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<document id="ABE8023E6237A3161D5B10A241E785E9" ID-CLB-Dataset="276817" ID-DOI="10.11646/zootaxa.5178.2.3" ID-GBIF-Dataset="72f5400a-d642-4d8e-94bf-e82318daf384" ID-ISSN="1175-5326" ID-Zenodo-Dep="7022237" ID-ZooBank="B466BB45-91F1-456E-971A-E3FAA38CBCA9" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1661413289198" checkinUser="plazi" docAuthor="Zamani, Alireza &amp; Marusik, Yuri M." docDate="2022" docId="76698788FFA3E31BFF6CFF7845772B73" docLanguage="en" docName="zootaxa.5178.2.3.pdf" docOrigin="Zootaxa 5178 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Zodariellum turanicum Zamani &amp; Marusik 2022, sp. n." docType="treatment" docVersion="4" lastPageNumber="175" masterDocId="8A50FFF0FFADE315FFFBFFEF40132F46" masterDocTitle="New taxonomic considerations in Zodariellum Andreeva &amp; Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera" masterLastPageNumber="177" masterPageNumber="161" pageNumber="175" updateTime="1699377370295" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F25E0BD6DA06ADC6614047CDF738E580">New taxonomic considerations in Zodariellum Andreeva &amp; Tyshchenko, 1968 (Araneae: Zodariidae), with notes on the presence of cymbial diverticulum in different zodariid genera</mods:title>
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<mods:namePart id="E493D88E788ECFE918D5480847BA4786">Zamani, Alireza</mods:namePart>
<mods:affiliation id="A48B3A5C2CFA7FAE177113D7C1D9C794">Zoological Museum, Biodiversity Unit, University of Turku, FI- 20014, Finland.</mods:affiliation>
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<mods:namePart id="4AD482529009BD29B4413C62646337F9">Marusik, Yuri M.</mods:namePart>
<mods:affiliation id="2F4075D04BC01C5F722B0783F09D0AE6">Institute for Biological Problems of the North RAS, Portovaya Str. 18, Magadan, Russia. &amp; Department of Zoology &amp; Entomology, University of the Free State, Bloemfontein 9300, South Africa.</mods:affiliation>
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<mods:title id="B0AF81A3D504A4A01A963CF409A0FCD0">Zootaxa</mods:title>
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<mods:date id="CF13CDCF09CA8226507D715A7EF73F2F">2022</mods:date>
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<mods:number id="FDCA7D98E13B0F3A99F51813049D0928">2022-08-25</mods:number>
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<treatment id="76698788FFA3E31BFF6CFF7845772B73" ID-GBIF-Taxon="217308878" LSID="urn:lsid:plazi:treatment:76698788FFA3E31BFF6CFF7845772B73" httpUri="http://treatment.plazi.org/id/76698788FFA3E31BFF6CFF7845772B73" lastPageNumber="175" pageId="14" pageNumber="175">
<subSubSection id="B6DA6515FFA3E31BFF6CFF7841EE2FF7" box="[151,509,151,178]" pageId="14" pageNumber="175" type="nomenclature">
<paragraph id="FE7F369EFFA3E31BFF6CFF7841EE2FF7" blockId="14.[151,590,151,214]" box="[151,509,151,178]" pageId="14" pageNumber="175">
<heading id="A53781F2FFA3E31BFF6CFF7841EE2FF7" bold="true" box="[151,509,151,178]" fontSize="11" level="1" pageId="14" pageNumber="175" reason="1">
<emphasis id="CCB4EA8CFFA3E31BFF6CFF7841EE2FF7" bold="true" box="[151,509,151,178]" pageId="14" pageNumber="175">
<taxonomicName id="39C04D1DFFA3E31BFF6CFF7841A02FF7" authority="Zamani &amp; Marusik, 2022" authorityName="Zamani &amp; Marusik" authorityYear="2022" box="[151,435,151,178]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="175" phylum="Arthropoda" rank="species" species="turanicum" status="sp. nov.">
<emphasis id="CCB4EA8CFFA3E31BFF6CFF7841A02FF7" bold="true" box="[151,435,151,178]" italics="true" pageId="14" pageNumber="175">Zodariellum turanicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D78757F7FFA3E31BFE40FF7841EE2FF7" box="[443,509,151,177]" pageId="14" pageNumber="175" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="B6DA6515FFA3E31BFF6CFF54425D2F93" box="[151,590,187,214]" pageId="14" pageNumber="175" type="description">
<paragraph id="FE7F369EFFA3E31BFF6CFF54425D2F93" blockId="14.[151,590,151,214]" box="[151,590,187,214]" pageId="14" pageNumber="175">
<figureCitation id="66FB2A1BFFA3E31BFF6CFF5440E32F93" box="[151,240,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1977]" captionTargetBox="[151,1436,605,1926]" captionTargetId="figure-210@2.[151,1436,594,1926]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Zodariellum asiaticum (A), Z. bactrianum (B), Z. turanicum sp. n. (C), Z. nenilini (DE) and Z. turkmenicum sp. n. (F), dorsal view. AD, F males; E female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7022241" httpUri="https://zenodo.org/record/7022241/files/figure.png" pageId="14" pageNumber="175">Figs 1C</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFF06FF5441312F93" box="[253,290,187,213]" captionStart="FIGURE 2" captionStartId="3.[151,250,1852,1876]" captionTargetBox="[191,1400,424,1829]" captionTargetId="figure-65@3.[173,1414,412,1829]" captionTargetPageId="3" captionText="FIGURE 2. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B, F), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E). AE ventral view; F dorsal view of cymbium, showing cymbial diverticulum. Scale bars = 0.2 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Eb—base of embolus, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branch(es) of retrolateral tibial apophysis, Sp—spur of tegular apophysis, Tg—tegular apophysis, To—triangular outgrowth, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022243" httpUri="https://zenodo.org/record/7022243/files/figure.png" pageId="14" pageNumber="175">2D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFED6FF5441412F93" box="[301,338,187,213]" captionStart="FIGURE 3" captionStartId="4.[151,250,1555,1579]" captionTargetBox="[152,1436,181,1531]" captionTargetId="figure-21@4.[152,1436,181,1531]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of the male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), ventral view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Eb—base of embolus, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Sp—spur of tegular apophysis, Tg—tegular apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022245" httpUri="https://zenodo.org/record/7022245/files/figure.png" pageId="14" pageNumber="175">3D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFEA5FF5441912F93" box="[350,386,187,213]" captionStart="FIGURE 4" captionStartId="5.[151,250,1907,1931]" captionTargetBox="[192,1383,207,1870]" captionTargetId="figure-17@5.[175,1412,181,1884]" captionTargetPageId="5" captionText="FIGURE 4. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), retrolateral view. Scale bars = 0.2 mm. Abbreviations: Ap—anterodorsal part of retrolateral tibial apophysis, Cn—conductor, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022249" httpUri="https://zenodo.org/record/7022249/files/figure.png" pageId="14" pageNumber="175">4D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFE75FF5441A02F90" box="[398,435,187,214]" captionStart="FIGURE 5" captionStartId="6.[151,250,1584,1608]" captionTargetBox="[177,1411,181,1560]" captionTargetId="figure-21@6.[177,1411,181,1560]" captionTargetPageId="6" captionText="FIGURE 5. SEM images of the male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), retrolateral view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Di—diverticulum, Eb—base of embolus, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022251" httpUri="https://zenodo.org/record/7022251/files/figure.png" pageId="14" pageNumber="175">5D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFE45FF5441F12F93" box="[446,482,187,213]" captionStart="FIGURE 6" captionStartId="7.[151,250,1878,1902]" captionTargetBox="[192,1423,460,1840]" captionTargetId="figure-119@7.[151,1436,450,1840]" captionTargetPageId="7" captionText="FIGURE 6. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), dorsal view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7022257" httpUri="https://zenodo.org/record/7022257/files/figure.png" pageId="14" pageNumber="175">6D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFE15FF5442012F93" box="[494,530,187,213]" captionStart="FIGURE 9" captionStartId="10.[151,250,1757,1781]" captionTargetBox="[153,1434,345,1733]" captionTargetId="figure-84@10.[153,1434,345,1733]" captionTargetPageId="10" captionText="FIGURE 9. SEM images of the male palps of Zodariellum asiaticum (A), Z. bactrianum (BC) and Z. turanicum sp. n. (D). A anteroretrolateral view; B ventroretrolateral view; C anteroventral view; D anterior view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Eb—base of embolus, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr— retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022263" httpUri="https://zenodo.org/record/7022263/files/figure.png" pageId="14" pageNumber="175">9D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFDE5FF54425D2F93" box="[542,590,187,213]" captionStart="FIGURE 12" captionStartId="12.[151,250,1866,1890]" captionTargetBox="[164,1424,453,1842]" captionTargetId="figure-144@12.[164,1424,453,1842]" captionTargetPageId="12" captionText="FIGURE 12. Distribution records of Zodariellum spp. A: 1) Z. asiaticum, 2) Z. bactrianum, 3) Z. bekuzini, 4) Z. continentale, 5) Z. proszynskii, 6) Z. spasskyi and Z. zebra, 7) Z. surprisum, 8) Z. sytchevskajae, 9) Z. turanicum sp. n., 10) Z. turkmenicum sp. n., 11) Z. tadzhikum, 12) Z. testaceofasciatum, 13) Z. volgouralense, 14) Z. nenilini (in part, excluding records from Mongolia). B: 1) Z. nenilini (in part, records from Mongolia), 2) Z. schmidti, 3) Z. mongolicum, 4) Z. chaoyangense, 5) Z. furcum, 6) Z. hunanense, 7) Z. planum. A single point may refer to several closely located records." figureDoi="http://doi.org/10.5281/zenodo.7022273" httpUri="https://zenodo.org/record/7022273/files/figure.png" pageId="14" pageNumber="175">12A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="B6DA6515FFA3E31BFF6CFEEC449D2E07" pageId="14" pageNumber="175" type="materials_examined">
<paragraph id="FE7F369EFFA3E31BFF6CFEEC449D2E07" blockId="14.[151,1437,259,1078]" pageId="14" pageNumber="175">
<emphasis id="CCB4EA8CFFA3E31BFF6CFEEC41552E5B" bold="true" box="[151,326,259,285]" pageId="14" pageNumber="175">Type material.</emphasis>
<materialsCitation id="4EA83CC3FFA3E31BFEB5FEEC449D2E07" ID-GBIF-Occurrence="4435949301" collectingDate="1985-05-20" collectionCode="ZMMU" collectorName="A. V. Tanasevitch" country="Turkmenistan" elevation="1100" latitude="37.933334" location="Khodzhapil" longLatPrecision="1178" longitude="66.61667" pageId="14" pageNumber="175" specimenCount="1" specimenCount-male="1" stateProvince="Lebap Welayaty" typeStatus="holotype">
<typeStatus id="217B883CFFA3E31BFEB5FEEC41A52E5B" box="[334,438,259,285]" pageId="14" pageNumber="175" type="holotype">Holotype</typeStatus>
<specimenCount id="E8C6FD17FFA3E31BFE44FEEB41C72E58" box="[447,468,260,286]" pageId="14" pageNumber="175" type="male"></specimenCount>
(
<collectionCode id="98D1AE5BFFA3E31BFE1FFEEC42522E5B" box="[484,577,259,285]" country="Russia" name="Zoological Museum, Moscow Lomonosov State University" pageId="14" pageNumber="175" type="Museum">ZMMU</collectionCode>
),
<collectingCountry id="86D7760EFFA3E31BFDAEFEEC43272E5B" box="[597,820,259,285]" name="Turkmenistan" pageId="14" pageNumber="175">TURKMENISTAN</collectingCountry>
:
<emphasis id="CCB4EA8CFFA3E31BFCBBFEEC43CE2E5B" box="[832,989,259,285]" italics="true" pageId="14" pageNumber="175">
<collectingRegion id="3C04F87CFFA3E31BFCBBFEEC43CE2E5B" box="[832,989,259,285]" country="Turkmenistan" name="Lebap Welayaty" pageId="14" pageNumber="175">Lebap Region</collectingRegion>
</emphasis>
: Kugitang-Tau Mnt. range, environs of
<location id="FB1F6045FFA3E31BFF6CFEC8410E2E07" LSID="urn:lsid:plazi:treatment:76698788FFA3E31BFF6CFF7845772B73:FB1F6045FFA3E31BFF6CFEC8410E2E07" box="[151,285,295,321]" country="Turkmenistan" latitude="37.933334" longLatPrecision="1178" longitude="66.61667" name="Khodzhapil" pageId="14" pageNumber="175" stateProvince="Lebap Welayaty">Khodzhapil</location>
, [
<geoCoordinate id="9BF45059FFA3E31BFED4FEC8419D2E04" box="[303,398,295,322]" degrees="37" direction="north" minutes="56" orientation="latitude" pageId="14" pageNumber="175" precision="925" value="37.933334">37°56'N</geoCoordinate>
<geoCoordinate id="9BF45059FFA3E31BFE6EFEC841E02E07" box="[405,499,295,321]" degrees="66" direction="east" minutes="37" orientation="longitude" pageId="14" pageNumber="175" precision="925" value="66.61667">66°37'E</geoCoordinate>
], ca
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<elevation id="75EDD1ADFFA3E31BFDDEFEC842682E04" box="[549,635,295,322]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="14" pageNumber="175" unit="m" value="1100.0">1100 m</elevation>
</quantity>
, on the ground,
<date id="8A7E105EFFA3E31BFCCBFEC843BD2E04" box="[816,942,295,322]" pageId="14" pageNumber="175" value="1985-05-20">
<collectingDate id="9A3AE9B6FFA3E31BFCCBFEC843BD2E04" box="[816,942,295,322]" pageId="14" pageNumber="175" value="1985-05-20">20.05.1985</collectingDate>
</date>
(
<collectorName id="53355348FFA3E31BFC45FEC844902E07" box="[958,1155,295,321]" pageId="14" pageNumber="175">A.V. Tanasevitch</collectorName>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="B6DA6515FFA3E31BFF3CFEA4447F2E23" box="[199,1132,331,358]" pageId="14" pageNumber="175" type="etymology">
<paragraph id="FE7F369EFFA3E31BFF3CFEA4447F2E23" blockId="14.[151,1437,259,1078]" box="[199,1132,331,358]" pageId="14" pageNumber="175">
<emphasis id="CCB4EA8CFFA3E31BFF3CFEA4415D2E23" bold="true" box="[199,334,331,357]" pageId="14" pageNumber="175">Etymology.</emphasis>
The specific epithet refers to Turan, a historical region in Central Asia.
</paragraph>
</subSubSection>
<subSubSection id="B6DA6515FFA3E31BFF3CFE8041712DC3" pageId="14" pageNumber="175" type="diagnosis">
<paragraph id="FE7F369EFFA3E31BFF3CFE8041712DC3" blockId="14.[151,1437,259,1078]" pageId="14" pageNumber="175">
<emphasis id="CCB4EA8CFFA3E31BFF3CFE8041512ECF" bold="true" box="[199,322,367,393]" pageId="14" pageNumber="175">Diagnosis.</emphasis>
The new species has abdominal pattern similar to those of
<taxonomicName id="39C04D1DFFA3E31BFC1DFE9F44502ECF" authorityName="Zamani &amp; Marusik" authorityYear="2022" baseAuthorityName="Charitonov" baseAuthorityYear="1946" box="[998,1091,367,393]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="175" phylum="Arthropoda" rank="species" species="zebra">
<emphasis id="CCB4EA8CFFA3E31BFC1DFE9F44502ECF" box="[998,1091,367,393]" italics="true" pageId="14" pageNumber="175">Z. zebra</emphasis>
</taxonomicName>
and
<taxonomicName id="39C04D1DFFA3E31BFB87FE9F45702ECF" authorityName=": Brignoli" authorityYear="1983" box="[1148,1379,367,393]" class="Arachnida" family="Zodariidae" genus="Zodarion" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="175" phylum="Arthropoda" rank="species" species="testaceofasciatum">
<emphasis id="CCB4EA8CFFA3E31BFB87FE9F45702ECF" box="[1148,1379,367,393]" italics="true" pageId="14" pageNumber="175">Z. testaceofasciatum</emphasis>
</taxonomicName>
, two species described not far from each other (eastern
<collectingCountry id="86D7760EFFA3E31BFD27FE7C43492EEB" box="[732,858,403,429]" name="Uzbekistan" pageId="14" pageNumber="175">Uzbekistan</collectingCountry>
and western
<collectingCountry id="86D7760EFFA3E31BFC0FFE7C44772EEB" box="[1012,1124,403,429]" name="Tajikistan" pageId="14" pageNumber="175">Tajikistan</collectingCountry>
) and currently known only from females. It differs from the males of its congeners by having two transverse light stripes on the abdomen (
<emphasis id="CCB4EA8CFFA3E31BFA85FE5745862E97" box="[1406,1429,440,465]" italics="true" pageId="14" pageNumber="175">vs</emphasis>
. lacking in other species), and a V-shaped dark marking on a light carapace (
<emphasis id="CCB4EA8CFFA3E31BFBF7FE3344302EB3" box="[1036,1059,476,501]" italics="true" pageId="14" pageNumber="175">vs</emphasis>
. carapace uniformly coloured or with small longitudinal spots along the fovea). The male palp of
<taxonomicName id="39C04D1DFFA3E31BFC7BFDEF44002D5F" authority="Zamani &amp; Marusik, 2022" authorityName="Zamani &amp; Marusik" authorityYear="2022" box="[896,1043,512,537]" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="175" phylum="Arthropoda" rank="species" species="turanicum" status="sp. nov.">
<emphasis id="CCB4EA8CFFA3E31BFC7BFDEF44002D5F" box="[896,1043,512,537]" italics="true" pageId="14" pageNumber="175">Z. turanicum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D78757F7FFA3E31BFBE7FDEF444A2D5C" box="[1052,1113,512,538]" pageId="14" pageNumber="175" rank="species">sp. n.</taxonomicNameLabel>
is most similar to that of
<taxonomicName id="39C04D1DFFA3E31BFA7DFDEF410B2D7B" authorityName=": Zamani &amp; Marusik" authorityYear="2021" class="Arachnida" family="Zodariidae" genus="Zodariellum" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="175" phylum="Arthropoda" rank="species" species="bactrianum">
<emphasis id="CCB4EA8CFFA3E31BFA7DFDEF410B2D7B" italics="true" pageId="14" pageNumber="175">Z. bactrianum</emphasis>
</taxonomicName>
by having two processes in the distal half of RTA, but can be differentiated by the angled anterodorsal part of RTA (
<emphasis id="CCB4EA8CFFA3E31BFED1FDA8415A2D27" box="[298,329,583,609]" italics="true" pageId="14" pageNumber="175">Ap</emphasis>
) (
<emphasis id="CCB4EA8CFFA3E31BFE98FDA741692D27" box="[355,378,584,609]" italics="true" pageId="14" pageNumber="175">vs</emphasis>
. rounded), and branches located at the same level (
<emphasis id="CCB4EA8CFFA3E31BFC4DFDA743DE2D27" box="[950,973,584,609]" italics="true" pageId="14" pageNumber="175">vs</emphasis>
. mesal one [=closer to cymbium] located more posteriorly).
</paragraph>
</subSubSection>
<subSubSection id="B6DA6515FFA3E31BFF3CFD60418D2B57" pageId="14" pageNumber="175" type="description">
<paragraph id="FE7F369EFFA3E31BFF3CFD6043552C1B" blockId="14.[151,1437,259,1078]" pageId="14" pageNumber="175">
<emphasis id="CCB4EA8CFFA3E31BFF3CFD60414A2DEF" bold="true" box="[199,345,655,681]" pageId="14" pageNumber="175">Description.</emphasis>
Male. Habitus as in
<figureCitation id="66FB2A1BFFA3E31BFDBEFD60428F2DEF" box="[581,668,655,681]" captionStart="FIGURE 1" captionStartId="2.[151,250,1953,1977]" captionTargetBox="[151,1436,605,1926]" captionTargetId="figure-210@2.[151,1436,594,1926]" captionTargetPageId="2" captionText="FIGURE 1. Habitus of Zodariellum asiaticum (A), Z. bactrianum (B), Z. turanicum sp. n. (C), Z. nenilini (DE) and Z. turkmenicum sp. n. (F), dorsal view. AD, F males; E female. Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7022241" httpUri="https://zenodo.org/record/7022241/files/figure.png" pageId="14" pageNumber="175">Fig. 1C</figureCitation>
. Total length 3.55. Carapace 1.60 long, 1.50 wide. Eye sizes: AME: 0.18, ALE: 0.10, PME: 0.10, PLE: 0.08. Carapace, sternum, chelicerae, labium and maxillae yellowish. Carapace light coloured with irregular V-shaped marking. Legs slightly lighter than carapace.Abdomen dorsally dark maroon, with 2 pale transverse stripes in anterior half and 3 obtuse triangular spots in anterior half; venter light beige. Measurements of legs: I: 6.78 (1.82, 0.62, 1.59, 1.77, 0.98), II: 6.23 (1.67, 0.55, 1.37, 1.78, 0.86), III: 6.57 (1.77, 0.61, 1.38, 1.91, 0.90), IV: 8.22 (2.34, 0.63, 1.88, 2.44, 0.93).
</paragraph>
<paragraph id="FE7F369EFFA3E31BFF3CFC8845512CAB" blockId="14.[151,1437,259,1078]" pageId="14" pageNumber="175">
Palp as in
<figureCitation id="66FB2A1BFFA3E31BFEBFFC8841B02CC7" box="[324,419,871,897]" captionStart="FIGURE 2" captionStartId="3.[151,250,1852,1876]" captionTargetBox="[191,1400,424,1829]" captionTargetId="figure-65@3.[173,1414,412,1829]" captionTargetPageId="3" captionText="FIGURE 2. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B, F), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E). AE ventral view; F dorsal view of cymbium, showing cymbial diverticulum. Scale bars = 0.2 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Eb—base of embolus, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branch(es) of retrolateral tibial apophysis, Sp—spur of tegular apophysis, Tg—tegular apophysis, To—triangular outgrowth, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022243" httpUri="https://zenodo.org/record/7022243/files/figure.png" pageId="14" pageNumber="175">Figs 2D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFE49FC8841C52CC7" box="[434,470,871,897]" captionStart="FIGURE 3" captionStartId="4.[151,250,1555,1579]" captionTargetBox="[152,1436,181,1531]" captionTargetId="figure-21@4.[152,1436,181,1531]" captionTargetPageId="4" captionText="FIGURE 3. SEM images of the male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), ventral view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Eb—base of embolus, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Sp—spur of tegular apophysis, Tg—tegular apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022245" httpUri="https://zenodo.org/record/7022245/files/figure.png" pageId="14" pageNumber="175">3D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFE1EFC8842192CC7" box="[485,522,871,897]" captionStart="FIGURE 4" captionStartId="5.[151,250,1907,1931]" captionTargetBox="[192,1383,207,1870]" captionTargetId="figure-17@5.[175,1412,181,1884]" captionTargetPageId="5" captionText="FIGURE 4. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), retrolateral view. Scale bars = 0.2 mm. Abbreviations: Ap—anterodorsal part of retrolateral tibial apophysis, Cn—conductor, Di—diverticulum, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022249" httpUri="https://zenodo.org/record/7022249/files/figure.png" pageId="14" pageNumber="175">4D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFDE3FC88422E2CC4" box="[536,573,871,898]" captionStart="FIGURE 5" captionStartId="6.[151,250,1584,1608]" captionTargetBox="[177,1411,181,1560]" captionTargetId="figure-21@6.[177,1411,181,1560]" captionTargetPageId="6" captionText="FIGURE 5. SEM images of the male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), retrolateral view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Cp—prolateral arm of conductor, Di—diverticulum, Eb—base of embolus, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr—retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022251" httpUri="https://zenodo.org/record/7022251/files/figure.png" pageId="14" pageNumber="175">5D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFDB7FC8842632CC7" box="[588,624,871,897]" captionStart="FIGURE 6" captionStartId="7.[151,250,1878,1902]" captionTargetBox="[192,1423,460,1840]" captionTargetId="figure-119@7.[151,1436,450,1840]" captionTargetPageId="7" captionText="FIGURE 6. Male palps of Zodariellum asiaticum (A), Z. bactrianum (B), Z. nenilini (C), Z. turanicum sp. n. (D) and Z. turkmenicum sp. n. (E), dorsal view. Scale bars = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.7022257" httpUri="https://zenodo.org/record/7022257/files/figure.png" pageId="14" pageNumber="175">6D</figureCitation>
,
<figureCitation id="66FB2A1BFFA3E31BFD84FC8842B72CC7" box="[639,676,871,897]" captionStart="FIGURE 9" captionStartId="10.[151,250,1757,1781]" captionTargetBox="[153,1434,345,1733]" captionTargetId="figure-84@10.[153,1434,345,1733]" captionTargetPageId="10" captionText="FIGURE 9. SEM images of the male palps of Zodariellum asiaticum (A), Z. bactrianum (BC) and Z. turanicum sp. n. (D). A anteroretrolateral view; B ventroretrolateral view; C anteroventral view; D anterior view. Scale bars = 0.1 mm. Abbreviations: Cn—conductor, Eb—base of embolus, Ra—retrolateral tibial apophysis, Rb—branches of retrolateral tibial apophysis, Tr— retrolateral arm of tegular apophysis, Tu—tutaculum." figureDoi="http://doi.org/10.5281/zenodo.7022263" httpUri="https://zenodo.org/record/7022263/files/figure.png" pageId="14" pageNumber="175">9D</figureCitation>
. Tibial apophysis longer than tibia, 2.3 times longer than wide in proximal part, tip inclined, with anterodorsal part (
<emphasis id="CCB4EA8CFFA3E31BFD3DFC6442F62CE3" box="[710,741,907,933]" italics="true" pageId="14" pageNumber="175">Ap</emphasis>
) angled; dorsal part with 2 processes originated at anterior 1/3, processes of subequal length and shape and located at the same level (one behind the other), ectal margin smoothly rounded in ventral view; cymbium length/width ratio 1.8; embolus originating at ca. 4:30 oclock position.
</paragraph>
<paragraph id="FE7F369EFFA3E31BFF3CFC18418D2B57" blockId="14.[151,1437,259,1078]" box="[199,414,1015,1041]" pageId="14" pageNumber="175">Female. Unknown.</paragraph>
</subSubSection>
<subSubSection id="B6DA6515FFA3E31BFF3CFBF445772B73" box="[199,1380,1051,1078]" pageId="14" pageNumber="175" type="distribution">
<paragraph id="FE7F369EFFA3E31BFF3CFBF445772B73" blockId="14.[151,1437,259,1078]" box="[199,1380,1051,1078]" pageId="14" pageNumber="175">
<emphasis id="CCB4EA8CFFA3E31BFF3CFBF441722B73" bold="true" box="[199,353,1051,1077]" pageId="14" pageNumber="175">Distribution.</emphasis>
Known only from the
<typeStatus id="217B883CFFA3E31BFD9BFBF342832B70" box="[608,656,1052,1078]" pageId="14" pageNumber="175">type</typeStatus>
locality in
<collectingRegion id="3C04F87CFFA3E31BFCF4FBF443A32B73" box="[783,944,1051,1077]" country="Turkmenistan" name="Lebap Welayaty" pageId="14" pageNumber="175">Lebap Region</collectingRegion>
, northeastern
<collectingCountry id="86D7760EFFA3E31BFBB7FBF444F42B73" box="[1100,1255,1051,1077]" name="Turkmenistan" pageId="14" pageNumber="175">Turkmenistan</collectingCountry>
(
<figureCitation id="66FB2A1BFFA3E31BFB0DFBF4454F2B73" box="[1270,1372,1051,1077]" captionStart="FIGURE 12" captionStartId="12.[151,250,1866,1890]" captionTargetBox="[164,1424,453,1842]" captionTargetId="figure-144@12.[164,1424,453,1842]" captionTargetPageId="12" captionText="FIGURE 12. Distribution records of Zodariellum spp. A: 1) Z. asiaticum, 2) Z. bactrianum, 3) Z. bekuzini, 4) Z. continentale, 5) Z. proszynskii, 6) Z. spasskyi and Z. zebra, 7) Z. surprisum, 8) Z. sytchevskajae, 9) Z. turanicum sp. n., 10) Z. turkmenicum sp. n., 11) Z. tadzhikum, 12) Z. testaceofasciatum, 13) Z. volgouralense, 14) Z. nenilini (in part, excluding records from Mongolia). B: 1) Z. nenilini (in part, records from Mongolia), 2) Z. schmidti, 3) Z. mongolicum, 4) Z. chaoyangense, 5) Z. furcum, 6) Z. hunanense, 7) Z. planum. A single point may refer to several closely located records." figureDoi="http://doi.org/10.5281/zenodo.7022273" httpUri="https://zenodo.org/record/7022273/files/figure.png" pageId="14" pageNumber="175">Fig. 12A</figureCitation>
)
</paragraph>
</subSubSection>
</treatment>
</document>