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<document ID-DOI="http://dx.doi.org/10.3897/jhr.84.68709" ID-GBIF-Dataset="b9e61bc3-4158-4bee-8d57-a65655588f70" ID-Pensoft-Pub="1314-2607-84-145" ID-Pensoft-UUID="AEDB6967BB17574BB6988187C56EE6CA" ID-Zenodo-Dep="5349694" ID-ZooBank="5F83EC637C7947FDBD29E7A2B9328518" ModsDocID="1314-2607-84-145" checkinTime="1629847073322" checkinUser="pensoft" docAuthor="Okayasu, Juriya" docDate="2021" docId="7603CBDB1C6855F4B10DBD3ED5EBBA9B" docLanguage="en" docName="JourHymenoptRes 84: 145-168" docOrigin="Journal of Hymenoptera Research 84" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/jhr.84.68709" docTitle="Arkaditilla nallinia Okayasu 2021, comb. nov." docType="treatment" docVersion="4" id="AEDB6967BB17574BB6988187C56EE6CA" lastPageNumber="145" masterDocId="AEDB6967BB17574BB6988187C56EE6CA" masterDocTitle="A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species" masterLastPageNumber="168" masterPageNumber="145" pageNumber="145" updateTime="1678756233000" updateUser="pensoft">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new velvet ant genus Arkaditilla (Hymenoptera, Mutillidae, Trogaspidiini) from the Oriental Region, with review of species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Okayasu, Juriya</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6307-2939</mods:nameIdentifier>
<mods:affiliation>Entomological Laboratory, Faculty of Agriculture, Ehime University, Tarumi 3 - 5 - 7, Matsuyama, 790 - 8566, Japan &amp; Systematic Entomology, Graduate School of Agriculture, Hokkaido University, Sapporo, 060 - 8589, Japan</mods:affiliation>
<mods:nameIdentifier type="email">mutiphiidae@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Hymenoptera Research</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-08-24</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>84</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>145</mods:start>
<mods:end>168</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/jhr.84.68709</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/jhr.84.68709</mods:identifier>
<mods:identifier type="Pensoft-Pub">1314-2607-84-145</mods:identifier>
<mods:identifier type="ZooBank">5F83EC637C7947FDBD29E7A2B9328518</mods:identifier>
<mods:identifier type="Pensoft-UUID">AEDB6967BB17574BB6988187C56EE6CA</mods:identifier>
<mods:identifier type="Zenodo-Dep">5349694</mods:identifier>
</mods:mods>
<subSection pageId="0" pageNumber="145" type="results">
<treatment ID-GBIF-Taxon="184941295" LSID="urn:lsid:plazi:treatment:7603CBDB1C6855F4B10DBD3ED5EBBA9B" httpUri="http://treatment.plazi.org/id/7603CBDB1C6855F4B10DBD3ED5EBBA9B" lastPageNumber="145" pageId="0" pageNumber="145">
<subSubSection pageId="0" pageNumber="145" type="nomenclature">
<paragraph pageId="0" pageNumber="145">
<taxonomicName LSID="7603CBDB-1C68-55F4-B10D-BD3ED5EBBA9B" authority="(Zavattari, 1914)" authorityName="Okayasu" authorityYear="2021" baseAuthorityName="Zavattari" baseAuthorityYear="1914" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia" status="comb. nov.">Arkaditilla nallinia (Zavattari, 1914)</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="145">comb. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="description">
<paragraph pageId="0" pageNumber="145">
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">Figures 11</figureCitation>
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">, 12</figureCitation>
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">, 18</figureCitation>
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 24</figureCitation>
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 35</figureCitation>
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">, 36</figureCitation>
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 47</figureCitation>
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">, 48</figureCitation>
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">, 64-66</figureCitation>
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">, 72</figureCitation>
<figureCitation captionStart="Figure 73–76" captionStartId="F8" captionText="Figure 73 - 76. Distributional map of Arkaditilla spp. based on specimens examined herein and the literature (Cameron 1902; Lelej 1996 b; Mickel 1935, 1937; Zavattari 1914) 73 A. bagrada (Cameron) (red circle) 74 A. depressicornis (Mickel) (blue circle) 75 A. frim sp. nov. (yellow circle), A. gibba sp. nov. (purple triangle), and A. leleji sp. nov. (light blue square) 76 A. nallinia (Zavattari) (orange circle)." figureDoi="10.3897/jhr.84.68709.figure73-76" httpUri="https://binary.pensoft.net/fig/579487" pageId="0" pageNumber="145">, 76</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="reference_group">
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Mutilla" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Mutilla nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia" status="comb. nov.">Mutilla nallinia</taxonomicName>
Zavattari, 1914: 100, ♂, syntypes ♂ (Giava [= Java], [Indonesia]; Bali, [Indonesia]) [Museo Regionale di Scienze Naturali, Torino, Italy].
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Timulla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Timulla (Trogaspidia) nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia" status="comb. nov." subGenus="Trogaspidia">Timulla (Trogaspidia) nallinia</taxonomicName>
: Mickel, 1935: 263, ♂.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia" status="comb. nov.">Krombeinidia nallinia</taxonomicName>
:
<bibRefCitation author="Lelej, AS" journalOrPublisher="Far Eastern Entomologist" pageId="0" pageNumber="145" pagination="1 - 24" refId="B10" refString="Lelej, AS, 1996a. To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species. Far Eastern Entomologist 30: 1 - 24" title="To the knowledge of the East Asian species of the tribe Trogaspidiini Bischoff, 1920 (Hymenoptera, Mutillidae) with description of eight new genera and two new species." volume="30" year="1996 a">Lelej 1996a</bibRefCitation>
: 11;
<bibRefCitation author="Lelej, AS" journalOrPublisher="Dalnauka, Vladivostok" pageId="0" pageNumber="145" refId="B12" refString="Lelej, AS, 2005. Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok" title="Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region." year="2005">2005</bibRefCitation>
: 66;
<bibRefCitation author="Lo Cascio, P" journalOrPublisher="Biodiversity Journal" pageId="0" pageNumber="145" pagination="529 - 592" refId="B15" refString="Lo Cascio, P, 2015. Worldwide checklist of the island mutillid wasps (Hymenoptera Mutillidae). Biodiversity Journal 6 (2): 529 - 592" title="Worldwide checklist of the island mutillid wasps (Hymenoptera Mutillidae)." volume="6" year="2015">Lo Cascio 2015</bibRefCitation>
: 551;
<bibRefCitation author="Pagliano, G" journalOrPublisher="Bollettino del Museo Regionale di Scienze Naturali di Torino" pageId="0" pageNumber="145" pagination="5 - 425" refId="B24" refString="Pagliano, G, Brothers, DJ, Cambra, R, Lelej, AS, Lo Cascio, P, Matteini, Palmerini M, Scaramozzino, PL, Williams, KA, Romano, M, 2020. Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species. Bollettino del Museo Regionale di Scienze Naturali di Torino 36 (1-2): 5 - 425" title="Checklist of names in Mutillidae (Hymenoptera), with illustrations of selected species." volume="36" year="2020">Pagliano et al. 2020</bibRefCitation>
: 251.
</paragraph>
<paragraph pageId="0" pageNumber="145">
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Petersenidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Petersenidia nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia" status="comb. nov.">Petersenidia nallinia</taxonomicName>
:
<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
: 94, ♂.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="diagnosis">
<paragraph pageId="0" pageNumber="145">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Male.</emphasis>
Clypeus dorsally elevated, sloping ventrally. Mandible with sharp subapical and large inner subbasal teeth (Fig.
<figureCitation captionStart="Figures 13–18" captionStartId="F3" captionText="Figures 13 - 18. Arkaditilla spp., ♂, face 13 A. bagrada (Cameron) 14 A. depressicornis (Mickel) 15 A. frim sp. nov., holotype 16 A. gibba sp. nov., paratype 17 A. leleji sp. nov., holotype 18 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures13-18" httpUri="https://binary.pensoft.net/fig/579482" pageId="0" pageNumber="145">18</figureCitation>
), ventrally deeply excised with large basal tooth (Fig.
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">24</figureCitation>
). Scape ventral carinae divergent apically (Fig.
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">35</figureCitation>
), densely punctate between them; F1 apical 1/3, F2 and F3 basal 1/3 whitish yellow on ventral half; F1 strongly depressed, F1 height ~0.8
<normalizedToken originalValue="×">x</normalizedToken>
pedicel diameter (Fig.
<figureCitation captionStart="Figures 19–36" captionStartId="F4" captionText="Figures 19 - 36. Arkaditilla spp., ♂ 19 - 24 mandible, lateral view 25, 27, 29, 31, 33, 35 scape, ventral view 26, 28, 30, 32, 34, 36 pedicel, F 1, and F 2, lateral view 19, 25, 26 A. bagrada (Cameron) 20, 27, 28 A. depressicornis (Mickel) 21, 29, 30 A. frim sp. nov., holotype 22, 31, 32 A. gibba sp. nov., holotype 23, 33, 34 A. leleji sp. nov., holotype 24, 35, 36 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures19-36" httpUri="https://binary.pensoft.net/fig/579483" pageId="0" pageNumber="145">36</figureCitation>
). T3 orange (Fig.
<figureCitation captionStart="Figures 7–12" captionStartId="F2" captionText="Figures 7 - 12. Arkaditilla spp., ♂, habitus 7, 9, 11 dorsal view 8, 10, 12 lateral view 7, 8 A. gibba sp. nov., holotype 9, 10 A. leleji sp. nov., holotype 11, 12 A. nallinia (Zavattari). Scale bars: 3 mm." figureDoi="10.3897/jhr.84.68709.figures7-12" httpUri="https://binary.pensoft.net/fig/579481" pageId="0" pageNumber="145">12</figureCitation>
); cuspis strongly widened on posterior 1/3 with rounded apex (Figs
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">64</figureCitation>
,
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">72</figureCitation>
); digitus posteriorly expanded, forming triangular projection (Fig.
<figureCitation captionStart="Figures 37–48" captionStartId="F5" captionText="Figures 37 - 48. Arkaditilla spp., ♂, genitalia 37, 39, 41, 43, 45, 47 dorsal view 38, 40, 42, 44, 46, 48 ventral view 37, 38 A. bagrada (Cameron) 39, 40 A. depressicornis (Mickel) 41, 42 A. frim sp. nov., holotype 43, 44 A. gibba sp. nov., holotype 45, 46 A. leleji sp. nov., holotype 47, 48 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures37-48" httpUri="https://binary.pensoft.net/fig/579484" pageId="0" pageNumber="145">47</figureCitation>
); paracuspis slightly tuberculate (Fig.
<figureCitation captionStart="Figures 67–72" captionStartId="F7" captionText="Figures 67 - 72. Arkaditilla spp., ♂, genital volsella, ventro-inner view 67 A. bagrada (Cameron) 68 A. depressicornis (Mickel) 69 A. frim sp. nov., holotype 70 A. gibba sp. nov., holotype 71 A. leleji sp. nov., holotype 72 A. nallinia (Zavattari). White arrows indicate the paracuspis." figureDoi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145">72</figureCitation>
); penis valve weakly expanded ventrally (Figs
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">65</figureCitation>
,
<figureCitation captionStart="Figures 49–66" captionStartId="F6" captionText="Figures 49 - 66. Arkaditilla spp., ♂, genitalia 49, 52, 55, 58, 61, 64 genital capsule and volsella, inner view 50, 53, 56, 59, 62, 65 left penis valve 51, 54, 57, 60, 63, 66 right penis valve 49 - 51 A. bagrada (Cameron) 52 - 54 A. depressicornis (Mickel) 55 - 57 A. frim sp. nov., holotype 58 - 60 A. gibba sp. nov., holotype 61 - 63 A. leleji sp. nov., holotype 64 - 66 A. nallinia (Zavattari)." figureDoi="10.3897/jhr.84.68709.figures49-66" httpUri="https://binary.pensoft.net/fig/579485" pageId="0" pageNumber="145">66</figureCitation>
).
<emphasis bold="true" pageId="0" pageNumber="145">Female.</emphasis>
Unknown.
</paragraph>
<caption ID-Zenodo-Dep="5349708" doi="10.3897/jhr.84.68709.figures67-72" httpUri="https://binary.pensoft.net/fig/579486" pageId="0" pageNumber="145" start="Figures 67–72" startId="F7">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figures 67-72.</emphasis>
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
</taxonomicName>
spp., ♂, genital volsella, ventro-inner view
<emphasis bold="true" pageId="0" pageNumber="145">67</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
</taxonomicName>
(Cameron)
<emphasis bold="true" pageId="0" pageNumber="145">68</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
</taxonomicName>
(Mickel)
<emphasis bold="true" pageId="0" pageNumber="145">69</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim">
<emphasis italics="true" pageId="0" pageNumber="145">A. frim</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">70</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. gibba" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="145">A. gibba</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">71</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
<emphasis italics="true" pageId="0" pageNumber="145">A. leleji</emphasis>
</taxonomicName>
sp. nov., holotype
<emphasis bold="true" pageId="0" pageNumber="145">72</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
</taxonomicName>
(Zavattari). White arrows indicate the paracuspis.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="materials_examined">
<paragraph pageId="0" pageNumber="145">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="145">
<materialsCitation collectingDate="2006-08-12" collectorName="K. Watanabe" country="Indonesia" county="Tabanan" elevation="682" latitude="-8.392166" location="Pura Batu Salahan" longLatPrecision="915" longitude="115.083336" municipality="Bengkel" specimenCount="3" specimenCount-male="3" stateProvince="Bali">
<specimenCount type="male">2♂</specimenCount>
.
<emphasis bold="true" pageId="0" pageNumber="145">
<collectingCountry name="Indonesia">Indonesia</collectingCountry>
</emphasis>
:
<collectingRegion country="Indonesia" name="Bali">Bali</collectingRegion>
:
<collectingCounty>Tabanan</collectingCounty>
,
<collectingMunicipality>Bengkel</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:7603CBDB1C6855F4B10DBD3ED5EBBA9B:DDA77259C82FE250305E102260A9238B" country="Indonesia" county="Tabanan" latitude="-8.392166" longLatPrecision="915" longitude="115.083336" municipality="Bengkel" name="Pura Batu Salahan" stateProvince="Bali">Pura Batu Salahan</location>
,
<geoCoordinate degrees="08" direction="south" minutes="23" orientation="latitude" precision="1" seconds="31.8" value="-8.392166">08°23'31.8&quot;S</geoCoordinate>
,
<geoCoordinate degrees="115" direction="east" minutes="05" orientation="longitude" precision="925" value="115.083336">115°05'E</geoCoordinate>
,
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.82" unit="m" value="682.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.82" unit="m" value="682.0">682 m</elevation>
</quantity>
alt.,
<collectingDate value="2006-08-12">12.VIII.2006</collectingDate>
,
<collectorName>K. Watanabe</collectorName>
leg. [
<specimenCount type="male">1♂</specimenCount>
TUA]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="identification source">
<paragraph pageId="0" pageNumber="145">Identification source.</paragraph>
<paragraph pageId="0" pageNumber="145">
Keyed using
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel (1935)</bibRefCitation>
and compared with the original description. No differences were found between the specimen examined herein and
<normalizedToken originalValue="Zavattari’s">Zavattari's</normalizedToken>
description.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="distribution">
<paragraph pageId="0" pageNumber="145">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="145">
Indonesia: Bali, Java (
<bibRefCitation author="Zavattari, E" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="145" pagination="61 - 114" refId="B26" refString="Zavattari, E, 1914. Mutille Austro-Malesi. Bollettino della Societa Entomologica Italiana 45: 61 - 114" title="Mutille Austro-Malesi." volume="45" year="1914">Zavattari 1914</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel 1935</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="145" type="remarks">
<paragraph pageId="0" pageNumber="145">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="145">
Only three mutillid species have been recorded from Bali, namely
<taxonomicName authorityName="Okayasu" authorityYear="2021" baseAuthorityName="Zavattari" baseAuthorityYear="1914" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla nallinia</emphasis>
</taxonomicName>
,
<taxonomicName class="Insecta" family="Mutillidae" genus="Krombeinidia" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Krombeinidia griseomaculata" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="griseomaculata">
<emphasis italics="true" pageId="0" pageNumber="145">Krombeinidia griseomaculata</emphasis>
</taxonomicName>
(
<normalizedToken originalValue="André">Andre</normalizedToken>
, 1898), and
<taxonomicName authorityName="Hammer" authorityYear="1962" class="Insecta" family="Mutillidae" genus="Mickelomyrme" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Mickelomyrme pusillaeformis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="pusillaeformis">
<emphasis italics="true" pageId="0" pageNumber="145">Mickelomyrme pusillaeformis</emphasis>
</taxonomicName>
(Hammer, 1962) (
<bibRefCitation author="Zavattari, E" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="145" pagination="61 - 114" refId="B26" refString="Zavattari, E, 1914. Mutille Austro-Malesi. Bollettino della Societa Entomologica Italiana 45: 61 - 114" title="Mutille Austro-Malesi." volume="45" year="1914">Zavattari 1914</bibRefCitation>
;
<bibRefCitation author="Lelej, AS" journalOrPublisher="Dalnauka, Vladivostok" pageId="0" pageNumber="145" refId="B12" refString="Lelej, AS, 2005. Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok" title="Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region." year="2005">Lelej 2005</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4723.1.1" author="Okayasu, J" journalOrPublisher="Zootaxa" pageId="0" pageNumber="145" pagination="1 - 110" refId="B22" refString="Okayasu, J, 2020. Velvet ants of the tribe Smicromyrmini Bischoff (Hymenoptera: Mutillidae) of Japan. Zootaxa 4723 (1): 1 - 110, DOI: https://doi.org/10.11646/zootaxa.4723.1.1" title="Velvet ants of the tribe Smicromyrmini Bischoff (Hymenoptera: Mutillidae) of Japan." url="https://doi.org/10.11646/zootaxa.4723.1.1" volume="4723" year="2020">Okayasu 2020</bibRefCitation>
; current data), although
<normalizedToken originalValue="Zavattari’s">Zavattari's</normalizedToken>
record of
<taxonomicName lsidName="K. griseomaculata" pageId="0" pageNumber="145" rank="species" species="griseomaculata">
<emphasis italics="true" pageId="0" pageNumber="145">K. griseomaculata</emphasis>
</taxonomicName>
was not accepted by
<bibRefCitation author="Lelej, AS" journalOrPublisher="Dalnauka, Vladivostok" pageId="0" pageNumber="145" refId="B12" refString="Lelej, AS, 2005. Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region. Dalnauka, Vladivostok" title="Catalogue of the Mutillidae (Hymenoptera) of the Oriental Region." year="2005">Lelej (2005)</bibRefCitation>
and
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel (1935)</bibRefCitation>
. Field surveys in this small island will be useful for discovery of the female of
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
</taxonomicName>
.
</paragraph>
<caption ID-Zenodo-Dep="5349710" doi="10.3897/jhr.84.68709.figure73-76" httpUri="https://binary.pensoft.net/fig/579487" pageId="0" pageNumber="145" start="Figure 73–76" startId="F8">
<paragraph pageId="0" pageNumber="145">
<emphasis bold="true" pageId="0" pageNumber="145">Figure 73-76.</emphasis>
Distributional map of
<taxonomicName authorityName="Okayasu" authorityYear="2021" class="Insecta" family="Mutillidae" genus="Arkaditilla" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Arkaditilla" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="145">Arkaditilla</emphasis>
</taxonomicName>
spp. based on specimens examined herein and the literature (
<bibRefCitation author="Cameron, P" journalOrPublisher="Journal of the Straits Branch of the Royal Asiatic Society" pageId="0" pageNumber="145" pagination="29 - 131" refId="B2" refString="Cameron, P, 1902. On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum. Journal of the Straits Branch of the Royal Asiatic Society 37: 29 - 131" title="On the Hymenoptera collected by Mr. Robert Shelford at Sarawak, and on the Hymenoptera of the Sarawak Museum." volume="37" year="1902">Cameron 1902</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.3759/tropics.6.91" author="Lelej, AS" journalOrPublisher="Tropics" pageId="0" pageNumber="145" pagination="91 - 104" refId="B11" refString="Lelej, AS, 1996b. Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae). Tropics 6 (1-2): 91 - 104, DOI: https://doi.org/10.3759/tropics.6.91" title="Mutillid wasps collected in Malaysia and Indonesia by Dr. Sk. Yamane (Hymenoptera, Mutillidae)." url="https://doi.org/10.3759/tropics.6.91" volume="6" year="1996 b">Lelej 1996b</bibRefCitation>
;
<bibRefCitation DOI="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" author="Mickel, CE" journalOrPublisher="Transactions of the Royal Entomological Society of London" pageId="0" pageNumber="145" pagination="177 - 312" refId="B20" refString="Mickel, CE, 1935. The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae). Transactions of the Royal Entomological Society of London 83 (2): 177 - 312, DOI: https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" title="The mutillid wasps of the islands of the Pacific Ocean (Hymenoptera, Mutillidae)." url="https://doi.org/10.1111/j.1365-2311.1935.tb01207.x" volume="83" year="1935">Mickel 1935</bibRefCitation>
,
<bibRefCitation DOI="https://doi.org/10.1080/00222933708655284" author="Mickel, CE" journalOrPublisher="The Annals and Magazine of Natural History" pageId="0" pageNumber="145" pagination="441 - 456" refId="B21" refString="Mickel, CE, 1937. New species and records of Mutillidae (Hymenoptera) from Borneo and the Solomon Islands. The Annals and Magazine of Natural History 10 (19): 441 - 456, DOI: https://doi.org/10.1080/00222933708655284" title="New species and records of Mutillidae (Hymenoptera) from Borneo and the Solomon Islands." url="https://doi.org/10.1080/00222933708655284" volume="10" year="1937">1937</bibRefCitation>
;
<bibRefCitation author="Zavattari, E" journalOrPublisher="Bollettino della Societa Entomologica Italiana" pageId="0" pageNumber="145" pagination="61 - 114" refId="B26" refString="Zavattari, E, 1914. Mutille Austro-Malesi. Bollettino della Societa Entomologica Italiana 45: 61 - 114" title="Mutille Austro-Malesi." volume="45" year="1914">Zavattari 1914</bibRefCitation>
)
<emphasis bold="true" pageId="0" pageNumber="145">73</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. bagrada" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bagrada">
<emphasis italics="true" pageId="0" pageNumber="145">A. bagrada</emphasis>
</taxonomicName>
(Cameron) (red circle)
<emphasis bold="true" pageId="0" pageNumber="145">74</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. depressicornis" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="depressicornis">
<emphasis italics="true" pageId="0" pageNumber="145">A. depressicornis</emphasis>
</taxonomicName>
(Mickel) (blue circle)
<emphasis bold="true" pageId="0" pageNumber="145">75</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. frim" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="frim">
<emphasis italics="true" pageId="0" pageNumber="145">A. frim</emphasis>
</taxonomicName>
sp. nov. (yellow circle),
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. gibba" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="gibba">
<emphasis italics="true" pageId="0" pageNumber="145">A. gibba</emphasis>
</taxonomicName>
sp. nov. (purple triangle), and
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. leleji" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="leleji">
<emphasis italics="true" pageId="0" pageNumber="145">A. leleji</emphasis>
</taxonomicName>
sp. nov. (light blue square)
<emphasis bold="true" pageId="0" pageNumber="145">76</emphasis>
<taxonomicName class="Insecta" kingdom="Animalia" lsidName="A. nallinia" order="Hymenoptera" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="nallinia">
<emphasis italics="true" pageId="0" pageNumber="145">A. nallinia</emphasis>
</taxonomicName>
(Zavattari) (orange circle).
</paragraph>
</caption>
</subSubSection>
</treatment>
</subSection>
</document>
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