forewing venation typically quadrifine with cubital vein appearing four branched; forewing with obtusely angled tornus and acutely angled apex giving wing an elongated appearance; hindwing with typical trifine venation (i.e., M
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<subScript attach="left" box="[407,416,1030,1044]" fontSize="6" pageId="15" pageNumber="132">2</subScript>
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reduced to fold near middle of cell), but veins M
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<subScript attach="left" box="[979,988,1030,1044]" fontSize="6" pageId="15" pageNumber="132">2</subScript>
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and CuA
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<subScript attach="left" box="[1099,1108,1030,1044]" fontSize="6" pageId="15" pageNumber="132">1</subScript>
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on common stalk (extending 0.10–0.25 × length of M
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<subScript attach="left" box="[758,767,1065,1079]" fontSize="6" pageId="15" pageNumber="132">3</subScript>
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).
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<emphasis box="[791,893,1047,1073]" italics="true" pageId="15" pageNumber="132">Abdomen</emphasis>
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<emphasis bold="true" box="[901,916,1047,1073]" pageId="15" pageNumber="132">–</emphasis>
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basal abdominal brushes and pockets absent.
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<emphasis box="[474,737,1082,1108]" italics="true" pageId="15" pageNumber="132">
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Male genitalia
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<emphasis bold="true" box="[645,660,1082,1108]" italics="true" pageId="15" pageNumber="132">–</emphasis>
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Uncus
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</emphasis>
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: decurved, cylindrical, abruptly tapered to spine-like apex, with brush of long stout setae dorsally toward apex.
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<emphasis italics="true" pageId="15" pageNumber="132">Tegumen</emphasis>
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: broad, enlarged into posterior lobe ventrally, about 2 × as wide ventrally as mesially with tuft of long setae on ventral lobe; vinculum with short centrally angled saccus ventrally.
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<taxonomicName authorityName="Ward" authorityYear="1942" box="[294,351,1223,1248]" genus="Juxta" higherTaxonomySource="GBIF" kingdom="Animalia" pageId="15" pageNumber="132" phylum="Arthropoda" rank="genus">
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<emphasis box="[294,351,1223,1248]" italics="true" pageId="15" pageNumber="132">Juxta</emphasis>
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</taxonomicName>
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: small, heavily sclerotized diamond-shaped plate with deep central notch posteriorly.
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<emphasis box="[350,421,1258,1284]" italics="true" pageId="15" pageNumber="132">Valvae</emphasis>
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: symmetrical, basally broad, narrowed mesially, broadening to broadly rounded apex, without corona; sacculus triangular, widest at base; apex of sacculus separated by narrow membranous band (typical of Elaphrini); clasper near dorsal margin of valva, arising from wider base immediately distal to sacculus, then narrow, curving and finger-like, about 1/4 × valva length; valva distal to sacculus lightly sclerotized except for band along costal margin.
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<emphasis box="[673,771,1434,1460]" italics="true" pageId="15" pageNumber="132">Aedeagus</emphasis>
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: smoothly sclerotized; everted vesica tubular, slightly longer than vesica with large subbasal diverticulum and large apical cornutus about 3/4 × as long as aedeagus.
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<emphasis box="[678,855,1504,1530]" italics="true" pageId="15" pageNumber="132">Female genitalia</emphasis>
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<emphasis bold="true" box="[860,874,1504,1530]" pageId="15" pageNumber="132">–</emphasis>
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corpus bursae gourd shaped, rounded posteriorly, abruptly tapered anteriorly into appendix-like tube; ductus bursae rugose with bands of sclerite, irregularly twisted to straight, heavily sclerotized posterior 1/3 of ductus bursae; apophyses long and narrow with triangular plate near middle of posterior apophysis; anal papillae broad, laterally flattened, setae sparse.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="16" pageNumber="133" type="distribution">
|
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<paragraph blockId="16.[140,1108,166,439]" pageId="16" pageNumber="133">
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<emphasis bold="true" box="[185,501,166,193]" pageId="16" pageNumber="133">
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<emphasis box="[185,336,166,192]" inLineHeading="true" pageId="16" pageNumber="133" reason="1">Distribution</emphasis>
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and biology.
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</emphasis>
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Known only from southwestern
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<collectingRegion box="[903,1053,167,193]" country="United States of America" name="New Mexico" pageId="16" pageNumber="133">New Mexico</collectingRegion>
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and southeastern
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<collectingRegion box="[289,381,202,228]" country="United States of America" name="Arizona" pageId="16" pageNumber="133">Arizona</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="16" pageNumber="133" type="discussion">
|
||
<paragraph blockId="16.[140,1108,166,439]" pageId="16" pageNumber="133">
|
||
<emphasis bold="true" box="[185,298,237,263]" pageId="16" pageNumber="133">
|
||
<emphasis box="[185,293,237,263]" inLineHeading="true" pageId="16" pageNumber="133" reason="1">Remarks</emphasis>
|
||
.
|
||
</emphasis>
|
||
This species would not normally be associated with the
|
||
<taxonomicName authorityName="Latreille" authorityYear="1809" box="[936,1055,237,263]" class="Insecta" family="Noctuidae" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="subFamily" subFamily="Noctuinae">Noctuinae</taxonomicName>
|
||
Elaphrini, except by the genitalia and DNA. It is confused with
|
||
<taxonomicName box="[861,987,272,298]" class="Insecta" family="Noctuidae" genus="Marimatha" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[861,987,272,298]" italics="true" pageId="16" pageNumber="133">Marimatha</emphasis>
|
||
</taxonomicName>
|
||
species in all collections where we have found material. It can readily be distinguished from
|
||
<taxonomicName box="[140,266,343,369]" class="Insecta" family="Noctuidae" genus="Marimatha" kingdom="Animalia" order="Lepidoptera" pageId="16" pageNumber="133" phylum="Arthropoda" rank="genus">
|
||
<emphasis box="[140,266,343,369]" italics="true" pageId="16" pageNumber="133">Marimatha</emphasis>
|
||
</taxonomicName>
|
||
by the narrow, acutely pointed forewing, the lack of a dark discal spot, the raised nodular sclerite of the metathoracic ear, and the characteristics of the genitalia given above.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |