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<document id="25488519C44568717C0D93808C9A4FA1" ID-CLB-Dataset="25474" ID-DOI="10.11646/zootaxa.4668.3.7" ID-GBIF-Dataset="ddf01fc8-a165-492e-aa9a-25316f7f4ab6" ID-ISSN="1175-5326" ID-Zenodo-Dep="3449692" ID-ZooBank="AA9BB1DA-8E27-48E2-9A4A-7428C45F6291" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1568897931647" checkinUser="plazi" docAuthor="Volynkin, Anton V. &amp; László, Gyula M." docDate="2019" docId="102387C1FFE6426966F0F976ACC5F882" docLanguage="en" docName="zootaxa.4668.3.7.pdf" docOrigin="Zootaxa 4668 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tumicla mbeghai Volynkin &amp; László 2019, sp. nov." docType="treatment" docUuid="231915D5-68F0-45B6-9524-8D0CBF3892C2" docUuidSource="ZooBank" docVersion="13" lastPageNumber="427" masterDocId="EC1AFFB9FFE5426F6667FFC8AD35FFCD" masterDocTitle="Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini)" masterLastPageNumber="434" masterPageNumber="421" pageNumber="424" updateTime="1698755699668" updateUser="ExternalLinkService">
<mods:mods id="85DE01EA6098B5219B066C56C9DFF61C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DBE4AFC3EBF71390FB0058EE514204D7">
<mods:title id="FB1ED5309EC2E8F8B38E33C76D5119CB">Four new species of the genus Tumicla Wallengren, 1863 (Lepidoptera: Erebidae Arctiinae: Lithosiini)</mods:title>
</mods:titleInfo>
<mods:name id="795E69AE471C9D94DA85A3D9C466E283" type="personal">
<mods:role id="3A495B43CEEC861B7911F044F63B7ED5">
<mods:roleTerm id="DEF26EE674A5CAEBCC7B695D7B39AA2C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="626337D2B693E4305364523FD5CE16E1">Volynkin, Anton V.</mods:namePart>
</mods:name>
<mods:name id="7567521E7B720CC0C56EAA652544956E" type="personal">
<mods:role id="5D3E851DB6A6DF85188EF2E8D0FC5773">
<mods:roleTerm id="3206B25EA1532D8F24720F6C5205DCD4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5AFB7E66A520FD3E49D1AED41856DF76">László, Gyula M.</mods:namePart>
</mods:name>
<mods:typeOfResource id="0FEA706A407B8D1FC0883C47284DE04F">text</mods:typeOfResource>
<mods:relatedItem id="7865DEA17F942759D4BF3526541BCEE2" type="host">
<mods:titleInfo id="BAA45CFE206A8CDC68A5E4EEE448A251">
<mods:title id="ED1D8C681BAEC19F8560F43B1981A193">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="673218BC17440D9EA343EC4E31428695">
<mods:date id="5E9A6C313AAB18789675CCDF7E2E9E3A">2019</mods:date>
<mods:detail id="18471EECF8A31FC7BA89161AB4148584" type="pubDate">
<mods:number id="350071D1A4C06AA10CC43501BEEA2A72">2019-09-11</mods:number>
</mods:detail>
<mods:detail id="CAEE33F716DA46D4620414A65B55453A" type="volume">
<mods:number id="0EC94A3531780ECBCA84B2EC2E7B9501">4668</mods:number>
</mods:detail>
<mods:detail id="4878B0AEDBADF14182FDFBC04A30C939" type="issue">
<mods:number id="B03D366BE205761287D3DA04E1C12738">3</mods:number>
</mods:detail>
<mods:extent id="9AC8634C3D7905E1FC56618F72F5A9BC" unit="page">
<mods:start id="0BCAF0EC72850C671D433208CDFC8E71">421</mods:start>
<mods:end id="5FEAFF4415D8A7926307B679366203A6">434</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="4023B623073D37D6CD7FB1DB4E4EEEBE">journal article</mods:classification>
<mods:identifier id="7A0254CB9EB1905310279D089C959224" type="CLB-Dataset">25474</mods:identifier>
<mods:identifier id="A93BC942DAC5F7AE7AFFB6C7456B2FB3" type="DOI">10.11646/zootaxa.4668.3.7</mods:identifier>
<mods:identifier id="D891C9319AC0A4C4098E4ADAFCBDA2E9" type="GBIF-Dataset">ddf01fc8-a165-492e-aa9a-25316f7f4ab6</mods:identifier>
<mods:identifier id="61120E571228E493F6D2A15150805453" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="580BC8F2EE93A10592053B882878D60B" type="Zenodo-Dep">3449692</mods:identifier>
<mods:identifier id="CD5592A01BFD9270D57B8085C04CEF28" type="ZooBank">AA9BB1DA-8E27-48E2-9A4A-7428C45F6291</mods:identifier>
</mods:mods>
<treatment id="102387C1FFE6426966F0F976ACC5F882" ID-DOI="http://doi.org/10.5281/zenodo.5941043" ID-GBIF-Taxon="159548969" ID-Zenodo-Dep="5941043" LSID="urn:lsid:zoobank.org:act:231915D5-68F0-45B6-9524-8D0CBF3892C2" httpUri="http://treatment.plazi.org/id/102387C1FFE6426966F0F976ACC5F882" lastPageId="6" lastPageNumber="427" pageId="3" pageNumber="424">
<subSubSection id="D090655CFFE6426C66F0F976ACE5F914" box="[151,464,1726,1753]" pageId="3" pageNumber="424" type="nomenclature">
<paragraph id="983536D7FFE6426C66F0F976ACE5F914" blockId="3.[151,464,1726,1789]" box="[151,464,1726,1753]" pageId="3" pageNumber="424">
<heading id="C37D81BBFFE6426C66F0F976ACE5F914" bold="true" box="[151,464,1726,1753]" fontSize="11" level="1" pageId="3" pageNumber="424" reason="1">
<emphasis id="AAFEEAC5FFE6426C66F0F976ACE5F914" bold="true" box="[151,464,1726,1753]" pageId="3" pageNumber="424">
<taxonomicName id="5F8A4D54FFE6426C66F0F976AC5FF914" ID-CoL="BFXQY" authority="Volynkin &amp; László, 2019" authorityName="Volynkin &amp; László" authorityYear="2019" box="[151,362,1726,1753]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="mbeghai" status="sp. nov.">
<emphasis id="AAFEEAC5FFE6426C66F0F976AC5FF914" bold="true" box="[151,362,1726,1753]" italics="true" pageId="3" pageNumber="424">Tumicla mbeghai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B1CD57BEFFE6426C6715F977ACE5F914" box="[370,464,1727,1753]" pageId="3" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE6426C66F0F92AACA3F930" box="[151,406,1762,1789]" pageId="3" pageNumber="424" type="description">
<paragraph id="983536D7FFE6426C66F0F92AACA3F930" blockId="3.[151,464,1726,1789]" box="[151,406,1762,1789]" pageId="3" pageNumber="424">
(
<figureCitation id="00B12A52FFE6426C66F8F92AAC24F931" box="[159,273,1762,1788]" captionStart="FIGURES 9–18" captionStartId="4.[151,264,1812,1837]" captionTargetBox="[151,1435,181,1763]" captionTargetId="figure@4.[151,1435,181,1787]" captionTargetPageId="4" captionText="FIGURES 9–18. Tumicla spp.: adults. 9, T. mbeghai, paratype ♂, NE Tanzania (©NHMUK); 10, T. mbeghai, paratype ♂, paratype of T. doa, E Tanzania (ZMB); 11, T. mbeghai, holotype ♀, NE Tanzania (©NHMUK); 12, T. similis, holotype ♀, Kenya (©NHMUK); 13, T. similis, ♂, Kenya (MWM/ZSM); 14, T. similis, ♂, Kenya (MWM/ZSM); 15, T. similis, ♀, Kenya (MWM/ZSM); 16, T. rubea, holotype ♂, South Africa (©NHMUK); 17, T. rubea, ♂, South Africa (©NHMUK); 18, T. rubea, paratype ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449700" httpUri="https://zenodo.org/record/3449700/files/figure.png" pageId="3" pageNumber="424">Figs 9–11</figureCitation>
,
<figureCitation id="00B12A52FFE6426C677AF92AAC50F931" box="[285,357,1762,1788]" captionStart="FIGURES 27–30" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[151,1435,181,1875]" captionTargetId="figure@7.[151,1435,181,1894]" captionTargetPageId="7" captionText="FIGURES 27–30. Tumicla spp.: male genitalia. 27, T. elephantina, holotype, Mozambique, slide AV3365 (ANHRT); 28, T. mbeghai, paratype, NE Tanzania, slide NHMUK 010314593 Volynkin (©NHMUK); 29, T. mbeghai, paratype, paratype of T. doa, NE Tanzania, slide 386/2004 Kühne (ZMB); 30, T. tsonga, holotype, Mozambique, slide AV3035 (ANHRT)." figureDoi="http://doi.org/10.5281/zenodo.3449708" httpUri="https://zenodo.org/record/3449708/files/figure.png" pageId="3" pageNumber="424">28, 29</figureCitation>
,
<figureCitation id="00B12A52FFE6426C6716F92AACBBF930" box="[369,398,1762,1789]" captionStart="FIGURES 35–40" captionStartId="9.[151,264,1751,1776]" captionTargetBox="[157,1432,181,1705]" captionTargetId="figure@9.[151,1435,181,1725]" captionTargetPageId="9" captionText="FIGURES 35–40. Tumicla spp.: female genitalia. 35, T. elephantina, paratype, Mozambique, slide AV4399 (ANHRT); 36, T. mbeghai, holotype, NE Tanzania, slide NHMUK 010314594 Volynkin (©NHMUK); 37, T. doa, holotype, Tanzania, slide 388/2004 Kühne (ZMB); 38, T. tsonga, paratype, Mozambique, slide AV3036 (ANHRT); 39, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 40, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NH- MUK)." figureDoi="http://doi.org/10.5281/zenodo.3449716" httpUri="https://zenodo.org/record/3449716/files/figure.png" pageId="3" pageNumber="424">36</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE6426C66F0F8E2AE91F834" pageId="3" pageNumber="424" type="materials_examined">
<paragraph id="983536D7FFE6426C66F0F8E2ACF2F840" blockId="3.[151,1437,1833,2041]" pageId="3" pageNumber="424">
<emphasis id="AAFEEAC5FFE6426C66F0F8E2AC0AF889" bold="true" box="[151,319,1833,1860]" pageId="3" pageNumber="424">
<typeStatus id="47318875FFE6426C66F0F8E2ADE7F889" box="[151,210,1834,1860]" pageId="3" pageNumber="424">Type</typeStatus>
material
</emphasis>
.
<materialsCitation id="28E23C8AFFE6426C6729F8E2ACF6F840" ID-GBIF-Occurrence="2465044340" collectionCode="NHMUK" collectorName="G. Pringle coll. B. M." country="Tanzania" location="Malaria Institute" municipality="Amani" pageId="3" pageNumber="424" specimenCode="NHMUK010915998, NHMUK 010314594" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="AAFEEAC5FFE6426C6729F8E2AC8EF889" bold="true" box="[334,443,1834,1860]" pageId="3" pageNumber="424">
<typeStatus id="47318875FFE6426C6729F8E2AC8EF889" box="[334,443,1834,1860]" pageId="3" pageNumber="424" type="holotype">Holotype</typeStatus>
</emphasis>
(
<figureCitation id="00B12A52FFE6426C67ADF8E2AF14F889" box="[458,545,1834,1860]" captionStart="FIGURES 9–18" captionStartId="4.[151,264,1812,1837]" captionTargetBox="[151,1435,181,1763]" captionTargetId="figure@4.[151,1435,181,1787]" captionTargetPageId="4" captionText="FIGURES 9–18. Tumicla spp.: adults. 9, T. mbeghai, paratype ♂, NE Tanzania (©NHMUK); 10, T. mbeghai, paratype ♂, paratype of T. doa, E Tanzania (ZMB); 11, T. mbeghai, holotype ♀, NE Tanzania (©NHMUK); 12, T. similis, holotype ♀, Kenya (©NHMUK); 13, T. similis, ♂, Kenya (MWM/ZSM); 14, T. similis, ♂, Kenya (MWM/ZSM); 15, T. similis, ♀, Kenya (MWM/ZSM); 16, T. rubea, holotype ♂, South Africa (©NHMUK); 17, T. rubea, ♂, South Africa (©NHMUK); 18, T. rubea, paratype ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449700" httpUri="https://zenodo.org/record/3449700/files/figure.png" pageId="3" pageNumber="424">Figs 11</figureCitation>
,
<figureCitation id="00B12A52FFE6426C644AF8E2AF78F888" box="[557,589,1834,1861]" captionStart="FIGURES 35–40" captionStartId="9.[151,264,1751,1776]" captionTargetBox="[157,1432,181,1705]" captionTargetId="figure@9.[151,1435,181,1725]" captionTargetPageId="9" captionText="FIGURES 35–40. Tumicla spp.: female genitalia. 35, T. elephantina, paratype, Mozambique, slide AV4399 (ANHRT); 36, T. mbeghai, holotype, NE Tanzania, slide NHMUK 010314594 Volynkin (©NHMUK); 37, T. doa, holotype, Tanzania, slide 388/2004 Kühne (ZMB); 38, T. tsonga, paratype, Mozambique, slide AV3036 (ANHRT); 39, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 40, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NH- MUK)." figureDoi="http://doi.org/10.5281/zenodo.3449716" httpUri="https://zenodo.org/record/3449716/files/figure.png" pageId="3" pageNumber="424">36</figureCitation>
):
<specimenCount id="8E8CFD5EFFE6426C6405F8E3AF42F888" box="[610,631,1835,1861]" pageId="3" pageNumber="424" type="female">♀</specimenCount>
, “Amani P 269 4/61” / “
<collectingCountry id="E09D7647FFE6426C65E9F8E2AECFF889" box="[910,1018,1834,1860]" name="Tanzania" pageId="3" pageNumber="424" value="Tanzania">Tanzania</collectingCountry>
:
<collectingMunicipality id="7851ACADFFE6426C6263F8E2A966F889" box="[1028,1107,1834,1860]" pageId="3" pageNumber="424">Amani</collectingMunicipality>
,
<location id="9D55600CFFE6426C6238F8E2A828F889" LSID="urn:lsid:plazi:treatment:102387C1FFE6426966F0F976ACC5F882:9D55600CFFE6426C6238F8E2A828F889" box="[1119,1309,1834,1860]" country="Tanzania" municipality="Amani" name="Malaria Institute" pageId="3" pageNumber="424">Malaria Institute</location>
,
<collectorName id="357F5301FFE6426C634FF8E2AC3CF8A4" pageId="3" pageNumber="424">G. Pringle coll. B.M.</collectorName>
1966-281” / “
<collectorName id="357F5301FFE6426C67CEF886AFA3F8A4" box="[425,662,1870,1897]" pageId="3" pageNumber="424">G. Pringle coll. B.M.</collectorName>
1966-281” / “
<specimenCode id="C82C9EACFFE6426C6553F885A911F8A5" box="[820,1060,1869,1896]" pageId="3" pageNumber="424">NHMUK010915998</specimenCode>
”, slide No.:
<specimenCode id="C82C9EACFFE6426C62CAF887A8A9F8A5" box="[1197,1436,1869,1897]" pageId="3" pageNumber="424">NHMUK 010314594</specimenCode>
Volynkin (Coll.
<collectionCode id="FE9BAE12FFE6426C672BF8BBAC8EF840" box="[332,443,1907,1933]" collectionName="NHMUK" pageId="3" pageNumber="424">NHMUK</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="983536D7FFE6426C66A0F85EAE91F834" blockId="3.[151,1437,1833,2041]" pageId="3" pageNumber="424">
<materialsCitation id="28E23C8AFFE6426C66A0F85EAC27F818" ID-GBIF-Occurrence="2465044333" collectionCode="NHMUK" country="Tanzania" pageId="3" pageNumber="424" specimenCode="NHMUK 010314593" specimenCount="4" specimenCount-female="3" specimenCount-male="1" typeStatus="paratype">
<emphasis id="AAFEEAC5FFE6426C66A0F85EAC0AF87D" bold="true" box="[199,319,1942,1968]" pageId="3" pageNumber="424">
<typeStatus id="47318875FFE6426C66A0F85EAC0AF87D" box="[199,319,1942,1968]" pageId="3" pageNumber="424" type="paratype">Paratypes</typeStatus>
</emphasis>
:
<specimenCount id="8E8CFD5EFFE6426C6736F85EAC4EF87D" box="[337,379,1941,1968]" pageId="3" pageNumber="424" type="male">1 ♂</specimenCount>
,
<specimenCount id="8E8CFD5EFFE6426C67E8F85EAC89F87C" box="[399,444,1942,1969]" pageId="3" pageNumber="424" type="female">3 ♀</specimenCount>
, with same data as the
<typeStatus id="47318875FFE6426C64B5F85EAE02F87D" box="[722,823,1942,1968]" pageId="3" pageNumber="424" type="holotype">holotype</typeStatus>
, slide No.:
<specimenCode id="C82C9EACFFE6426C65D8F85FA985F87D" box="[959,1200,1941,1969]" pageId="3" pageNumber="424">NHMUK 010314593</specimenCode>
(
<specimenCount id="8E8CFD5EFFE6426C62A0F85DA9E0F87D" box="[1223,1237,1941,1968]" pageId="3" pageNumber="424" type="male">♂</specimenCount>
)
<collectingMunicipality id="7851ACADFFE6426C628BF85EA867F87D" box="[1260,1362,1942,1968]" pageId="3" pageNumber="424">Volynkin</collectingMunicipality>
(
<location id="9D55600CFFE6426C6303F85EA8ACF87D" LSID="urn:lsid:plazi:treatment:102387C1FFE6426966F0F976ACC5F882:9D55600CFFE6426C6303F85EA8ACF87D" box="[1380,1433,1942,1968]" pageId="3" pageNumber="424">Coll</location>
.
<collectionCode id="FE9BAE12FFE6426C66F0F873AC33F818" box="[151,262,1979,2005]" collectionName="NHMUK" pageId="3" pageNumber="424">NHMUK</collectionCode>
);
</materialsCitation>
<materialsCitation id="28E23C8AFFE6426C677DF872AE95F834" ID-GBIF-Occurrence="2465044334" collectingDate="2006-02" collectionCode="ZMB" collectorName="Chr. Schroder S. V." country="Tanzania" determinerName="L. Kuhne" location="Amani" municipality="Amani" pageId="3" pageNumber="424" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="8E8CFD5EFFE6426C677DF872AC77F819" box="[282,322,1977,2004]" pageId="3" pageNumber="424" type="male">1 ♂</specimenCount>
, [
<collectingCountry id="E09D7647FFE6426C673CF871ACF3F819" box="[347,454,1977,2004]" name="Tanzania" pageId="3" pageNumber="424" value="Tanzania">Tanzania</collectingCountry>
] D.O. Afrika,
<collectingMunicipality id="7851ACADFFE6426C6400F872AF83F819" box="[615,694,1978,2004]" pageId="3" pageNumber="424">Amani</collectingMunicipality>
,
<date id="EC341017FFE6426C64A4F872AFCFF819" box="[707,762,1978,2004]" pageId="3" pageNumber="424" value="2006-02">
<collectingDate id="FC70E9FFFFE6426C64A4F872AFCFF819" box="[707,762,1978,2004]" pageId="3" pageNumber="424" value="2006-02">II.06</collectingDate>
</date>
,
<collectorName id="357F5301FFE6426C6560F872AEEFF819" box="[775,986,1978,2004]" pageId="3" pageNumber="424">Chr. Schröder S.V.</collectorName>
/ Am. 706 / T 279 /
<typeStatus id="47318875FFE6426C62A1F873A801F818" box="[1222,1332,1979,2005]" pageId="3" pageNumber="424" type="paratype">Paratypus</typeStatus>
<specimenCount id="8E8CFD5EFFE6426C635BF871A864F819" box="[1340,1361,1977,2004]" pageId="3" pageNumber="424" type="male">♂</specimenCount>
<taxonomicName id="5F8A4D54FFE6426C633EF873ADF4F835" authorityName="Kuhne" authorityYear="2007" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="424" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="AAFEEAC5FFE6426C633EF873ADF4F835" italics="true" pageId="3" pageNumber="424">Asura doa</emphasis>
</taxonomicName>
by:
<determinerName id="6DE9DA13FFE6426C6694F817AC68F835" box="[243,349,2014,2041]" pageId="3" pageNumber="424">L. Kühne</determinerName>
/ L. Kühne Dauerpräparat/e 386/2004 (Coll.
<collectionCode id="FE9BAE12FFE6426C6530F817AEADF834" box="[855,920,2015,2041]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="3" pageNumber="424">ZMB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="CCF5665FFFE1426B66F0F8DCAF12F870" ID-DOI="http://doi.org/10.5281/zenodo.3449700" ID-Zenodo-Dep="3449700" httpUri="https://zenodo.org/record/3449700/files/figure.png" pageId="4" pageNumber="425" startId="4.[151,264,1812,1837]" targetBox="[151,1435,181,1763]" targetPageId="4">
<paragraph id="983536D7FFE1426B66F0F8DCAF12F870" blockId="4.[151,1436,1812,1981]" pageId="4" pageNumber="425">
<emphasis id="AAFEEAC5FFE1426B66F0F8DCAC71F8E0" bold="true" box="[151,324,1812,1837]" pageId="4" pageNumber="425">FIGURES 9–18</emphasis>
.
<taxonomicName id="5F8A4D54FFE1426B6734F8DDAC90F8E0" authorityName="Wallengren" authorityYear="1863" box="[339,421,1813,1837]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="genus">
<emphasis id="AAFEEAC5FFE1426B6734F8DDAC90F8E0" box="[339,421,1813,1837]" italics="true" pageId="4" pageNumber="425">Tumicla</emphasis>
</taxonomicName>
spp.: adults. 9,
<taxonomicName id="5F8A4D54FFE1426B6437F8DEAFF7F8E0" authorityName="Volynkin &amp; László" authorityYear="2019" box="[592,706,1813,1837]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE1426B6437F8DEAFF7F8E0" box="[592,706,1813,1837]" italics="true" pageId="4" pageNumber="425">T. mbeghai</emphasis>
</taxonomicName>
, paratype ♂, NE Tanzania (©NHMUK); 10,
<taxonomicName id="5F8A4D54FFE1426B62C7F8DEA826F8E0" authorityName="Volynkin &amp; László" authorityYear="2019" box="[1184,1299,1813,1837]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE1426B62C7F8DEA826F8E0" box="[1184,1299,1813,1837]" italics="true" pageId="4" pageNumber="425">T. mbeghai</emphasis>
</taxonomicName>
, paratype ♂, paratype of
<taxonomicName id="5F8A4D54FFE1426B6771F8F2AC62F89C" authorityName="Kuhne" authorityYear="2007" box="[278,343,1849,1873]" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="AAFEEAC5FFE1426B6771F8F2AC62F89C" box="[278,343,1849,1873]" italics="true" pageId="4" pageNumber="425">T. doa</emphasis>
</taxonomicName>
, E Tanzania (ZMB); 11,
<taxonomicName id="5F8A4D54FFE1426B6405F8F2AFE0F89C" authorityName="Volynkin &amp; László" authorityYear="2019" box="[610,725,1849,1873]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE1426B6405F8F2AFE0F89C" box="[610,725,1849,1873]" italics="true" pageId="4" pageNumber="425">T. mbeghai</emphasis>
</taxonomicName>
, holotype ♀, NE Tanzania (©NHMUK); 12,
<taxonomicName id="5F8A4D54FFE1426B62D3F8F2A827F89C" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[1204,1298,1849,1873]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE1426B62D3F8F2A827F89C" box="[1204,1298,1849,1873]" italics="true" pageId="4" pageNumber="425">T. similis</emphasis>
</taxonomicName>
, holotype ♀, Kenya (©NHMUK); 13,
<taxonomicName id="5F8A4D54FFE1426B67FDF896ACC3F8B8" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[410,502,1885,1909]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE1426B67FDF896ACC3F8B8" box="[410,502,1885,1909]" italics="true" pageId="4" pageNumber="425">T. similis</emphasis>
</taxonomicName>
, ♂, Kenya (MWM/ZSM); 14,
<taxonomicName id="5F8A4D54FFE1426B6552F896AEA4F8B8" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[821,913,1885,1909]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE1426B6552F896AEA4F8B8" box="[821,913,1885,1909]" italics="true" pageId="4" pageNumber="425">T. similis</emphasis>
</taxonomicName>
, ♂, Kenya (MWM/ZSM); 15,
<taxonomicName id="5F8A4D54FFE1426B62A8F896A81EF8B8" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[1231,1323,1885,1909]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE1426B62A8F896A81EF8B8" box="[1231,1323,1885,1909]" italics="true" pageId="4" pageNumber="425">T. similis</emphasis>
</taxonomicName>
, ♀, Kenya (MWM/ZSM); 16,
<taxonomicName id="5F8A4D54FFE1426B673AF84AAC84F854" box="[349,433,1921,1945]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="rubea">
<emphasis id="AAFEEAC5FFE1426B673AF84AAC84F854" box="[349,433,1921,1945]" italics="true" pageId="4" pageNumber="425">T. rubea</emphasis>
</taxonomicName>
, holotype ♂, South Africa (©NHMUK); 17,
<taxonomicName id="5F8A4D54FFE1426B6518F84AAEE6F854" box="[895,979,1921,1945]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="rubea">
<emphasis id="AAFEEAC5FFE1426B6518F84AAEE6F854" box="[895,979,1921,1945]" italics="true" pageId="4" pageNumber="425">T. rubea</emphasis>
</taxonomicName>
, ♂, South Africa (©NHMUK); 18,
<taxonomicName id="5F8A4D54FFE1426B6326F84AA8A3F854" box="[1345,1430,1921,1945]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="425" phylum="Arthropoda" rank="species" species="rubea">
<emphasis id="AAFEEAC5FFE1426B6326F84AA8A3F854" box="[1345,1430,1921,1945]" italics="true" pageId="4" pageNumber="425">T. rubea</emphasis>
</taxonomicName>
, paratype ♀, South Africa (©NHMUK).
</paragraph>
</caption>
<caption id="CCF5665FFFE0426A66F0F8A3A908F83D" ID-DOI="http://doi.org/10.5281/zenodo.3449704" ID-Zenodo-Dep="3449704" httpUri="https://zenodo.org/record/3449704/files/figure.png" pageId="5" pageNumber="426" startId="5.[151,264,1899,1924]" targetBox="[156,1435,181,1860]" targetPageId="5">
<paragraph id="983536D7FFE0426A66F0F8A3A908F83D" blockId="5.[151,1437,1899,2032]" pageId="5" pageNumber="426">
<emphasis id="AAFEEAC5FFE0426A66F0F8A3AC78F849" bold="true" box="[151,333,1899,1924]" pageId="5" pageNumber="426">FIGURES 19–26</emphasis>
.
<taxonomicName id="5F8A4D54FFE0426A673EF8A3AC9EF84E" authorityName="Wallengren" authorityYear="1863" box="[345,427,1899,1923]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="genus">
<emphasis id="AAFEEAC5FFE0426A673EF8A3AC9EF84E" box="[345,427,1899,1923]" italics="true" pageId="5" pageNumber="426">Tumicla</emphasis>
</taxonomicName>
spp.: adults. 19,
<taxonomicName id="5F8A4D54FFE0426A6431F8A4AFEBF84E" authorityName="Volynkin &amp; László" authorityYear="2019" box="[598,734,1899,1923]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="admiranda">
<emphasis id="AAFEEAC5FFE0426A6431F8A4AFEBF84E" box="[598,734,1899,1923]" italics="true" pageId="5" pageNumber="426">T. admiranda</emphasis>
</taxonomicName>
, holotype ♂, DR Congo (©NHMUK); 20,
<taxonomicName id="5F8A4D54FFE0426A62F5F8A4A82CF84E" authorityName="Volynkin &amp; László" authorityYear="2019" box="[1170,1305,1899,1923]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="admiranda">
<emphasis id="AAFEEAC5FFE0426A62F5F8A4A82CF84E" box="[1170,1305,1899,1923]" italics="true" pageId="5" pageNumber="426">T. admiranda</emphasis>
</taxonomicName>
, paratype ♀, DR Congo (©NHMUK); 21,
<taxonomicName id="5F8A4D54FFE0426A67DAF858AF10F86A" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[445,549,1935,1959]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="AAFEEAC5FFE0426A67DAF858AF10F86A" box="[445,549,1935,1959]" italics="true" pageId="5" pageNumber="426">T. fasciata</emphasis>
</taxonomicName>
, holotype ♀, Central African Republic (©NHMUK); 22,
<taxonomicName id="5F8A4D54FFE0426A6204F858A9FEF86A" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[1123,1227,1935,1959]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="fasciata">
<emphasis id="AAFEEAC5FFE0426A6204F858A9FEF86A" box="[1123,1227,1935,1959]" italics="true" pageId="5" pageNumber="426">T. fasciata</emphasis>
</taxonomicName>
, paratype ♀, Central African Republic (©NHMUK); 23,
<taxonomicName id="5F8A4D54FFE0426A6463F87CAF6EF806" authorityName="Volynkin &amp; László" authorityYear="2019" box="[516,603,1971,1995]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="AAFEEAC5FFE0426A6463F87CAF6EF806" box="[516,603,1971,1995]" italics="true" pageId="5" pageNumber="426">T. smithi</emphasis>
</taxonomicName>
, holotype ♀, Zambia (ANHRT); 24,
<taxonomicName id="5F8A4D54FFE0426A65B3F87CA91EF806" authorityName="Volynkin &amp; László" authorityYear="2019" box="[980,1067,1971,1995]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="smithi">
<emphasis id="AAFEEAC5FFE0426A65B3F87CA91EF806" box="[980,1067,1971,1995]" italics="true" pageId="5" pageNumber="426">T. smithi</emphasis>
</taxonomicName>
, paratype ♀, Zambia (ANHRT); 25,
<taxonomicName id="5F8A4D54FFE0426A66F0F810AC20F822" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[151,277,2008,2031]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="sagenaria">
<emphasis id="AAFEEAC5FFE0426A66F0F810AC20F822" box="[151,277,2008,2031]" italics="true" pageId="5" pageNumber="426">T. sagenaria</emphasis>
</taxonomicName>
, ♂, South Africa (©NHMUK); 26,
<taxonomicName id="5F8A4D54FFE0426A64E7F810AFCBF822" baseAuthorityName="Wallengren" baseAuthorityYear="1860" box="[640,766,2008,2031]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="426" phylum="Arthropoda" rank="species" species="sagenaria">
<emphasis id="AAFEEAC5FFE0426A64E7F810AFCBF822" box="[640,766,2008,2031]" italics="true" pageId="5" pageNumber="426">T. sagenaria</emphasis>
</taxonomicName>
, ♀, South Africa (©NHMUK).
</paragraph>
</caption>
<subSubSection id="D090655CFFE3426966A0FF50AF79FC92" pageId="6" pageNumber="427" type="diagnosis">
<paragraph id="983536D7FFE3426966A0FF50AF79FC92" blockId="6.[151,1437,152,1871]" pageId="6" pageNumber="427">
<emphasis id="AAFEEAC5FFE3426966A0FF50AC0FFF7E" bold="true" box="[199,314,152,179]" pageId="6" pageNumber="427">Diagnosis</emphasis>
.
<taxonomicName id="5F8A4D54FFE34269672EFF51AF3EFF7E" authorityName="Volynkin &amp; László" authorityYear="2019" box="[329,523,153,179]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE34269672EFF51AF3EFF7E" box="[329,523,153,179]" italics="true" pageId="6" pageNumber="427">Tumicla mbeghai</emphasis>
</taxonomicName>
is the closest relative of
<taxonomicName id="5F8A4D54FFE342696541FF52AEF3FF7E" authorityName="Volynkin &amp; László" authorityYear="2019" box="[806,966,153,179]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="elephantina">
<emphasis id="AAFEEAC5FFE342696541FF52AEF3FF7E" box="[806,966,153,179]" italics="true" pageId="6" pageNumber="427">T. elephantina</emphasis>
</taxonomicName>
. The differences between the two species are discussed above in the diagnosis of
<taxonomicName id="5F8A4D54FFE34269643DFF76AFCEFF1A" authorityName="Volynkin &amp; László" authorityYear="2019" box="[602,763,189,215]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="elephantina">
<emphasis id="AAFEEAC5FFE34269643DFF76AFCEFF1A" box="[602,763,189,215]" italics="true" pageId="6" pageNumber="427">T. elephantina</emphasis>
</taxonomicName>
. Female of
<taxonomicName id="5F8A4D54FFE3426965E2FF76A934FF1A" authorityName="Volynkin &amp; László" authorityYear="2019" box="[901,1025,189,215]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE3426965E2FF76A934FF1A" box="[901,1025,189,215]" italics="true" pageId="6" pageNumber="427">T. mbeghai</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE342696275FF75A956FF1A" box="[1042,1123,189,215]" captionStart="FIGURES 9–18" captionStartId="4.[151,264,1812,1837]" captionTargetBox="[151,1435,181,1763]" captionTargetId="figure@4.[151,1435,181,1787]" captionTargetPageId="4" captionText="FIGURES 9–18. Tumicla spp.: adults. 9, T. mbeghai, paratype ♂, NE Tanzania (©NHMUK); 10, T. mbeghai, paratype ♂, paratype of T. doa, E Tanzania (ZMB); 11, T. mbeghai, holotype ♀, NE Tanzania (©NHMUK); 12, T. similis, holotype ♀, Kenya (©NHMUK); 13, T. similis, ♂, Kenya (MWM/ZSM); 14, T. similis, ♂, Kenya (MWM/ZSM); 15, T. similis, ♀, Kenya (MWM/ZSM); 16, T. rubea, holotype ♂, South Africa (©NHMUK); 17, T. rubea, ♂, South Africa (©NHMUK); 18, T. rubea, paratype ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449700" httpUri="https://zenodo.org/record/3449700/files/figure.png" pageId="6" pageNumber="427">Fig. 11</figureCitation>
) differs from that of
<taxonomicName id="5F8A4D54FFE342696331FF76A8A9FF1A" authorityName="Kuhne" authorityYear="2007" box="[1366,1436,189,215]" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="AAFEEAC5FFE342696331FF76A8A9FF1A" box="[1366,1436,189,215]" italics="true" pageId="6" pageNumber="427">T. doa</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE3426966C7FF29ADD4FF36" box="[160,225,225,251]" captionStart="FIGURES 1–8" captionStartId="2.[151,264,1911,1936]" captionTargetBox="[164,1419,179,1860]" captionTargetId="figure@2.[164,1423,179,1883]" captionTargetPageId="2" captionText="FIGURES 1–8. Tumicla spp.: adults. 1, T. elephantina, holotype ♂, Mozambique (ANHRT); 2, T. elephantina, paratype ♀, Mo- zambique (ANHRT); 3, T. elephantina, paratype ♀, Mozambique (ANHRT); 4, T. tsonga, holotype ♂, Mozambique (ANHRT); 5, T. tsonga, paratype ♀, Mozambique (ANHRT); 6, T. tsonga, paratype ♀, Mozambique (ANHRT); 7, T. rosea, holotype ♀, Malawi, Nyassaland (©NHMUK); 8, T. doa, holotype ♀, [Tanzania] Nguru Mts. (ZMB)." figureDoi="http://doi.org/10.5281/zenodo.3449696" httpUri="https://zenodo.org/record/3449696/files/figure.png" pageId="6" pageNumber="427">Fig. 8</figureCitation>
) by its larger size, broader forewing with more elongated apex, paler forewing ground colour, finer elements of wing pattern, and hindwing cilia being blackish only at apex (whereas in
<taxonomicName id="5F8A4D54FFE342696598FECEA973FED2" authorityName="Kuhne" authorityYear="2007" box="[1023,1094,261,287]" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="AAFEEAC5FFE342696598FECEA973FED2" box="[1023,1094,261,287]" italics="true" pageId="6" pageNumber="427">T. doa</emphasis>
</taxonomicName>
the cilia is blackish along the entire outer margin). Externally,
<taxonomicName id="5F8A4D54FFE342696460FEE2AFB6FE8E" authorityName="Volynkin &amp; László" authorityYear="2019" box="[519,643,297,323]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE342696460FEE2AFB6FE8E" box="[519,643,297,323]" italics="true" pageId="6" pageNumber="427">T. mbeghai</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE3426964F4FEE1AE31FE8E" box="[659,772,297,323]" captionStart="FIGURES 9–18" captionStartId="4.[151,264,1812,1837]" captionTargetBox="[151,1435,181,1763]" captionTargetId="figure@4.[151,1435,181,1787]" captionTargetPageId="4" captionText="FIGURES 9–18. Tumicla spp.: adults. 9, T. mbeghai, paratype ♂, NE Tanzania (©NHMUK); 10, T. mbeghai, paratype ♂, paratype of T. doa, E Tanzania (ZMB); 11, T. mbeghai, holotype ♀, NE Tanzania (©NHMUK); 12, T. similis, holotype ♀, Kenya (©NHMUK); 13, T. similis, ♂, Kenya (MWM/ZSM); 14, T. similis, ♂, Kenya (MWM/ZSM); 15, T. similis, ♀, Kenya (MWM/ZSM); 16, T. rubea, holotype ♂, South Africa (©NHMUK); 17, T. rubea, ♂, South Africa (©NHMUK); 18, T. rubea, paratype ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449700" httpUri="https://zenodo.org/record/3449700/files/figure.png" pageId="6" pageNumber="427">Figs 9–11</figureCitation>
) is most similar to
<taxonomicName id="5F8A4D54FFE3426965B9FEE2A9CBFE8E" authority="(Durante, 2008)" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[990,1278,297,323]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE3426965B9FEE2A976FE8E" box="[990,1091,297,323]" italics="true" pageId="6" pageNumber="427">T. similis</emphasis>
