171 lines
20 KiB
XML
171 lines
20 KiB
XML
<document id="FCDF92C4A1C50EA3A890011567BBA0C4" ID-DOI="10.11646/zootaxa.2944.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="5283420" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629731250194" checkinUser="felipe" docAuthor="Perkins, Philip D." docDate="2011" docId="03B087E55B2EFFDFFF79F14BFE50FEA9" docLanguage="en" docName="zt02944p417.pdf" docOrigin="Zootaxa 2944 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1" docStyle="DocumentStyle:D239614CE4198176A422035174489AB1.4:Zootaxa.2009-2012.monograph" docStyleId="D239614CE4198176A422035174489AB1" docStyleName="Zootaxa.2009-2012.monograph" docStyleVersion="4" docTitle="Hydraena akameku Perkins 2011, new species" docType="treatment" docVersion="5" lastPageNumber="77" masterDocId="FF89FF9D5B65FF93FFEEF22DFFB6FF96" masterDocTitle="New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944" masterLastPageNumber="417" masterPageNumber="1" pageNumber="76" updateTime="1699103370621" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B37781AFA634A4B77DC63D61EAB27E18">New species (130) of the hyperdiverse aquatic beetle genus Hydraena Kugelann from Papua New Guinea, and a preliminary analysis of areas of endemism (Coleoptera: Hydraenidae) 2944</mods:title>
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<mods:namePart id="C8182713622F8A3233B938A149B76443">Perkins, Philip D.</mods:namePart>
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<mods:title id="C86AA2197A42ADC3A82F2C0618887CF1">Zootaxa</mods:title>
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<mods:part id="9C4A45096BDE38DD448F76FA9E60782F">
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<mods:date id="36A1D0DED881B165A19449E512D82ABF">2011</mods:date>
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<mods:number id="9F8E3C1C957D97EB77BFC7A25F6CD945">2011-07-08</mods:number>
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<mods:number id="02A43B2D19A9F3DEA3A7CC64F5CC9AE0">2944</mods:number>
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<mods:url id="0B4520009AA12A053912EF8A739FE031">https://biotaxa.org/Zootaxa/article/view/zootaxa.2944.1.1</mods:url>
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<mods:identifier id="DF9E4443DFC74E069A33FCA2AAAB8523" type="DOI">10.11646/zootaxa.2944.1.1</mods:identifier>
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<treatment id="03B087E55B2EFFDFFF79F14BFE50FEA9" ID-DOI="http://doi.org/10.5281/zenodo.5291612" ID-Zenodo-Dep="5291612" LSID="urn:lsid:plazi:treatment:03B087E55B2EFFDFFF79F14BFE50FEA9" httpUri="http://treatment.plazi.org/id/03B087E55B2EFFDFFF79F14BFE50FEA9" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">
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<subSubSection id="C30365785B2EFFD8FF79F14BFD92FC16" box="[151,548,870,896]" pageId="75" pageNumber="76" type="nomenclature">
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<paragraph id="8BA636F35B2EFFD8FF79F14BFD92FC16" blockId="75.[151,548,870,929]" box="[151,548,870,896]" pageId="75" pageNumber="76">
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<heading id="D0EE819F5B2EFFD8FF79F14BFD92FC16" bold="true" box="[151,548,870,896]" fontSize="11" level="1" pageId="75" pageNumber="76" reason="1">
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<emphasis id="B96DEAE15B2EFFD8FF79F14BFD92FC16" bold="true" box="[151,548,870,896]" pageId="75" pageNumber="76">
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<taxonomicName id="4C194D705B2EFFD8FF79F14BFE33FC16" ID-CoL="3N352" authority="Perkins, 2011" authorityName="Perkins" authorityYear="2011" box="[151,389,870,896]" class="Insecta" family="Hydraenidae" genus="Hydraena" kingdom="Animalia" order="Coleoptera" pageId="75" pageNumber="76" phylum="Arthropoda" rank="species" species="akameku" status="sp. nov.">
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<emphasis id="B96DEAE15B2EFFD8FF79F14BFE33FC16" bold="true" box="[151,389,870,896]" italics="true" pageId="75" pageNumber="76">Hydraena akameku</emphasis>
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</taxonomicName>
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,
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C30365785B2EFFD8FF79F1A4FE0CFC37" box="[151,442,905,929]" pageId="75" pageNumber="76" type="description">
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<paragraph id="8BA636F35B2EFFD8FF79F1A4FE0CFC37" blockId="75.[151,548,870,929]" box="[151,442,905,929]" pageId="75" pageNumber="76">
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(
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<figureCitation id="13222A765B2EFFD8FF71F1A4FEBDFC37" box="[159,267,905,929]" captionStart="FIGURE 126" captionStartId="239.[151,250,1753,1776]" captionTargetBox="[207,1375,193,1713]" captionTargetId="figure-26@239.[195,1391,193,1732]" captionTargetPageId="239" captionText="FIGURE 126. Hydraena akameku, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283664" httpUri="https://zenodo.org/record/5283664/files/figure.png" pageId="75" pageNumber="76">Figs. 126</figureCitation>
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,
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<figureCitation id="13222A765B2EFFD8FEF8F1A4FEF4FC37" box="[278,322,905,929]" captionStart="FIGURES 128–129" captionStartId="241.[151,264,1650,1673]" captionText="FIGURES 128–129. Aedeagi of holotypes. 128. H. akameku. 129. H. quadriplumipes." pageId="75" pageNumber="76">128</figureCitation>
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,
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<figureCitation id="13222A765B2EFFD8FEA0F1A4FECCFC37" box="[334,378,905,929]" captionStart="FIGURES 400–403" captionStartId="372.[151,264,1650,1673]" captionTargetBox="[271,1319,208,1611]" captionTargetId="figure-28@372.[235,1351,193,1629]" captionTargetPageId="372" captionText="FIGURES 400–403. Female tergite X, gonocoxite, and spermatheca of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284025" httpUri="https://zenodo.org/record/5284025/files/figure.png" pageId="75" pageNumber="76">403</figureCitation>
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,
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<figureCitation id="13222A765B2EFFD8FE68F1A4FE07FC37" box="[390,433,905,929]" captionStart="FIGURES 487–490" captionStartId="397.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1578]" captionTargetId="figure-20@397.[165,1421,193,1629]" captionTargetPageId="397" captionText="FIGURES 487–490. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284103" httpUri="https://zenodo.org/record/5284103/files/figure.png" pageId="75" pageNumber="76">489</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C30365785B2EFFD8FF79F1FCFE70FBA7" pageId="75" pageNumber="76" type="materials_examined">
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<paragraph id="8BA636F35B2EFFD8FF79F1FCFE70FBA7" blockId="75.[151,1436,977,2010]" pageId="75" pageNumber="76">
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Type Material.
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<materialsCitation id="3B713CAE5B2EFFD8FEA0F1FCFCC8FB9B" collectingDate="2006-11-25" collectionCode="ZSM" collectorName="Balke & Kinibel" country="Papua New Guinea" elevation="750" latitude="-5.821783" location="Akameku - Brahmin" longLatPrecision="1" longitude="145.40648" municipality="Bismarck Range" pageId="75" pageNumber="76" specimenCount="1" stateProvince="Madang" typeStatus="holotype">
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<typeStatus id="54A288515B2EFFD8FEA0F1FCFE0DFC7F" box="[334,443,977,1001]" pageId="75" pageNumber="76" type="holotype">
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<emphasis id="B96DEAE15B2EFFD8FEA0F1FCFE0DFC7F" bold="true" box="[334,443,977,1001]" pageId="75" pageNumber="76">Holotype</emphasis>
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</typeStatus>
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(male):
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<emphasis id="B96DEAE15B2EFFD8FDF1F1FCFD4DFC7C" bold="true" box="[543,763,977,1002]" pageId="75" pageNumber="76">
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<collectingRegion id="49DDF8115B2EFFD8FDF1F1FCFD41FC7C" box="[543,759,977,1002]" country="Papua New Guinea" name="Madang" pageId="75" pageNumber="76">Madang Province</collectingRegion>
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:
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</emphasis>
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<location id="8EC660285B2EFFD8FCE6F1FCFC51FC7F" LSID="urn:lsid:plazi:treatment:03B087E55B2EFFDFFF79F14BFE50FEA9:8EC660285B2EFFD8FCE6F1FCFC51FC7F" box="[776,999,977,1001]" country="Papua New Guinea" latitude="-5.821783" longLatPrecision="1" longitude="145.40648" municipality="Bismarck Range" name="Akameku - Brahmin" pageId="75" pageNumber="76" stateProvince="Madang">Akameku–Brahmin</location>
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,
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<collectingMunicipality id="6BC2AC895B2EFFD8FC1CF1FCFB18FC7C" box="[1010,1198,977,1002]" pageId="75" pageNumber="76">Bismarck Range</collectingMunicipality>
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,
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<quantity id="4CE19B165B2EFFD8FB57F1FCFAB4FC7F" box="[1209,1282,977,1001]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="75" pageNumber="76" unit="m" value="750.0">
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<elevation id="0034D1C05B2EFFD8FB57F1FCFAB4FC7F" box="[1209,1282,977,1001]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="75" pageNumber="76" unit="m" value="750.0">750 m</elevation>
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</quantity>
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,
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<geoCoordinate id="EE2D50345B2EFFD8FAE3F1FCFA20FC7F" box="[1293,1430,977,1001]" degrees="5" direction="south" minutes="49.307" orientation="latitude" pageId="75" pageNumber="76" precision="1" value="-5.821783">5° 49.307' S</geoCoordinate>
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,
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<geoCoordinate id="EE2D50345B2EFFD8FF79F1D8FEF2FB9B" box="[151,324,1013,1037]" degrees="145" direction="east" minutes="24.389" orientation="longitude" pageId="75" pageNumber="76" precision="1" value="145.40648">145° 24.389' E</geoCoordinate>
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,
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<date id="FFA710335B2EFFD8FEBCF1D8FE64FB9B" box="[338,466,1013,1037]" pageId="75" pageNumber="76" value="2006-11-25">
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<collectingDate id="EFE3E9DB5B2EFFD8FEBCF1D8FE64FB9B" box="[338,466,1013,1037]" pageId="75" pageNumber="76" value="2006-11-25">25 xi 2006</collectingDate>
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</date>
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,
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<collectorName id="26EC53255B2EFFD8FE0EF1D8FD97FB9B" box="[480,545,1013,1037]" pageId="75" pageNumber="76">Balke</collectorName>
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&
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<collectorName id="26EC53255B2EFFD8FDA4F1D8FD28FB9B" box="[586,670,1013,1037]" pageId="75" pageNumber="76">Kinibel</collectorName>
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(
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<collectingCountry id="F30E76635B2EFFD8FD41F1D8FD5FFB9B" box="[687,745,1013,1037]" name="Papua New Guinea" pageId="75" pageNumber="76">PNG</collectingCountry>
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114) (
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<collectionCode id="ED08AE365B2EFFD8FCD8F1D8FCC3FB9B" box="[822,885,1013,1037]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="75" pageNumber="76" type="Museum">ZSM</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B713CAE5B2EFFD8FC65F1D8FE74FBA7" collectingDate="2006-11-25" collectionCode="MCZ, NHM, NMW, ZSM" collectorName="Balke & Kinibel" country="Papua New Guinea" elevation="750" latitude="-5.821783" location="Same" longLatPrecision="1" longitude="145.40648" municipality="Bismarck Range" pageId="75" pageNumber="76" specimenCount="1" stateProvince="Paratypes" typeStatus="paratype">
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<emphasis id="B96DEAE15B2EFFD8FC65F1D8FBB5FB9B" bold="true" box="[907,1027,1013,1037]" pageId="75" pageNumber="76">
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<typeStatus id="54A288515B2EFFD8FC65F1D8FBB5FB9B" box="[907,1027,1013,1037]" pageId="75" pageNumber="76" type="paratype">Paratypes</typeStatus>
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</emphasis>
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:
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<location id="8EC660285B2EFFD8FBF3F1D8FBEDFB9B" LSID="urn:lsid:plazi:treatment:03B087E55B2EFFDFFF79F14BFE50FEA9:8EC660285B2EFFD8FBF3F1D8FBEDFB9B" box="[1053,1115,1013,1037]" country="Papua New Guinea" latitude="-5.821783" longLatPrecision="1" longitude="145.40648" municipality="Bismarck Range" name="Same" pageId="75" pageNumber="76" stateProvince="Paratypes">Same</location>
