treatments-xml/data/09/3C/E9/093CE911FFE11D5B17B698578FBEFB94.xml
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<document id="8DB3B9B4109DC5A0EF17DDBF4BDADC20" ID-DOI="10.11646/zootaxa.5296.1.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="7970231" ID-ZooBank="62B8599B-5E59-42B4-A15D-40A724181BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1685010340024" checkinUser="plazi" docAuthor="Huemer, Peter &amp; Mayr, Toni" docDate="2023" docId="093CE911FFE11D5B17B698578FBEFB94" docLanguage="en" docName="zootaxa.5296.1.7.pdf" docOrigin="Zootaxa 5296 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5296.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Exapate aidasi Huemer &amp; Mayr 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="81" masterDocId="F5059169FFE21D5D17219C488C3CFF85" masterDocTitle="A surprising new species of Exapate Hübner, 1825 (Lepidoptera, Tortricidae) from Armenia" masterLastPageNumber="82" masterPageNumber="75" pageNumber="78" updateTime="1685010744749" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="8AE04134BA9092CE2A1EF38F4E225F31">A surprising new species of Exapate Hübner, 1825 (Lepidoptera, Tortricidae) from Armenia</mods:title>
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<mods:namePart id="B3471E7B57767466C66BB167EDC0D190">Huemer, Peter</mods:namePart>
<mods:affiliation id="2F9A244950FE2E8986B89942DE5AF644">Tiroler Landesmuseen Betriebsges. m. b. H., Natural History Collections, Krajnc-Str. 1, A- 6060 Hall in Tirol, Austria</mods:affiliation>
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<mods:namePart id="F12E4F05CB3B406462368B1CA871B7DD">Mayr, Toni</mods:namePart>
<mods:affiliation id="73854038A0FFAE263730BA0606A93478">Egelseestrasse 21, A- 6800 Feldkirch, Austria</mods:affiliation>
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<mods:date id="B7F0A0B1D0ADF1050840952FB38AA9B0">2023</mods:date>
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<heading id="DA62EF6BFFE11D5E17B698578D93FBBC" bold="true" box="[151,431,1054,1081]" fontSize="11" level="1" pageId="3" pageNumber="78" reason="1">
<emphasis id="B3E18415FFE11D5E17B698578D93FBBC" bold="true" box="[151,431,1054,1081]" pageId="3" pageNumber="78">
<taxonomicName id="46952384FFE11D5E17B698578D76FBBC" authority="Huemer &amp; Mayr, 2023" authorityName="Huemer &amp; Mayr" authorityYear="2023" box="[151,330,1054,1081]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="aidasi" status="sp. nov.">
<emphasis id="B3E18415FFE11D5E17B698578D76FBBC" bold="true" box="[151,330,1054,1081]" italics="true" pageId="3" pageNumber="78">Exapate aidasi</emphasis>
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</emphasis>
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<subSubSection id="C98F0B8CFFE11D5E17B6980B8D37FBD8" box="[151,267,1091,1117]" pageId="3" pageNumber="78" type="description">
<paragraph id="812A5807FFE11D5E17B6980B8D37FBD8" blockId="3.[151,431,1054,1117]" box="[151,267,1091,1117]" pageId="3" pageNumber="78">
(
<figureCitation id="19AE4482FFE11D5E17BE980B8D3FFBD8" box="[159,259,1091,1117]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStart-2="FIGURES 46" captionStartId-0="3.[151,250,2018,2042]" captionStartId-1="4.[151,250,1891,1915]" captionStartId-2="5.[151,264,1935,1959]" captionTargetBox-0="[151,1436,1177,1994]" captionTargetBox-1="[151,1436,705,1868]" captionTargetBox-2="[354,1231,181,1908]" captionTargetId-0="figure-324@3.[151,1436,1177,1994]" captionTargetId-1="figure-283@4.[151,1436,705,1868]" captionTargetId-2="figure-20@5.[354,1233,181,1911]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionText-0="FIGURE 2. Exapate aidasi sp. nov., adult, holotype, male. Scale bar = 5 mm." captionText-1="FIGURE 3. Male adults of Exapate spp.; Exapate aidasi sp. nov. (upper row, holotype left); Exapate duratella (middle row); Exapate congelatella (lower row). Scale bar = 10mm." captionText-2="FIGURES 46. Exapate spp., male genitalia. 4. E. aidasi sp. nov., paratype (gen. slide TOR 486). 5. E. congelatella (gen. slide TOR 487); 6. E. duratella (gen. slide TOR 485). Scale bars = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7970238" figureDoi-1="http://doi.org/10.5281/zenodo.7970240" figureDoi-2="http://doi.org/10.5281/zenodo.7970242" httpUri-0="https://zenodo.org/record/7970238/files/figure.png" httpUri-1="https://zenodo.org/record/7970240/files/figure.png" httpUri-2="https://zenodo.org/record/7970242/files/figure.png" pageId="3" pageNumber="78">Figs 24</figureCitation>
)
</paragraph>
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<paragraph id="812A5807FFE11D5E17B69BAA8F8DF87C" blockId="3.[151,945,2018,2042]" box="[151,945,2018,2042]" pageId="3" pageNumber="78">
<emphasis id="B3E18415FFE11D5E17B69BAA8D31F87F" bold="true" box="[151,269,2018,2042]" pageId="3" pageNumber="78">FIGURE 2</emphasis>
.
<taxonomicName id="46952384FFE11D5E16389BAB8D8DF87F" authority="Huemer &amp; Mayr, 2023" authorityName="Huemer &amp; Mayr" authorityYear="2023" box="[281,433,2018,2042]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="aidasi" status="sp. nov.">
<emphasis id="B3E18415FFE11D5E16389BAB8D8DF87F" box="[281,433,2018,2042]" italics="true" pageId="3" pageNumber="78">Exapate aidasi</emphasis>
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<emphasis id="B3E18415FFE11D5E16969BAA8E3BF87F" bold="true" box="[439,519,2018,2042]" pageId="3" pageNumber="78">
<taxonomicNameLabel id="A8D2396EFFE11D5E16969BAA8E3BF87F" box="[439,519,2018,2042]" pageId="3" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, adult, holotype, male. Scale bar = 5 mm.
</paragraph>
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<emphasis id="B3E18415FFE61D5917B69CDF8D70FF35" bold="true" box="[151,332,150,177]" pageId="4" pageNumber="79">Type material.</emphasis>
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<typeStatus id="5E2EE6A5FFE61D59167B9CDF8DFEFF34" box="[346,450,151,177]" pageId="4" pageNumber="79">Holotype</typeStatus>
♁,
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<collectingCountry id="F9821897FFE61D5916DB9CDE8E58FF35" box="[506,612,150,176]" name="Armenia" pageId="4" pageNumber="79">Armenia</collectingCountry>
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,
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,
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&amp;
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leg.; DNA barcode ID
<collectionCode id="E784C0C2FFE61D5915C59CF38F13FF50" box="[740,815,187,213]" country="Austria" httpUri="http://grbio.org/cool/7159-6y1v" name="Tiroler Landesmuseum Ferdinandeum" pageId="4" pageNumber="79">TLMF</collectionCode>
_
<collectorName id="2C603DD1FFE61D59141F9CF38F55FF50" box="[830,873,187,213]" pageId="4" pageNumber="79">Lep</collectorName>
_32188 (
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.
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: 7 ♁,
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,
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road,
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&amp;
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leg.; genitalia slide TOR 486 ♁ (
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,
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). 1
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♁,
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<collectingCountry id="F9821897FFE61D5916969D4A8E1DFE99" box="[439,545,258,284]" name="Armenia" pageId="4" pageNumber="79">Armenia</collectingCountry>
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, near
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,
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riv. valley,
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<collectorName id="2C603DD1FFE61D5917B69D6F8CE8FEC4" box="[151,212,295,321]" pageId="4" pageNumber="79">Duda</collectorName>
&amp;
<collectorName id="2C603DD1FFE61D5917D99D6F8D64FEC4" box="[248,344,295,321]" pageId="4" pageNumber="79">Saldaitis</collectorName>
leg. (
<collectionCode id="E784C0C2FFE61D5916B69D6F8DD1FEC4" box="[407,493,295,321]" pageId="4" pageNumber="79">RCTM</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C98F0B8CFFE61D5917E69D028831FD2C" pageId="4" pageNumber="79" type="diagnosis">
<paragraph id="812A5807FFE61D5917E69D028831FD2C" blockId="4.[151,1437,150,681]" pageId="4" pageNumber="79">
<emphasis id="B3E18415FFE61D5917E69D028D7EFEE1" bold="true" box="[199,322,330,356]" pageId="4" pageNumber="79">Diagnosis.</emphasis>
The new species differs from both congeners externally by its smaller size and the darker grey hindwings, but to some extent also by the wing pattern (
<figureCitation id="19AE4482FFE61D59142C9D278F4FFE0C" box="[781,883,367,393]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,2018,2042]" captionStartId-1="4.[151,250,1891,1915]" captionTargetBox-0="[151,1436,1177,1994]" captionTargetBox-1="[151,1436,705,1868]" captionTargetId-0="figure-324@3.[151,1436,1177,1994]" captionTargetId-1="figure-283@4.[151,1436,705,1868]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Exapate aidasi sp. nov., adult, holotype, male. Scale bar = 5 mm." captionText-1="FIGURE 3. Male adults of Exapate spp.; Exapate aidasi sp. nov. (upper row, holotype left); Exapate duratella (middle row); Exapate congelatella (lower row). Scale bar = 10mm." figureDoi-0="http://doi.org/10.5281/zenodo.7970238" figureDoi-1="http://doi.org/10.5281/zenodo.7970240" httpUri-0="https://zenodo.org/record/7970238/files/figure.png" httpUri-1="https://zenodo.org/record/7970240/files/figure.png" pageId="4" pageNumber="79">Figs 23</figureCitation>
). In
<taxonomicName id="46952384FFE61D5914879D278873FE0C" baseAuthorityName="Clerck" baseAuthorityYear="1759" box="[934,1103,367,393]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="congelatella">
<emphasis id="B3E18415FFE61D5914879D278873FE0C" box="[934,1103,367,393]" italics="true" pageId="4" pageNumber="79">E. congelatella</emphasis>
</taxonomicName>
the forewings are rather light brown to brownish grey, usually with well defined markings, whereas in
<taxonomicName id="46952384FFE61D5914F09DDB8864FE28" authorityName="Heyden" authorityYear="1864" box="[977,1112,403,429]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="duratella">
<emphasis id="B3E18415FFE61D5914F09DDB8864FE28" box="[977,1112,403,429]" italics="true" pageId="4" pageNumber="79">E. duratella</emphasis>
</taxonomicName>
the darker coloured veins of the terminal part of the forewing are characteristic. Both species are distinctly larger on average (wingspan c.
<quantity id="466DF5E2FFE61D5912779DFF8CFFFE70" metricMagnitude="-2" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="4" pageNumber="79" unit="mm" value="22.0" valueMax="23.0" valueMin="21.0">2123 mm</quantity>
in
<taxonomicName id="46952384FFE61D5917C79D938DB2FE70" baseAuthorityName="Clerck" baseAuthorityYear="1759" box="[230,398,475,501]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="congelatella">
<emphasis id="B3E18415FFE61D5917C79D938DB2FE70" box="[230,398,475,501]" italics="true" pageId="4" pageNumber="79">E. congelatella</emphasis>
</taxonomicName>
, c.
<quantity id="466DF5E2FFE61D5916969D928E13FE70" box="[439,559,474,501]" metricMagnitude="-2" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="4" pageNumber="79" unit="mm" value="24.5" valueMax="25.0" valueMin="24.0">2425 mm</quantity>
in
<taxonomicName id="46952384FFE61D5915739D938EEBFE70" authorityName="Heyden" authorityYear="1864" box="[594,727,475,501]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="duratella">
<emphasis id="B3E18415FFE61D5915739D938EEBFE70" box="[594,727,475,501]" italics="true" pageId="4" pageNumber="79">E. duratella</emphasis>
</taxonomicName>
, c.
<quantity id="466DF5E2FFE61D5915DE9D938F4BFE70" box="[767,887,475,501]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="1.9" metricValueMin="1.7" pageId="4" pageNumber="79" unit="mm" value="18.0" valueMax="19.0" valueMin="17.0">1719 mm</quantity>
in
<taxonomicName id="46952384FFE61D5914BB9D938FC1FE70" authority="Huemer &amp; Mayr, 2023" authorityName="Huemer &amp; Mayr" authorityYear="2023" box="[922,1021,475,501]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="aidasi" status="sp. nov.">
<emphasis id="B3E18415FFE61D5914BB9D938FC1FE70" box="[922,1021,475,501]" italics="true" pageId="4" pageNumber="79">E. aidasi</emphasis>
</taxonomicName>
<emphasis id="B3E18415FFE61D5913259D938861FE70" bold="true" box="[1028,1117,475,501]" pageId="4" pageNumber="79">
<taxonomicNameLabel id="A8D2396EFFE61D5913259D938861FE70" box="[1028,1117,475,501]" pageId="4" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
) and the hindwings are light grey to grey-brown. Further,
<taxonomicName id="46952384FFE61D5916F89DB78E00FD9C" authority="Huemer &amp; Mayr, 2023" authorityName="Huemer &amp; Mayr" authorityYear="2023" box="[473,572,511,537]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="aidasi" status="sp. nov.">
<emphasis id="B3E18415FFE61D5916F89DB78E00FD9C" box="[473,572,511,537]" italics="true" pageId="4" pageNumber="79">E. aidasi</emphasis>
</taxonomicName>
<emphasis id="B3E18415FFE61D5915639DB78EA7FD9C" bold="true" box="[578,667,511,537]" pageId="4" pageNumber="79">
<taxonomicNameLabel id="A8D2396EFFE61D5915639DB78EA7FD9C" box="[578,667,511,537]" pageId="4" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the related species by subtle but diagnostic characters of the male genitalia, particularly the narrower uncus and valva and the absence of a subapical thorn of the pallus and the presence of a medial thorn (
<figureCitation id="19AE4482FFE61D5916D49E0F8E67FDE4" box="[501,603,583,609]" captionStart="FIGURES 46" captionStartId="5.[151,264,1935,1959]" captionTargetBox="[354,1231,181,1908]" captionTargetId="figure-20@5.[354,1233,181,1911]" captionTargetPageId="5" captionText="FIGURES 46. Exapate spp., male genitalia. 4. E. aidasi sp. nov., paratype (gen. slide TOR 486). 5. E. congelatella (gen. slide TOR 487); 6. E. duratella (gen. slide TOR 485). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7970242" httpUri="https://zenodo.org/record/7970242/files/figure.png" pageId="4" pageNumber="79">Figs 46</figureCitation>
). Finally, the DNA barcode sequence is clearly divergent at a distance of c. 2.5%.
<taxonomicName id="46952384FFE61D5917FE9E238DB4FD00" baseAuthorityName="Clerck" baseAuthorityYear="1759" box="[223,392,619,645]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="congelatella">
<emphasis id="B3E18415FFE61D5917FE9E238DB4FD00" box="[223,392,619,645]" italics="true" pageId="4" pageNumber="79">E. congelatella</emphasis>
</taxonomicName>
and
<taxonomicName id="46952384FFE61D59169F9E238E78FD00" authorityName="Heyden" authorityYear="1864" box="[446,580,619,645]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="duratella">
<emphasis id="B3E18415FFE61D59169F9E238E78FD00" box="[446,580,619,645]" italics="true" pageId="4" pageNumber="79">E. duratella</emphasis>
</taxonomicName>
are extremely similar in genitalia and hardly discernible (
<bibRefCitation id="E50425F6FFE61D5913E89E238CEEFD2C" author="Bruun, H. H. &amp; Krogerus, H." pageId="4" pageNumber="79" pagination="77 - 86" refId="ref3643" refString="Bruun, H. H. &amp; Krogerus, H. (1996) Exapate bicuspidella sp. n., a new species of Tortricidae from northeastenr Finland (Lepidoptera). Entomologica Fennica, 7, 77 - 86. https: // doi. org / 10.33338 / ef. 83895" type="journal article" year="1996">Bruun &amp; Krogerus 1996</bibRefCitation>
,
<bibRefCitation id="E50425F6FFE61D5917FC9EC78DACFD2C" author="Razowski, J." box="[221,400,655,681]" pageId="4" pageNumber="79" refId="ref4250" refString="Razowski, J. (2002) Tortricidae (Lepidoptera) of Europe. Vol. 1. Tortricinae and Chlidanotinae. F. Slamka, Bratislava, 247 pp." type="book" year="2002">Razowski 2002</bibRefCitation>
). Male genitalia differ slightly by the shape of the uncus.
</paragraph>
</subSubSection>
<caption id="D5EA088FFFE61D5917B69B2B8E85F825" ID-DOI="http://doi.org/10.5281/zenodo.7970240" ID-Zenodo-Dep="7970240" httpUri="https://zenodo.org/record/7970240/files/figure.png" pageId="4" pageNumber="79" startId="4.[151,250,1891,1915]" targetBox="[151,1436,705,1868]" targetPageId="4" targetType="figure">
<paragraph id="812A5807FFE61D5917B69B2B8E85F825" blockId="4.[151,1436,1891,1952]" pageId="4" pageNumber="79">
<emphasis id="B3E18415FFE61D5917B69B2B8D32F8F9" bold="true" box="[151,270,1891,1916]" pageId="4" pageNumber="79">FIGURE 3</emphasis>
. Male adults of
<taxonomicName id="46952384FFE61D5916949B2C8E00F8FE" box="[437,572,1892,1915]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B3E18415FFE61D5916949B2C8E34F8FE" box="[437,520,1892,1915]" italics="true" pageId="4" pageNumber="79">Exapate</emphasis>
spp.
</taxonomicName>
;
<taxonomicName id="46952384FFE61D5915669B2C8EE3F8F9" authority="Huemer &amp; Mayr, 2023" authorityName="Huemer &amp; Mayr" authorityYear="2023" box="[583,735,1892,1916]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="aidasi" status="sp. nov.">
<emphasis id="B3E18415FFE61D5915669B2C8EE3F8F9" box="[583,735,1892,1916]" italics="true" pageId="4" pageNumber="79">Exapate aidasi</emphasis>
</taxonomicName>
<emphasis id="B3E18415FFE61D5915C49B2C8F0BF8F9" bold="true" box="[741,823,1892,1916]" pageId="4" pageNumber="79">
<taxonomicNameLabel id="A8D2396EFFE61D5915C49B2C8F0BF8F9" box="[741,823,1892,1916]" pageId="4" pageNumber="79" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(upper row, holotype left);
<taxonomicName id="46952384FFE61D5913739B2C8935F8F9" authorityName="Heyden" authorityYear="1864" box="[1106,1289,1892,1916]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="duratella">
<emphasis id="B3E18415FFE61D5913739B2C8935F8F9" box="[1106,1289,1892,1916]" italics="true" pageId="4" pageNumber="79">Exapate duratella</emphasis>
</taxonomicName>
(middle row);
<taxonomicName id="46952384FFE61D5917B69BC08D52F825" baseAuthorityName="Clerck" baseAuthorityYear="1759" box="[151,366,1928,1952]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="congelatella">
<emphasis id="B3E18415FFE61D5917B69BC08D52F825" box="[151,366,1928,1952]" italics="true" pageId="4" pageNumber="79">Exapate congelatella</emphasis>
</taxonomicName>
(lower row). Scale bar = 10mm.
</paragraph>
</caption>
<caption id="D5EA088FFFE71D5817B69BC78F62F84E" ID-DOI="http://doi.org/10.5281/zenodo.7970242" ID-Zenodo-Dep="7970242" httpUri="https://zenodo.org/record/7970242/files/figure.png" pageId="5" pageNumber="80" startId="5.[151,264,1935,1959]" targetBox="[354,1231,181,1908]" targetPageId="5" targetType="figure">
<paragraph id="812A5807FFE71D5817B69BC78F62F84E" blockId="5.[151,1436,1935,1996]" pageId="5" pageNumber="80">
<emphasis id="B3E18415FFE71D5817B69BC78D08F822" bold="true" box="[151,308,1935,1959]" pageId="5" pageNumber="80">FIGURES 46</emphasis>
.
<taxonomicName id="46952384FFE71D5816619BD88DF9F822" box="[320,453,1936,1959]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B3E18415FFE71D5816619BD88DAFF822" box="[320,403,1936,1959]" italics="true" pageId="5" pageNumber="80">Exapate</emphasis>
spp.
</taxonomicName>
, male genitalia. 4.
<taxonomicName id="46952384FFE71D5815A39BD88EE7F82D" authority="Huemer &amp; Mayr, 2023" authorityName="Huemer &amp; Mayr" authorityYear="2023" box="[642,731,1936,1960]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="aidasi" status="sp. nov.">
<emphasis id="B3E18415FFE71D5815A39BD88EE7F82D" box="[642,731,1936,1960]" italics="true" pageId="5" pageNumber="80">E. aidasi</emphasis>
</taxonomicName>
<emphasis id="B3E18415FFE71D5815C09BD88F0DF82D" bold="true" box="[737,817,1936,1960]" pageId="5" pageNumber="80">
<taxonomicNameLabel id="A8D2396EFFE71D5815C09BD88F0DF82D" box="[737,817,1936,1960]" pageId="5" pageNumber="80" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype (gen. slide TOR 486). 5.
<taxonomicName id="46952384FFE71D5813B49BD88911F82D" baseAuthorityName="Clerck" baseAuthorityYear="1759" box="[1173,1325,1936,1960]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="congelatella">
<emphasis id="B3E18415FFE71D5813B49BD88911F82D" box="[1173,1325,1936,1960]" italics="true" pageId="5" pageNumber="80">E. congelatella</emphasis>
</taxonomicName>
(gen. slide TOR 487); 6.
<taxonomicName id="46952384FFE71D5816049BFC8DA2F849" authorityName="Heyden" authorityYear="1864" box="[293,414,1972,1996]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="duratella">
<emphasis id="B3E18415FFE71D5816049BFC8DA2F849" box="[293,414,1972,1996]" italics="true" pageId="5" pageNumber="80">E. duratella</emphasis>
</taxonomicName>
(gen. slide TOR 485). Scale bars = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C98F0B8CFFE41D5B17E69CDE8E2CFD2C" pageId="6" pageNumber="81" type="description">
<paragraph id="812A5807FFE41D5B17E69CDE892BFEE0" blockId="6.[151,1437,150,1041]" pageId="6" pageNumber="81">
<emphasis id="B3E18415FFE41D5B17E69CDE8D89FF35" bold="true" box="[199,437,150,176]" pageId="6" pageNumber="81">Description. Adult (</emphasis>
<figureCitation id="19AE4482FFE41D5B16949CDF8E27FF34" box="[437,539,151,177]" captionStart-0="FIGURE 2" captionStart-1="FIGURE 3" captionStartId-0="3.[151,250,2018,2042]" captionStartId-1="4.[151,250,1891,1915]" captionTargetBox-0="[151,1436,1177,1994]" captionTargetBox-1="[151,1436,705,1868]" captionTargetId-0="figure-324@3.[151,1436,1177,1994]" captionTargetId-1="figure-283@4.[151,1436,705,1868]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="FIGURE 2. Exapate aidasi sp. nov., adult, holotype, male. Scale bar = 5 mm." captionText-1="FIGURE 3. Male adults of Exapate spp.; Exapate aidasi sp. nov. (upper row, holotype left); Exapate duratella (middle row); Exapate congelatella (lower row). Scale bar = 10mm." figureDoi-0="http://doi.org/10.5281/zenodo.7970238" figureDoi-1="http://doi.org/10.5281/zenodo.7970240" httpUri-0="https://zenodo.org/record/7970238/files/figure.png" httpUri-1="https://zenodo.org/record/7970240/files/figure.png" pageId="6" pageNumber="81">Figs 23</figureCitation>
). Head, thorax, tegulae and abdomen grey, frons and neck covered with brushlike hairy scales, labial palpi well developed, turned upwards; antenna dark grey, scape scaled, pedicellus and flagellomeres ciliated, with cilia of about half the length of the width of the flagellomeres; wingspan
<quantity id="466DF5E2FFE41D5B13D29C9789A4FF7C" box="[1267,1432,223,249]" metricMagnitude="-2" metricUnit="m" metricValue="1.825" metricValueMax="1.92" metricValueMin="1.73" pageId="6" pageNumber="81" unit="mm" value="18.25" valueMax="19.2" valueMin="17.3">17.319.2 mm</quantity>
; forewing grey with irregular dark grey mottling, particularly in the discal and discocellular area, on veins Sc and R
<subScript id="1D115A42FFE41D5B12AC9D5D89A9FEA6" attach="left" box="[1421,1429,277,291]" fontSize="6" pageId="6" pageNumber="81">1</subScript>
, and in the terminal area; base of fringes with dark grey dots at end of veins, basal cilia line light grey-brown, distal part darker grey; hindwings dark greyish, cilia light grey brown and darker grey with darker basal line.
</paragraph>
<paragraph id="812A5807FFE41D5B17E69D278CC4FE28" blockId="6.[151,1437,150,1041]" pageId="6" pageNumber="81">Variation. The extent of the darker markings on the forewing varies considerably and is occasionally largely reduced.</paragraph>
<paragraph id="812A5807FFE41D5B17E69DFF8ED0FD00" blockId="6.[151,1437,150,1041]" pageId="6" pageNumber="81">
<emphasis id="B3E18415FFE41D5B17E69DFF8D48FE55" bold="true" box="[199,372,438,465]" pageId="6" pageNumber="81">Male genitalia</emphasis>
(
<figureCitation id="19AE4482FFE41D5B16A49DFF8DF0FE54" box="[389,460,439,465]" captionStart="FIGURES 46" captionStartId="5.[151,264,1935,1959]" captionTargetBox="[354,1231,181,1908]" captionTargetId="figure-20@5.[354,1233,181,1911]" captionTargetPageId="5" captionText="FIGURES 46. Exapate spp., male genitalia. 4. E. aidasi sp. nov., paratype (gen. slide TOR 486). 5. E. congelatella (gen. slide TOR 487); 6. E. duratella (gen. slide TOR 485). Scale bars = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7970242" httpUri="https://zenodo.org/record/7970242/files/figure.png" pageId="6" pageNumber="81">Fig. 4</figureCitation>
). Uncus short and with a broad base about two-fifths the width of tegumen, medially abruptly widened with gradually constricted distal half; socii small, haired; gnathos medially short, ovate, lateral arms long, with short thorns; transtilla well developed, band-like, densely covered with medium-sized stiff setae in middle; juxta subtriangular, with long digitate appendices posteriorly; valva long and moderately slender, sacculus more strongly sclortised, with subbasal emargination and short finger-like appendix in middle; phallus long, slender, apically evenly pointed, with small postmedial thorn.
</paragraph>
<paragraph id="812A5807FFE41D5B17E69EC78E2CFD2C" blockId="6.[151,1437,150,1041]" box="[199,528,654,681]" pageId="6" pageNumber="81">
<emphasis id="B3E18415FFE41D5B17E69EC78DAFFD2D" bold="true" box="[199,403,654,681]" pageId="6" pageNumber="81">Female genitalia.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C98F0B8CFFE41D5B17E69EFA8DEEFCD8" pageId="6" pageNumber="81" type="biology_ecology">
<paragraph id="812A5807FFE41D5B17E69EFA8DEEFCD8" blockId="6.[151,1437,150,1041]" pageId="6" pageNumber="81">
<emphasis id="B3E18415FFE41D5B17E69EFA8D1BFD49" bold="true" box="[199,295,690,716]" pageId="6" pageNumber="81">Biology.</emphasis>
Adults have been collected in early and late October at light. Host-plant and early stages are unknown, however, the congeneric species mainly feed on woody plants and the new species probably also feeds on such. One of the localities is a semi-desert with only a few woody plants which may be potential host-plants, e.g.
<taxonomicName id="46952384FFE41D5B126A9EB38CDFFCBD" class="Magnoliopsida" family="Rosaceae" genus="Prunus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="species" species="incana">
<emphasis id="B3E18415FFE41D5B126A9EB38CDFFCBD" italics="true" pageId="6" pageNumber="81">Prunus incana</emphasis>
</taxonomicName>
or
<taxonomicName id="46952384FFE41D5B16299F578DE9FCBC" box="[264,469,799,825]" class="Magnoliopsida" family="Rhamnaceae" genus="Rhamnus" kingdom="Plantae" order="Rosales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="species" species="pallassii">
<emphasis id="B3E18415FFE41D5B16299F578DE9FCBC" box="[264,469,799,825]" italics="true" pageId="6" pageNumber="81">Rhamnus pallassii</emphasis>
</taxonomicName>
.
<taxonomicName id="46952384FFE41D5B16C29F578E1CFCBD" box="[483,544,799,824]" class="Pinopsida" family="Pinaceae" genus="Larix" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="genus">
<emphasis id="B3E18415FFE41D5B16C29F578E1CFCBD" box="[483,544,799,824]" italics="true" pageId="6" pageNumber="81">Larix</emphasis>
</taxonomicName>
, the main host-plant for
<taxonomicName id="46952384FFE41D5B14109F578F8BFCBC" authorityName="Heyden" authorityYear="1864" box="[817,951,799,825]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="duratella">
<emphasis id="B3E18415FFE41D5B14109F578F8BFCBC" box="[817,951,799,825]" italics="true" pageId="6" pageNumber="81">E. duratella</emphasis>
</taxonomicName>
according to
<bibRefCitation id="E50425F6FFE41D5B136F9F5788C1FCBC" author="Deutsch, H." box="[1102,1277,799,825]" pageId="6" pageNumber="81" pagination="351 - 356" refId="ref3697" refString="Deutsch, H. (2008) Exapate duratella Heyden 1864: Beitrag zur Biologie und Zuchtbericht (Lepidoptera, Tortricidae). Carinthia II, 198 / 118, 351 - 356." type="journal article" year="2008">Deutsch (2008)</bibRefCitation>
, is not present in
<collectingCountry id="F9821897FFE41D5B17939F0B8D28FCD8" box="[178,276,835,861]" name="Armenia" pageId="6" pageNumber="81">Armenia</collectingCountry>
(
<bibRefCitation id="E50425F6FFE41D5B16029F0B8DF9FCD8" author="Schmidt, W. C." box="[291,453,834,861]" pageId="6" pageNumber="81" pagination="6 - 18" refId="ref4373" refString="Schmidt, W. C. (1995) Around the world with Larix: an introduction. In: Schmidt, W. C. &amp; McDonald, K. J. (Eds.), Ecology and management of Larix forests: a look ahead. U. S. A. Gen. Tech. Rep. INT-GTR- 319. Intermountain Research Station, Ogden, Utah, pp. 6 - 18." type="book chapter" year="1995">Schmidt 1995</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C98F0B8CFFE41D5B17E69F2E8FB1FC4C" pageId="6" pageNumber="81" type="distribution">
<paragraph id="812A5807FFE41D5B17E69F2E8FB1FC4C" blockId="6.[151,1437,150,1041]" pageId="6" pageNumber="81">
<emphasis id="B3E18415FFE41D5B17E69F2E8D5DFC05" bold="true" box="[199,353,870,896]" pageId="6" pageNumber="81">Distribution.</emphasis>
<collectingCountry id="F9821897FFE41D5B16479F2F8DF0FC04" box="[358,460,871,897]" name="Armenia" pageId="6" pageNumber="81">Armenia</collectingCountry>
.
<taxonomicName id="46952384FFE41D5B16F79F2F8EBCFC04" baseAuthorityName="Clerck" baseAuthorityYear="1759" box="[470,640,871,897]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="congelatella">
<emphasis id="B3E18415FFE41D5B16F79F2F8EBCFC04" box="[470,640,871,897]" italics="true" pageId="6" pageNumber="81">E. congelatella</emphasis>
</taxonomicName>
is widespread in temperate Europe, extending its eastern range to Ural mts. (
<bibRefCitation id="E50425F6FFE41D5B17F49FC38DA8FC20" author="Sinev, S. Yu. &amp; Nedoshivina, S. V. &amp; Dubatolov, V. V. &amp; Zoological Institute RAS" box="[213,404,907,933]" pageId="6" pageNumber="81" pagination="120 - 156" refId="ref4451" refString="Sinev, S. Yu., Nedoshivina, S. V. &amp; Dubatolov, V. V. (2019) Tortricidae. In: Sinev, S. Yu. (Ed.), Catalogue of the Lepidoptera of Russia. 2 nd Edition. Zoological Institute RAS, St. Petersburg, pp. 120 - 156. [in Russian]" type="book chapter" year="2019">
Sinev
<emphasis id="B3E18415FFE41D5B163D9FC38D68FC20" box="[284,340,907,933]" italics="true" pageId="6" pageNumber="81">et al.</emphasis>
2019
</bibRefCitation>
) and allegedly
<collectingCountry id="F9821897FFE41D5B151C9FC38EFCFC20" box="[573,704,907,933]" name="Kazakhstan" pageId="6" pageNumber="81">Kazakhstan</collectingCountry>
(
<bibRefCitation id="E50425F6FFE41D5B15EC9FC38FBDFC20" author="Razowski, J." box="[717,897,907,933]" pageId="6" pageNumber="81" refId="ref4250" refString="Razowski, J. (2002) Tortricidae (Lepidoptera) of Europe. Vol. 1. Tortricinae and Chlidanotinae. F. Slamka, Bratislava, 247 pp." type="book" year="2002">Razowski 2002</bibRefCitation>
).
<taxonomicName id="46952384FFE41D5B14B39FC38824FC20" authorityName="Heyden" authorityYear="1864" box="[914,1048,907,933]" class="Insecta" family="Tortricidae" genus="Exapate" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="duratella">
<emphasis id="B3E18415FFE41D5B14B39FC38824FC20" box="[914,1048,907,933]" italics="true" pageId="6" pageNumber="81">E. duratella</emphasis>
</taxonomicName>
is restricted to European mountain systems: Alps, Pyrenees, Carpathians (
<bibRefCitation id="E50425F6FFE41D5B15669FE78FBEFC4C" author="Karle-Fendt, A. &amp; Wolf, W." box="[583,898,942,969]" pageId="6" pageNumber="81" pagination="39 - 42" refId="ref3970" refString="Karle-Fendt, A. &amp; Wolf, W. (2015) Ein Vorkommen von Exapate duratella Heyden, 1864 in den Allgauer Hochalpen (Insecta: Lepidoptera: Tortricidae). Beitrage zur bayerischen Entomofaunistik, 15, 39 - 42." type="journal article" year="2015">Karle-Fendt and Wolf 2015</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C98F0B8CFFE41D5B17E69F9B8FBEFB94" pageId="6" pageNumber="81" type="etymology">
<paragraph id="812A5807FFE41D5B17E69F9B8FBEFB94" blockId="6.[151,1437,150,1041]" pageId="6" pageNumber="81">
<emphasis id="B3E18415FFE41D5B17E69F9B8D72FC68" bold="true" box="[199,334,979,1005]" pageId="6" pageNumber="81">Etymology.</emphasis>
The new species is named in honour of Aidas Saldaitis, co-collector of the
<typeStatus id="5E2EE6A5FFE41D5B13A19F9B888CFC68" box="[1152,1200,979,1005]" pageId="6" pageNumber="81">type</typeStatus>
series, in recognition of his contribution in exploring
<taxonomicName id="46952384FFE41D5B16DD9FBF8EBFFB94" authorityName="Linnaeus" authorityYear="1758" box="[508,643,1015,1041]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Lepidoptera" pageId="6" pageNumber="81" phylum="Tracheophyta" rank="order">Lepidoptera</taxonomicName>
in many remote areas.
</paragraph>
</subSubSection>
</treatment>
</document>