(
<bibRefCitation id="FC1B4B26FFE342696235FEE1A9C3FE8E" author="Durante, A." box="[1106,1270,297,323]" pageId="6" pageNumber="427" pagination="53 - 68" refId="ref6776" refString="Durante, A. (2008) Asuroides, a new genus of lithosiine moths (Lepidoptera, Arctiidae, Lithosiinae). Zootaxa, 1713, 53 - 68." type="journal article" year="2008">Durante, 2008</bibRefCitation>
)
</taxonomicName>
(
<figureCitation id="00B12A52FFE34269636AFEE1A8BAFE8E" box="[1293,1423,296,323]" captionStart="FIGURES 9–18" captionStartId="4.[151,264,1812,1837]" captionTargetBox="[151,1435,181,1763]" captionTargetId="figure@4.[151,1435,181,1787]" captionTargetPageId="4" captionText="FIGURES 9–18. Tumicla spp.: adults. 9, T. mbeghai, paratype ♂, NE Tanzania (©NHMUK); 10, T. mbeghai, paratype ♂, paratype of T. doa, E Tanzania (ZMB); 11, T. mbeghai, holotype ♀, NE Tanzania (©NHMUK); 12, T. similis, holotype ♀, Kenya (©NHMUK); 13, T. similis, ♂, Kenya (MWM/ZSM); 14, T. similis, ♂, Kenya (MWM/ZSM); 15, T. similis, ♀, Kenya (MWM/ZSM); 16, T. rubea, holotype ♂, South Africa (©NHMUK); 17, T. rubea, ♂, South Africa (©NHMUK); 18, T. rubea, paratype ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449700" httpUri="https://zenodo.org/record/3449700/files/figure.png" pageId="6" pageNumber="427">Figs 12–15</figureCitation>
), from which it differs by its larger size, longer postmedial longitudinal dashes and the absence of blackish dashes at hindwing apex. In the male genitalia,
<taxonomicName id="5F8A4D54FFE34269643DFEBAAFE3FE46" authorityName="Volynkin &amp; László" authorityYear="2019" box="[602,726,369,395]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE34269643DFEBAAFE3FE46" box="[602,726,369,395]" italics="true" pageId="6" pageNumber="427">T. mbeghai</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE342696481FEB9AE53FE46" box="[742,870,369,395]" captionStart="FIGURES 27–30" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[151,1435,181,1875]" captionTargetId="figure@7.[151,1435,181,1894]" captionTargetPageId="7" captionText="FIGURES 27–30. Tumicla spp.: male genitalia. 27, T. elephantina, holotype, Mozambique, slide AV3365 (ANHRT); 28, T. mbeghai, paratype, NE Tanzania, slide NHMUK 010314593 Volynkin (©NHMUK); 29, T. mbeghai, paratype, paratype of T. doa, NE Tanzania, slide 386/2004 Kühne (ZMB); 30, T. tsonga, holotype, Mozambique, slide AV3035 (ANHRT)." figureDoi="http://doi.org/10.5281/zenodo.3449708" httpUri="https://zenodo.org/record/3449708/files/figure.png" pageId="6" pageNumber="427">Figs 28, 29</figureCitation>
) differs clearly from
<taxonomicName id="5F8A4D54FFE342696231FEBAA98FFE46" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[1110,1210,369,395]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE342696231FEBAA98FFE46" box="[1110,1210,369,395]" italics="true" pageId="6" pageNumber="427">T. similis</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE3426962ADFEB9A82EFE46" box="[1226,1307,369,395]" captionStart="FIGURES 31–34" captionStartId="8.[151,264,1808,1833]" captionTargetBox="[155,1418,182,1761]" captionTargetId="figure@8.[151,1435,181,1782]" captionTargetPageId="8" captionText="FIGURES 31–34. Tumicla spp.: male genitalia. 31, T. similis, Kenya, slide MWM 33734 Volynkin (MWM/ZSM); 32, T. rubea, holotype, South Africa, slide BMNH(E) Arct-5868 (©NHMUK); 33, T. admiranda, holotype, DR Congo, slide NHMUK 010314599 Volynkin (©NHMUK); 34, T. sagenaria, South Africa, slide BMNH(E) Arct-5861 (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449712" httpUri="https://zenodo.org/record/3449712/files/figure.png" pageId="6" pageNumber="427">Fig. 31</figureCitation>
) by its narrower uncus, narrower tegumen, wider, U-shaped vinculum (that is V-shaped in
<taxonomicName id="5F8A4D54FFE34269624EFE5EA9BAFE62" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[1065,1167,405,431]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE34269624EFE5EA9BAFE62" box="[1065,1167,405,431]" italics="true" pageId="6" pageNumber="427">T. similis</emphasis>
</taxonomicName>
), more elongated valva apex, much longer aedeagus with a sclerotised carina, and broader vesica with a robust cornutus (whereas the vesica of
<taxonomicName id="5F8A4D54FFE3426966D1FE16AC29FE3A" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[182,284,477,503]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE3426966D1FE16AC29FE3A" box="[182,284,477,503]" italics="true" pageId="6" pageNumber="427">T. similis</emphasis>
</taxonomicName>
is narrow, lacking any sclerotised elements). The female genitalia of
<taxonomicName id="5F8A4D54FFE342696257FE16A998FE3A" authorityName="Volynkin &amp; László" authorityYear="2019" box="[1072,1197,477,503]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE342696257FE16A998FE3A" box="[1072,1197,477,503]" italics="true" pageId="6" pageNumber="427">T. mbeghai</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE3426962D9FE15A825FE35" box="[1214,1296,477,504]" captionStart="FIGURES 35–40" captionStartId="9.[151,264,1751,1776]" captionTargetBox="[157,1432,181,1705]" captionTargetId="figure@9.[151,1435,181,1725]" captionTargetPageId="9" captionText="FIGURES 35–40. Tumicla spp.: female genitalia. 35, T. elephantina, paratype, Mozambique, slide AV4399 (ANHRT); 36, T. mbeghai, holotype, NE Tanzania, slide NHMUK 010314594 Volynkin (©NHMUK); 37, T. doa, holotype, Tanzania, slide 388/2004 Kühne (ZMB); 38, T. tsonga, paratype, Mozambique, slide AV3036 (ANHRT); 39, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 40, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NH- MUK)." figureDoi="http://doi.org/10.5281/zenodo.3449716" httpUri="https://zenodo.org/record/3449716/files/figure.png" pageId="6" pageNumber="427">Fig. 36</figureCitation>
) differ from those of