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data as holotype (17
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<collectionCode id="ED08AE365B2EFFD8FAB6F1D8FA21FB9B" box="[1368,1431,1013,1037]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="75" pageNumber="76" type="Museum">MCZ</collectionCode>
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,
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<collectionCode id="ED08AE365B2EFFD8FF79F637FF6DFBA7" box="[151,219,1050,1073]" pageId="75" pageNumber="76">NHM</collectionCode>
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,
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<collectionCode id="ED08AE365B2EFFD8FF08F637FE98FBA7" box="[230,302,1050,1073]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="75" pageNumber="76">NMW</collectionCode>
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,
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<collectingCountry id="F30E76635B2EFFD8FED7F634FEC7FBA7" box="[313,369,1049,1073]" name="Papua New Guinea" pageId="75" pageNumber="76">PNG</collectingCountry>
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,
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<collectionCode id="ED08AE365B2EFFD8FE92F634FE0CFBA7" box="[380,442,1049,1073]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="75" pageNumber="76" type="Museum">ZSM</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C30365785B2EFFD8FF29F610FD78FB0B" pageId="75" pageNumber="76" type="diagnosis">
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<paragraph id="8BA636F35B2EFFD8FF29F610FD78FB0B" blockId="75.[151,1436,977,2010]" pageId="75" pageNumber="76">
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<emphasis id="B96DEAE15B2EFFD8FF29F610FE65FBC0" bold="true" box="[199,467,1085,1110]" pageId="75" pageNumber="76">Differential Diagnosis.</emphasis>
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Recognized by the narrow body form, the coarsely densely punctate dorsum, and the very narrow mesoventral intercoxal process (P2). Neither the dorsal habitus nor the male genitalia show a close affinity with other species groups (
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<figureCitation id="13222A765B2EFFD8FDF2F6A8FD31FB0B" box="[540,647,1157,1181]" captionStart="FIGURE 126" captionStartId="239.[151,250,1753,1776]" captionTargetBox="[207,1375,193,1713]" captionTargetId="figure-26@239.[195,1391,193,1732]" captionTargetPageId="239" captionText="FIGURE 126. Hydraena akameku, holotype, dorsal and lateral habitus, and venter." figureDoi="http://doi.org/10.5281/zenodo.5283664" httpUri="https://zenodo.org/record/5283664/files/figure.png" pageId="75" pageNumber="76">Figs. 126</figureCitation>
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,
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<figureCitation id="13222A765B2EFFD8FD7DF6A8FD77FB0B" box="[659,705,1157,1181]" captionStart="FIGURES 128–129" captionStartId="241.[151,264,1650,1673]" captionText="FIGURES 128–129. Aedeagi of holotypes. 128. H. akameku. 129. H. quadriplumipes." pageId="75" pageNumber="76">128</figureCitation>
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).
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</paragraph>
|
||
</subSubSection>
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||
<subSubSection id="C30365785B2EFFDFFF29F684FA9AFF42" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76" type="description">
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||
<paragraph id="8BA636F35B2EFFD8FF29F684FCBDFB73" blockId="75.[151,1436,977,2010]" pageId="75" pageNumber="76">
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||
<emphasis id="B96DEAE15B2EFFD8FF29F684FEEEFB54" bold="true" box="[199,344,1193,1218]" inLineHeading="true" pageId="75" pageNumber="76" reason="1">Description.</emphasis>
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Size:
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<typeStatus id="54A288515B2EFFD8FE42F684FDB8FB57" box="[428,526,1193,1217]" pageId="75" pageNumber="76" type="holotype">holotype</typeStatus>
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(length/width, mm): body (length to elytral apices) 1.39/0.56; head 0.23/0.33; pronotum 0.34/0.44, PA 0.36, PB 0.34; elytra 0.83/0.56.
