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<mods:title id="F4C07E2B3B95FC5E4B746E45925184CC">New species of Burmeistera (Campanulaceae: Lobelioideae) from Ecuador</mods:title>
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<mods:namePart id="5B35E4D024F15C18AFE8F2F724326229">Pérez, Álvaro J.</mods:namePart>
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<mods:namePart id="294C91C0EB62EB114FB0692BDA4771A5">Cevallos, Daniela</mods:namePart>
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<mods:namePart id="2231A4DCBE9A437909C1F3916844C3AC">Muchhala, Nathan</mods:namePart>
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<taxonomicName id="73B9B989FFF1AF1AFF41FE59FD1FC9EA" ID-CoL="8WRY5" authority="A. J. Perez &amp; N. Muchhala" authorityName="A. J. Perez &amp; N. Muchhala" authorityYear="2018" box="[136,698,410,437]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="draconis" status="sp. nov.">
<emphasis id="86CD1E18FFF1AF1AFF41FE59FE27C9EA" bold="true" box="[136,386,410,437]" italics="true" pageId="4" pageNumber="267">Burmeistera draconis</emphasis>
A. J. Pérez &amp; N. Muchhala
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, 
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(
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, 
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)
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Type:— 
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. 
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: Nudo de Sabanilla, east slope ca. 
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from pass on road Yangana-Valladolid, montane rain forest, 
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, 
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, 
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, 
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(fl,fr), 
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Harling 
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. &amp; Andersson 
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. 23645
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(
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: 
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26693!, 
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: 
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1290161!, 
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1185765!).
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<emphasis id="86CD1E18FFF1AF1AFF41FA03FF52CD8A" bold="true" box="[136,247,1472,1493]" pageId="4" pageNumber="267">FIGURE 3</emphasis>
. 
<taxonomicName id="73B9B989FFF1AF1AFECAFA03FD7BCD8A" authority="A. J. Perez &amp; N. Muchhala" authorityName="A. J. Perez &amp; N. Muchhala" authorityYear="2018" box="[259,734,1472,1493]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="267" phylum="Tracheophyta" rank="species" species="draconis" status="sp. nov.">
<emphasis id="86CD1E18FFF1AF1AFECAFA03FE71CD8A" box="[259,468,1472,1493]" italics="true" pageId="4" pageNumber="267">Burmeistera draconis</emphasis>
A. J. Pérez &amp; N. Muchhala
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. 
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.
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<emphasis id="86CD1E18FFF1AF1AFCF1FA03FCECCD8A" bold="true" box="[824,841,1472,1493]" pageId="4" pageNumber="267">A</emphasis>
. Dorso-lateral view of leaves and male flower. 
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. Abaxial view of leaves and fruit. 
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. Fruits. 
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. Female flower, lateral view. Photos by Álvaro J. Pérez.
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<emphasis id="86CD1E18FFF1AF1AFF41F9EAFED8CE1D" box="[136,381,1577,1602]" italics="true" pageId="4" pageNumber="267">Burmeistera draconis</emphasis>
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differs from other species of 
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<emphasis id="86CD1E18FFF1AF1AFD07F9EAFCFDCE1D" box="[718,856,1577,1602]" italics="true" pageId="4" pageNumber="267">Burmeistera</emphasis>
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by having elliptic leaves with an irregularly toothed margin and cylindrical inflated fruits with short calyx lobes.
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Freestanding shrub, 1.0– 
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with multiple stems, to scandent hemi-epiphytic shrub, climbing to 
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; stems 8.0– 
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diam., hispidulous, pale green; branches 2.0–5.0 mm diam., hispidulous, pale green; latex white. Leaves alternate, spirally arranged, chartaceous; lamina elliptic, glabrous, apex attenuate, base decurrent, 18–80 × 7.0– 
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, distal leaves gradually smaller than proximal leaves, leaf surfaces pale green when fresh, olive-green when dried; margin irregularly bidentate, the larger up to 6.0 mm long, with obtuse to rounded apices; venation semicraspedodromous, hispidulous on both sides, with 8 to 14 pair of secondary veins; petiole 5.0– 
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long × 
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diam., hispidulous. Flowers solitary in upper leaf axils, light green suffused with maroon; pedicels 
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long × 0.6–2.0 mm diam., hispidulous, pale green, curved to sinuate and ascending at anthesis, declined in fruit, ebracteolate. Hypanthium cup-shaped, 
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long × 5.0–7.0 mm diam., glabrous, light green. Calyx lobes triangular, 3.0–5.0 × 2.0–3.0 mm, glabrous, patent to arcuate at anthesis, with fine reticulate venation, margin finely denticulate, with 4.0– 5.0 teeth per side; apex acute. Corolla glabrous, 
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long, light green suffused with maroon externally, pale green within, corolla base slightly wider than hypanthium, corolla tube slightly curved, 
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long dorsally, narrowing from the base to the middle to 
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long; the two dorsal lobes lanceolate, falcate, 13–17 × 3.0–5.0 mm, acute at apex; the three ventral lobes narrowly triangular, falcate, 12 × 3.0 mm, acute at apex. Staminal column long-exerted, to ca. 
<quantity id="73416FEFFFF0AF1BFF06FF2BFEBAC95E" box="[207,287,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="268" unit="mm" value="18.0">18 mm</quantity>
beyond dorsal lobes; filament tube 
<quantity id="73416FEFFFF0AF1BFD70FF2BFC96C95E" box="[697,819,232,257]" metricMagnitude="-2" metricUnit="m" metricValue="2.65" metricValueMax="2.8" metricValueMin="2.5" pageId="5" pageNumber="268" unit="mm" value="26.5" valueMax="28.0" valueMin="25.0">25–28 mm</quantity>
long × 1.0– 
<quantity id="73416FEFFFF0AF1BFC09FF2BFBB3C95E" box="[960,1046,232,257]" metricMagnitude="-3" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="268" unit="mm" value="1.3">1.3 mm</quantity>
diam., slightly curved, glabrescent, pale green; anther tube curved-cylindrical, 9.0 mm long × 4.0 mm diam., dorsal and ventral anthers barbate, pale green suffused with maroon; dorsal anthers 9.0– 
<quantity id="73416FEFFFF0AF1BFDA2FEF3FD1DC916" box="[619,696,304,329]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" pageId="5" pageNumber="268" unit="mm" value="11.0">11 mm</quantity>
long; ventral anthers 6.0–7.0 mm long. Berry cylindrical, inflated with 1.0 mm thick walls, light green suffused with maroon, 
<quantity id="73416FEFFFF0AF1BFCFFFE97FC23C932" box="[822,902,340,365]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="268" unit="mm" value="29.0">29 mm</quantity>
long × 
<quantity id="73416FEFFFF0AF1BFC28FE97FB94C932" box="[993,1073,340,365]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="5" pageNumber="268" unit="mm" value="24.0">24 mm</quantity>
diam., crowned by the persistent calyx lobes; seeds elliptic, 0.8–1.0 mm.
</paragraph>
<paragraph id="B406C20AFFF0AF1BFF74FE5FFEA2C9A2" blockId="5.[136,1452,160,1625]" pageId="5" pageNumber="268">
<emphasis id="86CD1E18FFF0AF1BFF74FE5FFE90C9EA" box="[189,309,412,437]" italics="true" pageId="5" pageNumber="268">Etymology</emphasis>
:—The epithet is the genitive of the latin noun 
<emphasis id="86CD1E18FFF0AF1BFC9FFE5FFC32C9EA" box="[854,919,412,437]" italics="true" pageId="5" pageNumber="268">draco</emphasis>
, which means dragon, in reference to the way the bidentate leaves arranged around long stems with distal flowers resemble the serpentine dragons of Chinese mythology.
</paragraph>
<paragraph id="B406C20AFFF0AF1BFF74FDCBFEFACB42" blockId="5.[136,1452,160,1625]" pageId="5" pageNumber="268">
<emphasis id="86CD1E18FFF0AF1BFF74FDCBFD98CA7E" box="[189,573,520,545]" italics="true" pageId="5" pageNumber="268">Distribution, habitat and ecology</emphasis>
:— 
<taxonomicName id="73B9B989FFF0AF1BFDABFDCBFCFCCA7E" authorityName="A. J. Perez &amp; N. Muchhala" authorityYear="2018" box="[610,857,520,545]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="draconis">
<emphasis id="86CD1E18FFF0AF1BFDABFDCBFCFCCA7E" box="[610,857,520,545]" italics="true" pageId="5" pageNumber="268">Burmeistera draconis</emphasis>
</taxonomicName>
is endemic to the montane cloud forest of 
<collectingRegion id="767D0CE8FFF0AF1BFA9DFDCBFF5ECA1A" country="Ecuador" name="Zamora-Chinchipe" pageId="5" pageNumber="268">Zamora Chinchipe</collectingRegion>
between 
<quantity id="73416FEFFFF0AF1BFEAEFDEFFDA7CA1A" box="[359,514,556,581]" metricMagnitude="3" metricUnit="m" metricValue="2.55" metricValueMax="2.7" metricValueMin="2.4" pageId="5" pageNumber="268" unit="m" value="2550.0" valueMax="2700.0" valueMin="2400.0">2400–2700 m</quantity>
(
<figureCitation id="2C82DE8FFFF0AF1BFDDBFDEFFDF2CA1A" box="[530,599,556,581]" captionStart="FIGURE 2" captionStartId="3.[136,229,1079,1100]" captionTargetBox="[348,1238,187,1051]" captionTargetId="figure-14@3.[348,1238,187,1051]" captionTargetPageId="3" captionText="FIGURE 2. Distribution map of Burmeistera pterifolia (circle, Santa Lucia Cloud Forest Reserve in Pichincha province) and B. draconis (star, Reserva Tapichalaca in Zamora Chinchipe province) in Ecuador." figureDoi="http://doi.org/10.5281/zenodo.13703688" httpUri="https://zenodo.org/record/13703688/files/figure.png" pageId="5" pageNumber="268">Fig. 2</figureCitation>
). According to the Ministerio del Ambiente del 
<collectingCountry id="CCAE829AFFF0AF1BFBB8FDEFFB6BCA1A" box="[1137,1230,556,581]" name="Ecuador" pageId="5" pageNumber="268">Ecuador</collectingCountry>
(2013) this locality is within a much larger zone dominated by bosque siempreverde montano del sur de la cordillera oriental de los Andes (BsMn02). This species is sympatric with 
<taxonomicName id="73B9B989FFF0AF1BFDA2FDB7FB79CAD2" authority="Muchhala &amp; Perez (2015: 36)" authorityName="Muchhala &amp; Perez" authorityPageNumber="36" authorityYear="2015" box="[619,1244,628,653]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="zamorensis">
<emphasis id="86CD1E18FFF0AF1BFDA2FDB7FCDECAD2" box="[619,891,628,653]" italics="true" pageId="5" pageNumber="268">Burmeistera zamorensis</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFC4DFDB7FB79CAD2" author="Muchhala, N. &amp; Perez, A. J." box="[900,1244,628,653]" pageId="5" pageNumber="268" pagination="36 - 38" refId="ref5043" refString="Muchhala, N. &amp; Perez, A. J. (2015) Burmeistera zamorensis (Campanulaceae, Lobelioideae), a New Species from Southern Ecuador. Novon 24: 36 - 38. https: // doi. org / 10.3417 / 2014009" type="journal article" year="2015">Muchhala &amp; Pérez (2015: 36)</bibRefCitation>
</taxonomicName>
, and associated to species such as 
<taxonomicName id="73B9B989FFF0AF1BFE89FD5BFC81CAEE" authority="Gleasson (1925: 13)" authorityName="Gleasson" authorityPageNumber="13" authorityYear="1925" box="[320,804,664,689]" class="Magnoliopsida" family="Campanulaceae" genus="Centropogon" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="comosus">
<emphasis id="86CD1E18FFF0AF1BFE89FD5BFD99CAEE" box="[320,572,664,689]" italics="true" pageId="5" pageNumber="268">Centropogon comosus</emphasis>
Gleasson (1925:13)
</taxonomicName>
and 
<taxonomicName id="73B9B989FFF0AF1BFC96FD5BFEA5CA8A" authority="Santamaria-Aguilar &amp; Lagomarsino (2015: 92)" authorityName="Santamaria-Aguilar &amp; Lagomarsino" authorityPageNumber="92" authorityYear="2015" class="Magnoliopsida" family="Pentaphylacaceae" genus="Freziera" kingdom="Plantae" order="Ericales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="neillii">
<emphasis id="86CD1E18FFF0AF1BFC96FD5BFBACCAEE" box="[863,1033,664,689]" italics="true" pageId="5" pageNumber="268">Freziera neillii</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFBDBFD5BFEA5CA8A" author="Santamaria-Aguilar, D. &amp; Lagomarsino, L. P." pageId="5" pageNumber="268" pagination="89 - 102" refId="ref5146" refString="Santamaria-Aguilar, D. &amp; Lagomarsino, L. P. (2015) Cuatro nuevas especies y un nuevo registro de Freziera (Pentaphylacaceae) de Ecuador y Peru. Journal of the Botanical Research Institute of Texas 9: 89 - 102." type="journal article" year="2015">Santamaria-Aguilar &amp; Lagomarsino (2015: 92)</bibRefCitation>
</taxonomicName>
. It is likely that 
<taxonomicName id="73B9B989FFF0AF1BFE78FD7FFD97CA8A" authorityName="A. J. Perez &amp; N. Muchhala" authorityYear="2018" box="[433,562,700,725]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="draconis">
<emphasis id="86CD1E18FFF0AF1BFE78FD7FFD97CA8A" box="[433,562,700,725]" italics="true" pageId="5" pageNumber="268">B. draconis</emphasis>
</taxonomicName>
also occurs in the adjacent Podocarpus National Park and Yacuri National Park, where similar environmental conditions and forest 
<typeStatus id="6B027CA8FFF0AF1BFD0EFD23FD52CAA6" box="[711,759,736,761]" pageId="5" pageNumber="268">type</typeStatus>
are found. Flowers open at dusk and are pollinated by nectar bats (
<taxonomicName id="73B9B989FFF0AF1BFF0CFCC7FEF7CB42" box="[197,338,772,797]" class="Mammalia" family="Phyllostomidae" genus="Anoura" kingdom="Animalia" order="Chiroptera" pageId="5" pageNumber="268" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="86CD1E18FFF0AF1BFF0CFCC7FEBCCB42" box="[197,281,772,797]" italics="true" pageId="5" pageNumber="268">Anoura</emphasis>
spp.
</taxonomicName>
).
</paragraph>
<paragraph id="B406C20AFFF0AF1BFF74FCEBFECFCB3A" blockId="5.[136,1452,160,1625]" pageId="5" pageNumber="268">
<emphasis id="86CD1E18FFF0AF1BFF74FCEBFE90CB1E" box="[189,309,808,833]" italics="true" pageId="5" pageNumber="268">Phenology</emphasis>
:—Flowers and fruits are found from June to November, but some individuals were observed flowering throughout the year.
</paragraph>
<paragraph id="B406C20AFFF0AF1BFF74FCB3FB30CC46" blockId="5.[136,1452,160,1625]" pageId="5" pageNumber="268">
<emphasis id="86CD1E18FFF0AF1BFF74FCB3FE3ACBD6" box="[189,415,880,905]" italics="true" pageId="5" pageNumber="268">Conservation status</emphasis>
:—This species was found at two localities, close to each other. One is in the Tapichalaca Reserve, a privately owned protected area of 3500 ha managed by the Jocotoco Foundation since 1998, and the other one is a collection along the road from Yangana to Valladolid. The limited collections and the need to explore other nearby areas, including the adjacent Podocarpus National Park and Yacuri National Park, prevent us to accurately assess its conservation status. Thus we proposed to rank this species as Data Deficient (
<collectionCode id="D2A85ACFFFF0AF1BFB92FBC3FB2CCC46" box="[1115,1161,1024,1049]" country="India" lsid="urn:lsid:biocol.org:col:13033" name="Forest Research Institute, Indian Council of Forestry Research and Education" pageId="5" pageNumber="268" type="Herbarium">DD</collectionCode>
).
</paragraph>
<paragraph id="B406C20AFFF0AF1BFF74FBE7FCE1CDFA" blockId="5.[136,1452,160,1625]" pageId="5" pageNumber="268">
<emphasis id="86CD1E18FFF0AF1BFF74FBE7FE9DCC62" box="[189,312,1060,1085]" italics="true" pageId="5" pageNumber="268">Discussion</emphasis>
:— 
<taxonomicName id="73B9B989FFF0AF1BFE95FBE7FDEACC62" authorityName="A. J. Perez &amp; N. Muchhala" authorityYear="2018" box="[348,591,1060,1085]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="draconis">
<emphasis id="86CD1E18FFF0AF1BFE95FBE7FDEACC62" box="[348,591,1060,1085]" italics="true" pageId="5" pageNumber="268">Burmeistera draconis</emphasis>
</taxonomicName>
is easily recognizable by its elliptic leaves with irregularly bidentate margins and cylindrical inflated fruits with short calyx lobes. Preliminary phylogenetic studies (Muchhala, unpublished) suggest it is closely related to a group of 
<taxonomicName id="73B9B989FFF0AF1BFDBEFBAFFCF8CCDA" box="[631,861,1132,1157]" pageId="5" pageNumber="268">
<emphasis id="86CD1E18FFF0AF1BFDBEFBAFFCA4CCDA" box="[631,769,1132,1157]" italics="true" pageId="5" pageNumber="268">Burmeistera</emphasis>
species
</taxonomicName>
with inflated fruits, including 
<taxonomicName id="73B9B989FFF0AF1BFB0EFBAFFEAACCF6" authority="Wimmer (1932: 124)" authorityName="Wimmer" authorityPageNumber="124" authorityYear="1932" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="ramosa">
<emphasis id="86CD1E18FFF0AF1BFB0EFBAFFA9BCCDA" box="[1223,1342,1132,1157]" italics="true" pageId="5" pageNumber="268">B. ramosa</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFA80FBAFFEAACCF6" author="Wimmer, F. E." pageId="5" pageNumber="268" pagination="123 - 126" refId="ref5413" refString="Wimmer, F. E. (1932) Burmeistera. Eine umstrittene Pflanzengattung und ihre Arten. Repertorium Specierum Novarum Regni Vegetabilis 30. Dahlem, Berlin, pp. 123 - 126." type="book chapter" year="1932">Wimmer (1932: 124)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="73B9B989FFF0AF1BFED3FB53FD2FCCF6" authority="Wimmer (1932: 124)" authorityName="Wimmer" authorityPageNumber="124" authorityYear="1932" box="[282,650,1168,1193]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="refracta">
<emphasis id="86CD1E18FFF0AF1BFED3FB53FE37CCF6" box="[282,402,1168,1193]" italics="true" pageId="5" pageNumber="268">B. refracta</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFE50FB53FD2FCCF6" author="Wimmer, F. E." box="[409,650,1168,1193]" pageId="5" pageNumber="268" pagination="123 - 126" refId="ref5413" refString="Wimmer, F. E. (1932) Burmeistera. Eine umstrittene Pflanzengattung und ihre Arten. Repertorium Specierum Novarum Regni Vegetabilis 30. Dahlem, Berlin, pp. 123 - 126." type="book chapter" year="1932">Wimmer (1932: 124)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="73B9B989FFF0AF1BFD5CFB53FB7ACCF6" authority="(Kunth 1818: 307) Hook. f. &amp; B. D. Jacks" authorityName="Hook. f. &amp; B. D. Jacks" baseAuthorityName="Kunth" baseAuthorityPageNumber="307" baseAuthorityYear="1818" box="[661,1247,1168,1193]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="glabrata">
<emphasis id="86CD1E18FFF0AF1BFD5CFB53FCB0CCF6" box="[661,789,1168,1193]" italics="true" pageId="5" pageNumber="268">B. glabrata</emphasis>
(
<bibRefCitation id="D028BFFBFFF0AF1BFCEAFB53FC41CCF6" author="Kunth, C. S." box="[803,996,1168,1193]" pageId="5" pageNumber="268" refId="ref4479" refString="Kunth, C. S. (1818) Nova Genera et Species Pantarum. A La Libreirie Grecque-Latine-Allemende, Paris, 456 pp." type="book" year="1818">Kunth 1818: 307</bibRefCitation>
) Hook.f. &amp; B.D.Jacks
</taxonomicName>
in 
<bibRefCitation id="D028BFFBFFF0AF1BFACAFB53FF1BCC92" author="Jackson, B. D." pageId="5" pageNumber="268" refId="ref4343" refString="Jackson, B. D. (1893) Index kewensis. Clarendon Press, Oxford." type="book" year="1893">Jackson (1893: 361)</bibRefCitation>
, 
<taxonomicName id="73B9B989FFF0AF1BFF02FB77FDE4CC92" authority="Wimmer (1932: 123)" authorityName="Wimmer" authorityPageNumber="123" authorityYear="1932" box="[203,577,1204,1229]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="vulgaris">
<emphasis id="86CD1E18FFF0AF1BFF02FB77FEE2CC92" box="[203,327,1204,1229]" italics="true" pageId="5" pageNumber="268">B. vulgaris</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFE86FB77FDE4CC92" author="Wimmer, F. E." box="[335,577,1204,1229]" pageId="5" pageNumber="268" pagination="123 - 126" refId="ref5413" refString="Wimmer, F. E. (1932) Burmeistera. Eine umstrittene Pflanzengattung und ihre Arten. Repertorium Specierum Novarum Regni Vegetabilis 30. Dahlem, Berlin, pp. 123 - 126." type="book chapter" year="1932">Wimmer (1932: 123)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="73B9B989FFF0AF1BFD87FB77FC56CC92" authority="Jeppesen (1981: 40)" authorityName="Jeppesen" authorityPageNumber="40" authorityYear="1981" box="[590,1011,1204,1229]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="oyacachensis">
<emphasis id="86CD1E18FFF0AF1BFD87FB77FCA1CC92" box="[590,772,1204,1229]" italics="true" pageId="5" pageNumber="268">B. oyacachensis</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFCC5FB77FC56CC92" author="Jeppesen, S." box="[780,1011,1204,1229]" pageId="5" pageNumber="268" pagination="9 - 170" refId="ref4360" refString="Jeppesen, S. (1981) Lobeliaceae. In: Harling, G. &amp; Andersson, L. (Eds.) Flora of Ecuador, No 14. Swedish Natural Science Research Council, Stockholm, pp. 9 - 170." type="book chapter" year="1981">Jeppesen (1981: 40)</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="73B9B989FFF0AF1BFBC9FB77FAD3CC92" authority="Jeppesen (1981: 29)" authorityName="Jeppesen" authorityPageNumber="29" authorityYear="1981" box="[1024,1398,1204,1229]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="borjensis">
<emphasis id="86CD1E18FFF0AF1BFBC9FB77FB22CC92" box="[1024,1159,1204,1229]" italics="true" pageId="5" pageNumber="268">B. borjensis</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFB46FB77FAD3CC92" author="Jeppesen, S." box="[1167,1398,1204,1229]" pageId="5" pageNumber="268" pagination="9 - 170" refId="ref4360" refString="Jeppesen, S. (1981) Lobeliaceae. In: Harling, G. &amp; Andersson, L. (Eds.) Flora of Ecuador, No 14. Swedish Natural Science Research Council, Stockholm, pp. 9 - 170." type="book chapter" year="1981">Jeppesen (1981: 29)</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="73B9B989FFF0AF1BFF41FB1BFD8DCCAE" authority="Wimmer (1955: 108)" authorityName="Wimmer" authorityPageNumber="108" authorityYear="1955" box="[136,552,1240,1265]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis id="86CD1E18FFF0AF1BFF41FB1BFE94CCAE" box="[136,305,1240,1265]" italics="true" pageId="5" pageNumber="268">B. oblongifolia</emphasis>
<bibRefCitation id="D028BFFBFFF0AF1BFEFEFB1BFD8DCCAE" author="Wimmer, F. E." box="[311,552,1240,1265]" pageId="5" pageNumber="268" pagination="107 - 111" refId="ref5448" refString="Wimmer, F. E. (1955) Lobeliacearum species Novae Austro. Americanae. Brittonia 8: 107 - 111. https: // doi. org / 10.2307 / 2804853" type="journal article" year="1955">Wimmer (1955: 108)</bibRefCitation>
</taxonomicName>
. Based on calyx lobes and other traits, it is most similar to 
<taxonomicName id="73B9B989FFF0AF1BFB08FB1BFACCCCAE" authorityName="Wimmer" authorityYear="1955" box="[1217,1385,1240,1265]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis id="86CD1E18FFF0AF1BFB08FB1BFACCCCAE" box="[1217,1385,1240,1265]" italics="true" pageId="5" pageNumber="268">B. oblongifolia</emphasis>
</taxonomicName>
(both have short triangular calyx lobes, &lt;
<quantity id="73416FEFFFF0AF1BFDE0FB3FFDD4CD4A" box="[553,625,1276,1301]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="268" unit="mm" value="5.0">5 mm</quantity>
), but it differs in the bidentate leaf margin (vs. callose-denticulate in 
<taxonomicName id="73B9B989FFF0AF1BFA5AFB3FFEB4CD66" authorityName="Wimmer" authorityYear="1955" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="oblongifolia">
<emphasis id="86CD1E18FFF0AF1BFA5AFB3FFEB4CD66" italics="true" pageId="5" pageNumber="268">B. oblongifolia</emphasis>
</taxonomicName>
). All other species in this group have calyx lobes&gt; 
<quantity id="73416FEFFFF0AF1BFC80FAE3FC2CCD66" box="[841,905,1312,1337]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="268" unit="mm" value="5.0">5 mm</quantity>
in length. Based on leaf margin, it is most similar to 
<taxonomicName id="73B9B989FFF0AF1BFF6EFA87FEF9CD02" authorityName="Jeppesen" authorityYear="1981" box="[167,348,1348,1373]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="oyacachensis">
<emphasis id="86CD1E18FFF0AF1BFF6EFA87FEF9CD02" box="[167,348,1348,1373]" italics="true" pageId="5" pageNumber="268">B. oyacachensis</emphasis>
</taxonomicName>
, which has coarsely repand-lacerate margins. This species is endemic to the 
<collectingRegion id="767D0CE8FFF0AF1BFB05FA87FAD3CD02" box="[1228,1398,1348,1373]" country="Ecuador" name="Napo" pageId="5" pageNumber="268">Napo province</collectingRegion>
, and differs from 
<taxonomicName id="73B9B989FFF0AF1BFED1FAABFE3FCDDE" authorityName="A. J. Perez &amp; N. Muchhala" authorityYear="2018" box="[280,410,1384,1409]" class="Magnoliopsida" family="Campanulaceae" genus="Burmeistera" kingdom="Plantae" order="Asterales" pageId="5" pageNumber="268" phylum="Tracheophyta" rank="species" species="draconis">
<emphasis id="86CD1E18FFF0AF1BFED1FAABFE3FCDDE" box="[280,410,1384,1409]" italics="true" pageId="5" pageNumber="268">B. draconis</emphasis>
</taxonomicName>
by its long calyx lobes (
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). In terms of leaf margin shape, all other species in this group have almost entire to shallowly repand-dentate margins.
</paragraph>
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(
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)
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:— 
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: 
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, on trail starting from 
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. Muchhala 464, 465
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(
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!)
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; 
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en el sendero de las 
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. 
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. Pérez et al. 7141
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(
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!, 
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(fl, fr), 
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. 
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. Pérez et al. 9117
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(
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!, 
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!, 
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.
</paragraph>
</subSubSection>
</treatment>
</document>