<taxonomicName id="5F8A4D54FFE342696690FDCAAC09FDD6" authorityName="Kuhne" authorityYear="2007" box="[247,316,513,539]" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="AAFEEAC5FFE342696690FDCAAC09FDD6" box="[247,316,513,539]" italics="true" pageId="6" pageNumber="427">T. doa</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE34269672CFDC9ACA9FDD6" box="[331,412,513,539]" captionStart="FIGURES 35–40" captionStartId="9.[151,264,1751,1776]" captionTargetBox="[157,1432,181,1705]" captionTargetId="figure@9.[151,1435,181,1725]" captionTargetPageId="9" captionText="FIGURES 35–40. Tumicla spp.: female genitalia. 35, T. elephantina, paratype, Mozambique, slide AV4399 (ANHRT); 36, T. mbeghai, holotype, NE Tanzania, slide NHMUK 010314594 Volynkin (©NHMUK); 37, T. doa, holotype, Tanzania, slide 388/2004 Kühne (ZMB); 38, T. tsonga, paratype, Mozambique, slide AV3036 (ANHRT); 39, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 40, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NH- MUK)." figureDoi="http://doi.org/10.5281/zenodo.3449716" httpUri="https://zenodo.org/record/3449716/files/figure.png" pageId="6" pageNumber="427">Fig. 37</figureCitation>
) by their broader ostium bursae, broader and more heavily sclerotised ductus bursae having an anterio-lateral sclerotised protrusion to the appendix bursae, and the corpus bursae structure. In
<taxonomicName id="5F8A4D54FFE342696292FDEEA847FDF2" authorityName="Volynkin &amp; László" authorityYear="2019" box="[1269,1394,549,575]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE342696292FDEEA847FDF2" box="[1269,1394,549,575]" italics="true" pageId="6" pageNumber="427">T. mbeghai</emphasis>
</taxonomicName>
the corpus bursae is subdivided into two sections, the anterior one is globular and membranous with weak scobination, while the posterior one is dilated, dorso-ventrally flattened having conspicuous lateral protrusions, whereas in
<taxonomicName id="5F8A4D54FFE34269633FFDA6A8A9FD4A" authorityName="Kuhne" authorityYear="2007" box="[1368,1436,621,647]" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="AAFEEAC5FFE34269633FFDA6A8A9FD4A" box="[1368,1436,621,647]" italics="true" pageId="6" pageNumber="427">T. doa</emphasis>
</taxonomicName>
the corpus bursae is more or less pear-like with its anterior broadened section heavily sclerotised. In addition, the appendix bursae of
<taxonomicName id="5F8A4D54FFE342696712FD7EACC4FD02" authorityName="Volynkin &amp; László" authorityYear="2019" box="[373,497,693,719]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE342696712FD7EACC4FD02" box="[373,497,693,719]" italics="true" pageId="6" pageNumber="427">T. mbeghai</emphasis>
</taxonomicName>
is small, conical, heavily sclerotised, situated postero-laterally at left side, whereas in
<taxonomicName id="5F8A4D54FFE3426966D3FD12ADCFFD3E" authorityName="Kuhne" authorityYear="2007" box="[180,250,729,755]" class="Insecta" family="Arctiidae" genus="Asura" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="doa">
<emphasis id="AAFEEAC5FFE3426966D3FD12ADCFFD3E" box="[180,250,729,755]" italics="true" pageId="6" pageNumber="427">T. doa</emphasis>
</taxonomicName>
that is broadly conical, more weakly sclerotised, rugose and situated laterally at right side. In comparison with those of
<taxonomicName id="5F8A4D54FFE342696757FD36ACA1FCDA" baseAuthorityName="Durante" baseAuthorityYear="2008" box="[304,404,765,791]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="similis">
<emphasis id="AAFEEAC5FFE342696757FD36ACA1FCDA" box="[304,404,765,791]" italics="true" pageId="6" pageNumber="427">T. similis</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE3426967C3FD35ACC2FCDA" box="[420,503,765,791]" captionStart="FIGURES 41–45" captionStartId="11.[151,264,1898,1923]" captionTargetBox="[189,1394,242,1857]" captionTargetId="figure@11.[189,1398,237,1874]" captionTargetPageId="11" captionText="FIGURES 41–45. Tumicla spp.: female genitalia. 41, T. similis, paratype, Kenya, Kibwezi, slide NHMUK010314047 Volynkin (©NHMUK); 42, T. admiranda, paratype, DR Congo, slide NHMUK 010314600 Volynkin (©NHMUK); 43, T. fasciata, paratype, CentralAfrican Republic, slide NHMUK010314046 Volynkin (©NHMUK); 44, T. smithi, holotype, Zambia, slideAV5243 (ANHRT); 45, T. sagenaria, lectotype, [South Africa] Caffraria, slide NHRS 000011009 (photo by T. Malm, ©2017NHRS)." figureDoi="http://doi.org/10.5281/zenodo.3449718" httpUri="https://zenodo.org/record/3449718/files/figure.png" pageId="6" pageNumber="427">Fig. 41</figureCitation>
), the female genitalia of
<taxonomicName id="5F8A4D54FFE34269656AFD36AEBDFCDA" authorityName="Volynkin &amp; László" authorityYear="2019" box="[781,904,765,791]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE34269656AFD36AEBDFCDA" box="[781,904,765,791]" italics="true" pageId="6" pageNumber="427">T. mbeghai</emphasis>
</taxonomicName>
(
<figureCitation id="00B12A52FFE3426965FFFD35AEDCFCD5" box="[920,1001,765,792]" captionStart="FIGURES 35–40" captionStartId="9.[151,264,1751,1776]" captionTargetBox="[157,1432,181,1705]" captionTargetId="figure@9.[151,1435,181,1725]" captionTargetPageId="9" captionText="FIGURES 35–40. Tumicla spp.: female genitalia. 35, T. elephantina, paratype, Mozambique, slide AV4399 (ANHRT); 36, T. mbeghai, holotype, NE Tanzania, slide NHMUK 010314594 Volynkin (©NHMUK); 37, T. doa, holotype, Tanzania, slide 388/2004 Kühne (ZMB); 38, T. tsonga, paratype, Mozambique, slide AV3036 (ANHRT); 39, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 40, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NH- MUK)." figureDoi="http://doi.org/10.5281/zenodo.3449716" httpUri="https://zenodo.org/record/3449716/files/figure.png" pageId="6" pageNumber="427">Fig. 36</figureCitation>
) have longer and much broader ductus bursae, much broader and sclerotised posterior section of corpus bursae having sclerotised lateral protrusions, and larger and sclerotised appendix bursae.
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE3426966A0FCA0A952F972" pageId="6" pageNumber="427" type="description">
<paragraph id="983536D7FFE3426966A0FCA0A952F972" blockId="6.[151,1437,152,1871]" pageId="6" pageNumber="427">
<emphasis id="AAFEEAC5FFE3426966A0FCA0AC64FC4E" bold="true" box="[199,337,872,899]" pageId="6" pageNumber="427">Description</emphasis>
.
<emphasis id="AAFEEAC5FFE342696738FCA1AFFEFC4E" bold="true" box="[351,715,872,899]" pageId="6" pageNumber="427">External morphology of adults</emphasis>
(
<figureCitation id="00B12A52FFE3426964BEFCA1AE79FC4E" box="[729,844,873,899]" captionStart="FIGURES 9–18" captionStartId="4.[151,264,1812,1837]" captionTargetBox="[151,1435,181,1763]" captionTargetId="figure@4.[151,1435,181,1787]" captionTargetPageId="4" captionText="FIGURES 9–18. Tumicla spp.: adults. 9, T. mbeghai, paratype ♂, NE Tanzania (©NHMUK); 10, T. mbeghai, paratype ♂, paratype of T. doa, E Tanzania (ZMB); 11, T. mbeghai, holotype ♀, NE Tanzania (©NHMUK); 12, T. similis, holotype ♀, Kenya (©NHMUK); 13, T. similis, ♂, Kenya (MWM/ZSM); 14, T. similis, ♂, Kenya (MWM/ZSM); 15, T. similis, ♀, Kenya (MWM/ZSM); 16, T. rubea, holotype ♂, South Africa (©NHMUK); 17, T. rubea, ♂, South Africa (©NHMUK); 18, T. rubea, paratype ♀, South Africa (©NHMUK)." figureDoi="http://doi.org/10.5281/zenodo.3449700" httpUri="https://zenodo.org/record/3449700/files/figure.png" pageId="6" pageNumber="427">Figs 9–11</figureCitation>
). Forewing length 12–12.5 mm in males and 16–16.5 mm in females. Male antennae ciliate, female antennae filiform. Head pale mustard yellow with blackish dot on frons. Thorax pale mustard yellow, with four black spots medio-laterally and one dot medio-posteriorly. Forewing moderately broad, broader in female, with rounded apex and oblique tornus; forewing ground colour pale mustard yellow; wing pattern blackish, consists of one large subbasal spot, one short longitudinal subbasal dash, zigzagged ante-, postmedial and medial lines connecting each other, semilunar discal spot, and interrupted subterminal line consisting of long dashes of various length connected to postmedial line; cilia blackish. Hindwing pale mustard yellow, lighter than forewing; cilia as wing ground colour, sometimes with admixture of blackish scales at apex. Abdomen pale ochreous.
<emphasis id="AAFEEAC5FFE3426967DEFB41AF50FB6E" bold="true" box="[441,613,1160,1187]" pageId="6" pageNumber="427">Male genitalia</emphasis>
(
<figureCitation id="00B12A52FFE342696412FB41AFCFFB6E" box="[629,762,1161,1187]" captionStart="FIGURES 27–30" captionStartId="7.[151,264,1920,1945]" captionTargetBox="[151,1435,181,1875]" captionTargetId="figure@7.[151,1435,181,1894]" captionTargetPageId="7" captionText="FIGURES 27–30. Tumicla spp.: male genitalia. 27, T. elephantina, holotype, Mozambique, slide AV3365 (ANHRT); 28, T. mbeghai, paratype, NE Tanzania, slide NHMUK 010314593 Volynkin (©NHMUK); 29, T. mbeghai, paratype, paratype of T. doa, NE Tanzania, slide 386/2004 Kühne (ZMB); 30, T. tsonga, holotype, Mozambique, slide AV3035 (ANHRT)." figureDoi="http://doi.org/10.5281/zenodo.3449708" httpUri="https://zenodo.org/record/3449708/files/figure.png" pageId="6" pageNumber="427">Figs 28, 29</figureCitation>
). Uncus moderately long, narrow, distally hook-like curved and slightly broadened, apically pointed; tuba analis broad, scaphium thin, weakly sclerotised, subscaphium represented by a field of fine spinulose scobination; tegumen moderately long and broad; juxta broad, shield-like; vinculum robust, U-shaped; valva elongated, medially broadened, its distal half slightly curved ventrally, apically strongly tapered, with pointed tip; costal margin convex, with short irregular dentation on its distal third; sacculus short, narrow, weakly setose, without processes, with slightly convex ventral margin. Aedeagus short, straight; carina elongated ventrally, heavily sclerotised. Vesica broad, directed ventrad, membranous, with one small subapical diverticulum and one long and robust, spine-like subbasal cornutus situated on base connected to carina by a narrow sclerotised band.
<emphasis id="AAFEEAC5FFE34269673DFA61AF2AFA0E" bold="true" box="[346,543,1448,1475]" pageId="6" pageNumber="427">Female genitalia</emphasis>
(
<figureCitation id="00B12A52FFE342696449FA61AFB7FA09" box="[558,642,1449,1476]" captionStart="FIGURES 35–40" captionStartId="9.[151,264,1751,1776]" captionTargetBox="[157,1432,181,1705]" captionTargetId="figure@9.[151,1435,181,1725]" captionTargetPageId="9" captionText="FIGURES 35–40. Tumicla spp.: female genitalia. 35, T. elephantina, paratype, Mozambique, slide AV4399 (ANHRT); 36, T. mbeghai, holotype, NE Tanzania, slide NHMUK 010314594 Volynkin (©NHMUK); 37, T. doa, holotype, Tanzania, slide 388/2004 Kühne (ZMB); 38, T. tsonga, paratype, Mozambique, slide AV3036 (ANHRT); 39, T. rosea, holotype, Malawi, Nyassaland, slide BMNH(E) Arct-5855 (©NHMUK); 40, T. rubea, paratype, South Africa, slide BMNH(E) Arct-5869 (©NH- MUK)." figureDoi="http://doi.org/10.5281/zenodo.3449716" httpUri="https://zenodo.org/record/3449716/files/figure.png" pageId="6" pageNumber="427">Fig. 36</figureCitation>
). Ovipositor short and broad; papillae anales rectangular, with rounded angles; apophyses moderately long and thin; apophyses posteriores slightly longer and narrower than apophyses anteriores. Ostium bursae broad with slightly arched margin; ductus bursae broad, dorso-ventrally flattened, slightly rugose medially, heavily sclerotised, it’s anterior left corner heavily sclerotised and protruding to posterior margin of appendix bursae. Posterior section of corpus bursae elongated, very broad, dorso-ventrally flattened, moderately sclerotised, weakly rugose, with trigonal and heavily sclerotised right protrusion latero-anteriorly, and broad but short semi-globular left protrusion laterally. Anterior section of corpus bursae globular, weakly setose. Appendix bursae short, broadly trigonal, heavily sclerotised, situated postero-laterally at left side.
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE3426966A0F900AC56F8C5" pageId="6" pageNumber="427" type="distribution">
<paragraph id="983536D7FFE3426966A0F900AC56F8C5" blockId="6.[151,1437,152,1871]" pageId="6" pageNumber="427">
<emphasis id="AAFEEAC5FFE3426966A0F900AC6CF92E" bold="true" box="[199,345,1736,1763]" pageId="6" pageNumber="427">Distribution</emphasis>
.
<taxonomicName id="5F8A4D54FFE342696700F901AF1DF92E" authorityName="Volynkin &amp; László" authorityYear="2019" box="[359,552,1737,1763]" class="Insecta" family="Erebidae" genus="Tumicla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="427" phylum="Arthropoda" rank="species" species="mbeghai">
<emphasis id="AAFEEAC5FFE342696700F901AF1DF92E" box="[359,552,1737,1763]" italics="true" pageId="6" pageNumber="427">Tumicla mbeghai</emphasis>
</taxonomicName>
is only known to date from its
<typeStatus id="47318875FFE3426965E5F901AE87F92E" box="[898,946,1737,1763]" pageId="6" pageNumber="427">type</typeStatus>
locality in the Usambara Mountains (northeastern Tanzania).
</paragraph>
</subSubSection>
<subSubSection id="D090655CFFE3426966A0F8D9ACC5F882" pageId="6" pageNumber="427" type="etymology">
<paragraph id="983536D7FFE3426966A0F8D9ACC5F882" blockId="6.[151,1437,152,1871]" pageId="6" pageNumber="427">
<emphasis id="AAFEEAC5FFE3426966A0F8D9AC7DF8E6" bold="true" box="[199,328,1809,1835]" pageId="6" pageNumber="427">Etymology</emphasis>
. Mbegha is a mythic hero of the Shambaa people inhabiting the Usambara Mountains, considered founder of the Kilindi dynasty.
</paragraph>
</subSubSection>
</treatment>
</document>
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