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||
</paragraph>
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<paragraph id="8BA636F35B2EFFD8FF79F6DFFCB2FAC7" blockId="75.[151,1436,977,2010]" pageId="75" pageNumber="76">Dorsum of head dark brown to piceous; pronotum light brown to testaceous in front of and behind piceous fascia, ratios of color bands, as measured in midline, ca. 6/13/4; elytra dark brown; legs light brown; maxillary palpi testaceous, tip very slightly darker. Dorsum weakly shining.</paragraph>
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<paragraph id="8BA636F35B2EFFD8FF29F773FAB7F9E7" blockId="75.[151,1436,977,2010]" pageId="75" pageNumber="76">Frons punctures ca. 2xef near eyes, smaller and sparser medially; interstices shining, 1–3xpd. Clypeus effacedly microreticulate laterally, very finely sparsely punctulate and shining medially. Mentum very sparsely very finely punctulate, strongly shining. Postmentum effacedly microreticulate in posterior 1/2, otherwise finely sparsely punctulate. Genae very slightly raised, weakly shining, without posterior ridge. Pronotum cordiform, median 2/3 of anterior margin emarginate; sides straight or very slightly emarginate between anterior angle and widest part, sinuate between widest part and posterior angle; punctures on disc ca. 1xpd largest frons punctures, interstices shining, ca. 1–2xpd, punctures anteriorly and posteriorly denser than those on disc; PF1 and PF4 very shallow; PF2 moderately deep, medially conjointly shallowly confluent; PF3 moderately deep, wide.</paragraph>
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<paragraph id="8BA636F35B2EFFD8FF29F450FD0BF8B3" blockId="75.[151,1436,977,2010]" pageId="75" pageNumber="76">Elytra widest at about midlength; summit of posterior declivity at or very near midlength; lateral explanate margins moderately wide; on basal 1/3 punctures ca. 1xpd largest pronotal punctures, a few punctures subserial, punctures becoming gradually smaller toward posterior. Intervals not raised, very weakly shining, on disc ca. 1– 2xpd, as are interstices between punctures of a row. Apices in dorsal aspect conjointly rounded, in posterior aspect margins forming shallow angle with one another.</paragraph>
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<paragraph id="8BA636F35B2EFFDFFF29F51CFA9AFF42" blockId="75.[151,1436,977,2010]" lastBlockId="76.[151,1436,151,319]" lastPageId="76" lastPageNumber="77" pageId="75" pageNumber="76">Ratios of P2 width and plaque shape (P2/w/l/s) ca. 1/2.5/5/2. P1 laminate; median carina sinuate in profile, slightly raised between coxae. P2 very narrow, length/width ca. 6/1, sides very slightly converging toward blunt apex. Plaques large, slightly raised, at sides of narrow median depression, anteriorly slightly tapering and slightly convergent. Metaventrite flat between mesocoxae. AIS width at straight posterior margin ca. 4x P2. All legs of moderate length. Profemur (male) with minute tubercle on ventral margin near trochanter; protibia very slightly arcuate, increasing in width from base to apex. Mesotibia straight. Metatibia straight, very slightly increasing in width in distal 1/2. Abdominal apex symmetrical; last tergite (male) with large, deep apicomedian notch.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B29FFDFFF29F2CDFCB9FF61" box="[199,783,223,248]" pageId="76" pageNumber="77" type="etymology">
|
||
<paragraph id="8BA636F35B29FFDFFF29F2CDFCB9FF61" blockId="76.[151,1436,151,319]" box="[199,783,223,248]" pageId="76" pageNumber="77">
|
||
<emphasis id="B96DEAE15B29FFDFFF29F2CDFEFBFF6E" bold="true" box="[199,333,224,248]" inLineHeading="true" pageId="76" pageNumber="77" reason="1">Etymology.</emphasis>
|
||
Named in reference to the
|
||
<typeStatus id="54A288515B29FFDFFD90F2CDFD18FF6E" box="[638,686,224,248]" pageId="76" pageNumber="77">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C30365785B29FFDFFF29F32EFE50FEA9" pageId="76" pageNumber="77" type="distribution">
|
||
<paragraph id="8BA636F35B29FFDFFF29F32EFE50FEA9" blockId="76.[151,1436,151,319]" pageId="76" pageNumber="77">
|
||
<emphasis id="B96DEAE15B29FFDFFF29F32EFED6FE8A" bold="true" box="[199,352,259,284]" inLineHeading="true" pageId="76" pageNumber="77" reason="1">Distribution.</emphasis>
|
||
Currently known only from the
|
||
<typeStatus id="54A288515B29FFDFFD3EF329FCB6FE8A" box="[720,768,260,284]" pageId="76" pageNumber="77">type</typeStatus>
|
||
locality in the Bismarck Range, in the eastern part of Area 2; elevation
|
||
<quantity id="4CE19B165B29FFDFFECFF30AFEDEFEA9" box="[289,360,295,319]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="76" pageNumber="77" unit="m" value="750.0">750 m</quantity>
|
||
(
|
||
<figureCitation id="13222A765B29FFDFFE99F30AFE6DFEA9" box="[375,475,295,319]" captionStart="FIGURES 487–490" captionStartId="397.[151,264,1650,1673]" captionTargetBox="[165,1421,224,1578]" captionTargetId="figure-20@397.[165,1421,193,1629]" captionTargetPageId="397" captionText="FIGURES 487–490. Geographical distributions of Hydraena species." figureDoi="http://doi.org/10.5281/zenodo.5284103" httpUri="https://zenodo.org/record/5284103/files/figure.png" pageId="76" pageNumber="77">Fig. 489</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |