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<document ID-DOI="http://doi.org/10.5281/zenodo.3788372" ID-GBIF-Dataset="a757de0f-b7ba-4ef8-9c95-24cfc4226b76" ID-GBIF-Taxon="163747474" ID-ISSN="13132970" ID-Zenodo-Dep="3788372" ID-ZooBank="EB891295-3333-4BB1-B7EA-F118C4D6B13A" approvalRequired="26" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="23" approvalRequired_for_treatments="2" checkinTime="1587953814852" checkinUser="plazi" docAuthor="Paulsen, M. J. &amp; Smith, Andrew" docDate="2010" docId="AA0687ED8270FFA7FF63FEA53DCBFAA4" docLanguage="en" docName="ZK_article_2192.pdf" docOrigin="ZooKeys 43 (43)" docStyle="DocumentStylede.uka.ipd.idaho.easyIO.settings.Settings@6ed1fdb7" docStyleName="zookeys.2008.journal_article" docTitle="Chiasognathus mniszechii Thomson 1862" docType="treatment" docVersion="4" lastPageId="23" lastPageNumber="56" masterDocId="563FFF958266FFB0FFEFFFF33E69FFCA" masterDocTitle="Revision of the genus Chiasognathus Stephens of southern South America with the description of a new species (Coleoptera, Lucanidae, Lucaninae, Chiasognathini)" masterLastPageNumber="63" masterPageNumber="33" pageId="22" pageNumber="55" updateTime="1643557281858" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revision of the genus Chiasognathus Stephens of southern South America with the description of a new species (Coleoptera, Lucanidae, Lucaninae, Chiasognathini)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Paulsen, M. J.</mods:namePart>
<mods:affiliation>University of Nebraska State Museum, Lincoln, United States of America</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Smith, Andrew</mods:namePart>
<mods:affiliation>,, Canada</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2010</mods:date>
<mods:detail type="pubDate">
<mods:number>2010-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>43</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>43</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>33</mods:start>
<mods:end>63</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.3897/zookeys.43.397</mods:identifier>
<mods:identifier type="GBIF-Dataset">a757de0f-b7ba-4ef8-9c95-24cfc4226b76</mods:identifier>
<mods:identifier type="ISSN">13132970</mods:identifier>
<mods:identifier type="Zenodo-Dep">576657</mods:identifier>
<mods:identifier type="ZooBank">EB891295-3333-4BB1-B7EA-F118C4D6B13A</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3788372" ID-GBIF-Taxon="163747474" ID-Zenodo-Dep="3788372" LSID="urn:lsid:plazi:treatment:AA0687ED8270FFA7FF63FEA53DCBFAA4" httpUri="http://treatment.plazi.org/id/AA0687ED8270FFA7FF63FEA53DCBFAA4" lastPageId="23" lastPageNumber="56" pageId="22" pageNumber="55">
<subSubSection pageId="22" pageNumber="55" type="nomenclature">
<paragraph blockId="22.[140,643,342,404]" box="[140,643,342,369]" pageId="22" pageNumber="55">
<heading box="[140,643,342,369]" fontSize="11" level="2" pageId="22" pageNumber="55" reason="3">
<taxonomicName authority="Thomson, 1862" authorityName="Thomson" authorityYear="1862" box="[140,643,342,369]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="mniszechii">
<emphasis bold="true" box="[140,643,342,369]" pageId="22" pageNumber="55">
<emphasis bold="true" box="[140,447,342,369]" italics="true" pageId="22" pageNumber="55">Chiasognathus mniszechii</emphasis>
<bibRefCitation author="Thomson J" box="[454,643,342,369]" pageId="22" pageNumber="55" pagination="389 - 436" refId="ref15544" refString="Thomson J (1862) Catalogue des lucanides de la collection de M. James Thomson suivi d'un appendix renfermant la description des coupes generiques et specifiques nouvelles. Annales de la Societe Entomologique de France 4 (2): 389 - 436." type="journal article" year="1862">Thomson, 1862</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="22.[140,643,342,404]" box="[140,426,378,404]" pageId="22" pageNumber="55">Figs 6, 19, 21, 2324, 31</paragraph>
</subSubSection>
<subSubSection pageId="22" pageNumber="55" type="reference_group">
<paragraph blockId="22.[140,1108,448,1249]" pageId="22" pageNumber="55">
<treatmentCitationGroup pageId="22" pageNumber="55">
<treatmentCitation author="Thomson J" box="[140,672,448,474]" page="406" pageId="22" pageNumber="55" year="1862">
<taxonomicName authority="Thomson, 1862: 406" authorityName="Thomson" authorityPageNumber="406" authorityYear="1862" box="[140,672,448,474]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="mniszechii">
<emphasis box="[140,419,448,474]" italics="true" pageId="22" pageNumber="55">Chiasognathus mniszechii</emphasis>
<bibRefCitation author="Thomson J" box="[428,672,448,474]" pageId="22" pageNumber="55" pagination="389 - 436" refId="ref15544" refString="Thomson J (1862) Catalogue des lucanides de la collection de M. James Thomson suivi d'un appendix renfermant la description des coupes generiques et specifiques nouvelles. Annales de la Societe Entomologique de France 4 (2): 389 - 436." type="journal article" year="1862">Thomson, 1862: 406</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis bold="true" box="[685,845,448,474]" pageId="22" pageNumber="55">revised status</emphasis>
. Type material:
<typeStatus pageId="22" pageNumber="55">Lectotype</typeStatus>
male (MNHN) labeled: a) bordered label “Ex-Musaeo / Mniszech”; b) bordered label “
<emphasis box="[325,361,519,545]" italics="true" pageId="22" pageNumber="55">Th.</emphasis>
/ TYPE”; c) handwritten “
<emphasis box="[663,781,519,545]" italics="true" pageId="22" pageNumber="55">Mniszechii</emphasis>
/ Type Thoms / 4.1862
<collectingCountry name="Chile" pageId="22" pageNumber="55">Chili</collectingCountry>
/ [300]
<emphasis box="[288,299,554,580]" italics="true" pageId="22" pageNumber="55"></emphasis>
; d) “MUSÉUM PARIS / [280D.]”; e) red paper, “TYPE”; f) “[1993] /
<taxonomicName authorityName="Reiche" authorityYear="1850" box="[185,457,589,615]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="jousselinii">
<emphasis box="[185,457,589,615]" italics="true" pageId="22" pageNumber="55">Chiasognathus jousselinii</emphasis>
</taxonomicName>
/ Reiche / Det: F. Chalumeau &amp; B. Brochier”; g) handwritten “
<emphasis box="[289,454,624,650]" italics="true" pageId="22" pageNumber="55">
<taxonomicName box="[289,450,624,650]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="genus">Chiasognathus</taxonomicName>
.
</emphasis>
/
<emphasis box="[484,590,624,650]" italics="true" pageId="22" pageNumber="55">jousselini.</emphasis>
Reich. / Det: JP. LACROIX.”; h) bordered label “Southern Neotropical Scarabs / database #
<collectingCountry box="[780,815,659,685]" name="American Samoa" pageId="22" pageNumber="55">AS</collectingCountry>
2618226 /
<taxonomicName authorityName="Thomson" authorityYear="1862" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="mniszechii">
<emphasis italics="true" pageId="22" pageNumber="55">Chiasognathus mniszechii</emphasis>
</taxonomicName>
/
<bibRefCitation author="Thomson J" box="[324,506,695,721]" pageId="22" pageNumber="55" pagination="389 - 436" refId="ref15544" refString="Thomson J (1862) Catalogue des lucanides de la collection de M. James Thomson suivi d'un appendix renfermant la description des coupes generiques et specifiques nouvelles. Annales de la Societe Entomologique de France 4 (2): 389 - 436." type="journal article" year="1862">Thomson, 1862</bibRefCitation>
[male symbol] / DET: A.B.T. SMITH 2009”; i) red label “
<taxonomicName box="[257,414,730,756]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis box="[257,414,730,756]" italics="true" pageId="22" pageNumber="55">Chiasognathus</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Reiche" authorityYear="1850" box="[438,544,730,756]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="jousselinii">
<emphasis box="[438,544,730,756]" italics="true" pageId="22" pageNumber="55">jousselinii</emphasis>
</taxonomicName>
[male symbol] /
<bibRefCitation author="Thomson J" box="[737,918,730,756]" pageId="22" pageNumber="55" pagination="389 - 436" refId="ref15544" refString="Thomson J (1862) Catalogue des lucanides de la collection de M. James Thomson suivi d'un appendix renfermant la description des coupes generiques et specifiques nouvelles. Annales de la Societe Entomologique de France 4 (2): 389 - 436." type="journal article" year="1862">Thomson, 1862</bibRefCitation>
/
<typeStatus box="[942,1108,730,756]" pageId="22" pageNumber="55">LECTOTYPE</typeStatus>
/ Det. M.J. Paulsen”.
<typeStatus box="[442,553,766,792]" pageId="22" pageNumber="55">Lectotype</typeStatus>
designated by
<bibRefCitation author="Chalumeau F &amp; Brochier B" box="[730,1107,765,792]" pageId="22" pageNumber="55" pagination="18 - 24" refId="ref14099" refString="Chalumeau F, Brochier B (1995) Les Chiasognathinae: genres, sous-genres et synonymies (Coleoptera, Lucanidae). Bulletin de la Societe Sciences Nat 83: 18 - 24." type="journal article" year="1995">Chalumeau and Brochier (1995)</bibRefCitation>
by inference under ICZN Article 74.6 when they referred to the specimen as the
<typeStatus box="[185,288,835,861]" pageId="22" pageNumber="55">holotype</typeStatus>
.
<bibRefCitation author="Thomson J" box="[301,494,835,862]" pageId="22" pageNumber="55" pagination="389 - 436" refId="ref15544" refString="Thomson J (1862) Catalogue des lucanides de la collection de M. James Thomson suivi d'un appendix renfermant la description des coupes generiques et specifiques nouvelles. Annales de la Societe Entomologique de France 4 (2): 389 - 436." type="journal article" year="1862">Thomson (1862)</bibRefCitation>
gave no definitive indication that there was only one specimen in the type series. Type locality: “
<collectingCountry box="[674,738,871,897]" name="Chile" pageId="22" pageNumber="55">Chili</collectingCountry>
.”
</treatmentCitationGroup>
</paragraph>
<paragraph blockId="22.[140,1108,448,1249]" pageId="22" pageNumber="55">
<treatmentCitationGroup pageId="22" pageNumber="55">
<treatmentCitation author="Kriesche R" box="[140,680,906,932]" page="94" pageId="22" pageNumber="55" year="1919">
<taxonomicName authority="Kriesche, 1919: 94" authorityName="Kriesche" authorityPageNumber="94" authorityYear="1917" box="[140,680,906,932]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="schoenemanni" status="new synonymy">
<emphasis box="[140,455,906,932]" italics="true" pageId="22" pageNumber="55">Chiasognathus schoenemanni</emphasis>
<bibRefCitation author="Kriesche R" box="[462,680,906,932]" pageId="22" pageNumber="55" pagination="93 - 95" refId="ref14578" refString="Kriesche R (1919) Neue Chiasognathinen (Coleopt. Lucanid.). Archiv fur Naturgeschichte (A) 83: 93 - 95. [Dated 1917]." type="journal article" year="1919">Kriesche, 1919: 94</bibRefCitation>
</taxonomicName>
</treatmentCitation>
,
<emphasis bold="true" box="[692,871,906,932]" pageId="22" pageNumber="55">
<taxonomicNameLabel box="[692,871,906,932]" pageId="22" pageNumber="55">new synonymy</taxonomicNameLabel>
</emphasis>
. Type material:
<typeStatus pageId="22" pageNumber="55">Lectotype</typeStatus>
male (SMTD) designated by
<bibRefCitation author="Chalumeau F &amp; Brochier B" box="[609,987,941,968]" pageId="22" pageNumber="55" pagination="18 - 24" refId="ref14099" refString="Chalumeau F, Brochier B (1995) Les Chiasognathinae: genres, sous-genres et synonymies (Coleoptera, Lucanidae). Bulletin de la Societe Sciences Nat 83: 18 - 24." type="journal article" year="1995">Chalumeau and Brochier (1995)</bibRefCitation>
labeled a) handwritten “
<collectingCountry box="[341,405,976,1002]" name="Chile" pageId="22" pageNumber="55">Chile</collectingCountry>
/ Bader von Longavi, / Parral”; b) handwritten “
<taxonomicName box="[950,1107,976,1002]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis box="[950,1107,976,1002]" italics="true" pageId="22" pageNumber="55">Chiasognathus</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Kriesche" authorityYear="1917" box="[202,352,1011,1037]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="schoenemanni">
<emphasis box="[202,352,1011,1037]" italics="true" pageId="22" pageNumber="55">schoenemanni</emphasis>
</taxonomicName>
/
<emphasis box="[377,457,1012,1038]" italics="true" pageId="22" pageNumber="55">Cotyp!”</emphasis>
; c) “sammlung KRIESCHE / Ankauf 1974”; d) red label “Typ”; e) “Staatl. Museum für / Tierkunde. Dresden”; f) bordered “[-93] /
<taxonomicName class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="genus">Chiasognathus</taxonomicName>
/ schoenemanni / dés. F. Chalumeau &amp; / B. Brochier”; f) “[-94] /
<taxonomicName authorityName="Reiche" authorityYear="1850" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="jousselinii">Chiasognathus jousselinii</taxonomicName>
/ Reiche / Dét. F. Chalumeau &amp; B. Brochier”. g) red label “
<taxonomicName box="[197,354,1152,1178]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis box="[197,354,1152,1178]" italics="true" pageId="22" pageNumber="55">Chiasognathus</emphasis>
</taxonomicName>
/
<taxonomicName authorityName="Kriesche" authorityYear="1917" box="[384,534,1152,1178]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="55" phylum="Arthropoda" rank="species" species="schoenemanni">
<emphasis box="[384,534,1152,1178]" italics="true" pageId="22" pageNumber="55">schoenemanni</emphasis>
</taxonomicName>
[male symbol] /
<bibRefCitation author="Kriesche R" box="[739,912,1152,1178]" pageId="22" pageNumber="55" pagination="93 - 95" refId="ref14578" refString="Kriesche R (1919) Neue Chiasognathinen (Coleopt. Lucanid.). Archiv fur Naturgeschichte (A) 83: 93 - 95. [Dated 1917]." type="journal article" year="1919">Kriesche, 1919</bibRefCitation>
/
<typeStatus box="[942,1108,1152,1178]" pageId="22" pageNumber="55">LECTOTYPE</typeStatus>
/ Det. M.J. Paulsen”. Five
<typeStatus box="[507,658,1187,1213]" pageId="22" pageNumber="55">paralectotype</typeStatus>
males (ZMHB).Type locality: “
<collectingCountry box="[1037,1103,1187,1213]" name="Chile" pageId="22" pageNumber="55">Chile</collectingCountry>
, Bäder von Longavi, Parral.”
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="23" lastPageNumber="56" pageId="22" pageNumber="55" type="description">
<paragraph blockId="22.[140,1108,1292,1672]" lastBlockId="23.[140,1108,167,1390]" lastPageId="23" lastPageNumber="56" pageId="22" pageNumber="55">
<emphasis bold="true" box="[140,386,1292,1319]" pageId="22" pageNumber="55">
<emphasis box="[140,283,1293,1319]" inLineHeading="true" pageId="22" pageNumber="55" reason="1">Description</emphasis>
.
<emphasis bold="true" box="[298,377,1292,1319]" italics="true" pageId="22" pageNumber="55">Length</emphasis>
:
</emphasis>
22.0
<quantity box="[460,570,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="4.22" pageId="22" pageNumber="55" unit="mm" value="42.2">42.2 mm</quantity>
.
<emphasis bold="true" box="[581,667,1292,1319]" pageId="22" pageNumber="55">
<emphasis bold="true" box="[581,658,1292,1319]" italics="true" pageId="22" pageNumber="55">Width</emphasis>
:
</emphasis>
<quantity box="[675,851,1293,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.275" metricValueMax="1.48" metricValueMin="1.0699999999999998" pageId="22" pageNumber="55" unit="mm" value="12.75" valueMax="14.8" valueMin="10.7">10.714.8 mm</quantity>
.
<emphasis bold="true" box="[862,937,1292,1319]" pageId="22" pageNumber="55">
<emphasis bold="true" box="[862,928,1292,1319]" italics="true" pageId="22" pageNumber="55">Color</emphasis>
:
</emphasis>
Brown to reddish brown, with weak metallic reflections, appearing gray from dense covering of scale-like setae (Figs 2324); setae variably abraded or greased.
<emphasis bold="true" box="[852,925,1363,1390]" pageId="22" pageNumber="55">
<emphasis bold="true" box="[852,917,1363,1390]" italics="true" pageId="22" pageNumber="55">Head</emphasis>
:
</emphasis>
Form subquadrate. Surface punctate; punctures fine to coarse, generally setose with short to long setae, especially laterally. Anterior margin of head roundly produced beyond anterior angles and with median nasus; nasus variable with two distinct or conjoined teeth or reduced to a single process, in females nasus migrated downward from anterior margin. Anterior angles obtusely rounded in dorsal view. Male mandible (Fig. 21) setose, cylindrical, narrow, 23× as long as head, externally sinuate until abruptly, internally curved at apex; apex curved weakly upward, lacking patch of setae. Dentate carina of 1520 peg-like teeth beginning on dorsal margin in basal third (in basal third teeth variably reduced to form low carina), becoming internal in median third, but dorsal again in apical third. Base of mandibles with acute ventral tooth; tooth occasionally obsolete. Female mandible as long as head, externally weakly rounded, dorsally flattened, with median internal tooth. Galeal brush short, shorter than first and second labial palpomeres combined. Antennal scape in males with sparse area of long setae at apex.
<emphasis bold="true" box="[209,338,343,369]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[209,330,343,369]" italics="true" pageId="23" pageNumber="56">Pronotum</emphasis>
:
</emphasis>
Posterior and lateral angles obtusely angulate (rarely subdentate), posterior angle more strongly so in males, both angles almost obsolete in females. Lateral margins distinct, crenulate. Dorsal surface with strong transverse ridge either side of triangular median longitudinal furrow, weak depressions indicated either side of disc before middle in most specimens; furrow and depressions variably setose; setae short to long in males (3× longer than length of distal antennomere of club), short in females.
<emphasis bold="true" box="[240,322,553,580]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[240,314,553,580]" italics="true" pageId="23" pageNumber="56">Elytra</emphasis>
:
</emphasis>
Surface rough, slightly wrinkled, generally obscured by dense cover of scale-like setae (especially in males). Apex subdentate, but lacking spine; elytral margin narrow, beaded. Epipleuron strongly concave.
<emphasis bold="true" box="[752,810,624,651]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[752,803,625,651]" italics="true" pageId="23" pageNumber="56">Legs</emphasis>
:
</emphasis>
Protibiae elongate, serrate externally (Fig. 19); ventral surface along internal margin with teeth well developed in males, variably developed in females.
<emphasis bold="true" box="[592,772,694,721]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[592,764,694,721]" italics="true" pageId="23" pageNumber="56">Male genitalia</emphasis>
:
</emphasis>
Flagellum short, shorter than length of parameres and basal piece together (Fig. 6).
</paragraph>
</subSubSection>
<subSubSection box="[185,1010,765,791]" pageId="23" pageNumber="56" type="distribution">
<paragraph blockId="23.[140,1108,167,1390]" box="[185,1010,765,791]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[185,344,765,791]" pageId="23" pageNumber="56">
<emphasis box="[185,339,765,791]" inLineHeading="true" pageId="23" pageNumber="56" reason="1">Distribution</emphasis>
.
</emphasis>
Central
<collectingCountry box="[444,506,765,791]" name="Chile" pageId="23" pageNumber="56">Chile</collectingCountry>
and adjacent
<collectingRegion box="[664,773,765,791]" country="Argentina" name="Neuquen" pageId="23" pageNumber="56">Neuquén</collectingRegion>
,
<collectingCountry box="[785,897,765,791]" name="Argentina" pageId="23" pageNumber="56">Argentina</collectingCountry>
(Fig. 31).
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="56" type="materials_examined">
<paragraph blockId="23.[140,1108,167,1390]" box="[185,846,799,827]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[185,354,800,826]" pageId="23" pageNumber="56">
<collectingCountry box="[185,354,800,826]" name="Argentina" pageId="23" pageNumber="56">ARGENTINA</collectingCountry>
</emphasis>
(2):
<emphasis bold="true" box="[409,519,799,826]" pageId="23" pageNumber="56">
<collectingRegion box="[409,519,799,826]" country="Argentina" name="Neuquen" pageId="23" pageNumber="56">Neuquén</collectingRegion>
</emphasis>
(2): Parque Nacional Lanín.
</paragraph>
<paragraph blockId="23.[140,1108,167,1390]" pageId="23" pageNumber="56">
<materialsCitation ID-GBIF-Occurrence="2608811618" box="[185,635,835,862]" country="Chile" location="Maule" pageId="23" pageNumber="56" specimenCount="1" stateProvince="Maule">
<emphasis bold="true" box="[185,274,835,861]" pageId="23" pageNumber="56">
<collectingCountry box="[185,274,835,861]" name="Chile" pageId="23" pageNumber="56">CHILE</collectingCountry>
</emphasis>
(60):
<emphasis bold="true" box="[351,427,835,861]" pageId="23" pageNumber="56">
<collectingRegion box="[351,427,835,861]" country="Chile" name="Maule" pageId="23" pageNumber="56">Maule</collectingRegion>
</emphasis>
(1): Estero Leiva
</materialsCitation>
;
<emphasis bold="true" box="[649,730,834,861]" pageId="23" pageNumber="56">
<collectingRegion box="[649,730,834,861]" country="Chile" name="Biobio" pageId="23" pageNumber="56">Biobío</collectingRegion>
</emphasis>
(24): Atacalco, Cordillera Chíllan, Las Trancas, Pemehue;
<emphasis bold="true" box="[456,614,869,897]" pageId="23" pageNumber="56">
La
<collectingRegion box="[493,614,869,896]" country="Chile" name="Araucania" pageId="23" pageNumber="56">Araucanía</collectingRegion>
</emphasis>
(35): Cherquenco, Cunco, Curacautín, Las Raíces, Manzanar.
</paragraph>
<paragraph blockId="23.[140,1108,167,1390]" box="[185,1089,941,968]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[185,459,941,967]" pageId="23" pageNumber="56">
<emphasis box="[185,299,941,967]" inLineHeading="true" pageId="23" pageNumber="56" reason="1">Temporal</emphasis>
distribution.
</emphasis>
January (6), February (32), March (1), December (23).
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="56" type="diagnosis">
<paragraph blockId="23.[140,1108,167,1390]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[185,312,976,1002]" pageId="23" pageNumber="56">
<emphasis box="[185,307,976,1002]" inLineHeading="true" pageId="23" pageNumber="56" reason="1">Diagnosis</emphasis>
.
</emphasis>
This is the most commonly encountered species with dense, velvety pubescence on the elytra of males. If the scales are abraded, males of this species can still be recognized because they have narrower, more sinuate mandibles than the other species in the genus (Fig. 21). In addition,
<taxonomicName authorityName="Thomson" authorityYear="1862" box="[622,769,1082,1108]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="56" phylum="Arthropoda" rank="species" species="mniszechii">
<emphasis box="[622,769,1082,1108]" italics="true" pageId="23" pageNumber="56">C. mniszechii</emphasis>
</taxonomicName>
differs from the other densely scaled species,
<emphasis box="[306,453,1117,1144]" italics="true" pageId="23" pageNumber="56">
<taxonomicName authorityName="Reiche" authorityYear="1850" box="[306,449,1117,1144]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="56" phylum="Arthropoda" rank="species" species="jousselinii">C. jousselinii</taxonomicName>
,
</emphasis>
in having the basal part of the protibiae serrate instead of dentate (Fig. 19), strongly concave epipleura, and a marginal bead on a narrow elytral margin apically.
</paragraph>
</subSubSection>
<subSubSection pageId="23" pageNumber="56" type="discussion">
<paragraph blockId="23.[140,1108,167,1390]" pageId="23" pageNumber="56">
<emphasis bold="true" box="[185,298,1223,1249]" pageId="23" pageNumber="56">
<emphasis box="[185,293,1223,1249]" inLineHeading="true" pageId="23" pageNumber="56" reason="1">Remarks</emphasis>
.
</emphasis>
This species is one of the more commonly collected and widespread species in the genus, and it has been collected at mercury vapor lights. Because the species has for so long been incorrectly synonymized under
<emphasis box="[820,968,1293,1320]" italics="true" pageId="23" pageNumber="56">
<taxonomicName authorityName="Reiche" authorityYear="1850" box="[820,963,1293,1320]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="56" phylum="Arthropoda" rank="species" species="jousselinii">C. jousselinii</taxonomicName>
,
</emphasis>
all distributional records or information previously published for
<taxonomicName authorityName="Reiche" authorityYear="1850" box="[750,889,1328,1355]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="56" phylum="Arthropoda" rank="species" species="jousselinii">
<emphasis box="[750,889,1328,1355]" italics="true" pageId="23" pageNumber="56">C. jousselinii</emphasis>
</taxonomicName>
(
<emphasis box="[904,948,1329,1354]" italics="true" pageId="23" pageNumber="56">e.g.,</emphasis>
<bibRefCitation author="Benesh B" box="[954,1102,1328,1354]" pageId="23" pageNumber="56" refId="ref14030" refString="Benesh B (1960) Coleopterorum Catalogus Supplementa, Pars 8: Lucanidea (sic). W. Junk, Berlin, 178 pp." type="book" year="1960">Benesh 1960</bibRefCitation>
;
<bibRefCitation author="Ocampo FC &amp; Paulsen MJ" box="[140,450,1363,1390]" pageId="23" pageNumber="56" pagination="501 - 504" refId="ref15053" refString="Ocampo FC, Paulsen MJ (2008) Lucanidae. In Claps LE, Debandi G, Roig-Junent S (Eds) Biodiversidad de Artropodos Argentinos, volume 2. Sociedad Entomologica Argentina, 501 - 504." type="book chapter" year="2008">Ocampo and Paulsen 2008</bibRefCitation>
) is likely to apply instead to
<taxonomicName authorityName="Thomson" authorityYear="1862" box="[775,923,1363,1390]" class="Insecta" family="Lucanidae" genus="Chiasognathus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="56" phylum="Arthropoda" rank="species" species="mniszechii">
<emphasis box="[775,923,1363,1390]" italics="true" pageId="23" pageNumber="56">C. mniszechii</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>