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<mods:title id="3C0BA8BB44D498E86938D4B109F23486">Review of the millipede family Diplomaragnidae of Japan, with description of a new species and the restoration of the combination Diplomaragna hokkaidensis (Verhoeff, 1939) (Diplopoda, Chordeumatida, Diplomaragnidae)</mods:title>
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<mods:namePart id="4E3BE5FDA5BD3944877B660A445DEAEF">Mikhaljova, Elena V.</mods:namePart>
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<treatment id="03F087F6FFCC4F0C6FFDFB55511C4BDA" ID-DOI="http://doi.org/10.5281/zenodo.6094952" ID-GBIF-Taxon="119593295" ID-Zenodo-Dep="6094952" LSID="urn:lsid:plazi:treatment:03F087F6FFCC4F0C6FFDFB55511C4BDA" httpUri="http://treatment.plazi.org/id/03F087F6FFCC4F0C6FFDFB55511C4BDA" lastPageNumber="570" pageId="1" pageNumber="570">
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<paragraph id="8BE636E0FFCC4F0C6FFDFB5556904998" blockId="1.[151,700,1213,1239]" box="[151,700,1213,1239]" pageId="1" pageNumber="570">
<heading id="D0AE818CFFCC4F0C6FFDFB5556904998" bold="true" box="[151,700,1213,1239]" fontSize="11" level="1" pageId="1" pageNumber="570" reason="1">
<taxonomicName id="4C594D63FFCC4F0C6FFDFB5556904998" ID-CoL="36LVZ" authority="Verhoeff, 1939" authorityName="Verhoeff" authorityYear="1939" box="[151,700,1213,1239]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="species" species="hokkaidensis">
<emphasis id="B92DEAF2FFCC4F0C6FFDFB5556904998" bold="true" box="[151,700,1213,1239]" pageId="1" pageNumber="570">
<emphasis id="B92DEAF2FFCC4F0C6FFDFB5555C04998" bold="true" box="[151,492,1213,1239]" italics="true" pageId="1" pageNumber="570">Diplomaragna hokkaidensis</emphasis>
(
<bibRefCitation id="EFC84B11FFCC4F0C6E91FB55569F4998" author="Verhoeff" box="[507,691,1213,1239]" pageId="1" pageNumber="570" refString="Verhoeff, K. W. (1939) Zur Kenntnis ostasiatischer Diplopoden. III. Zoologischer Anzeiger, 127, 113 - 125." type="journal article" year="1939">Verhoeff, 1939</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BE636E0FFCC4F0C6FFDFAEC570E4855" blockId="1.[151,1333,1284,1400]" box="[151,802,1284,1306]" pageId="1" pageNumber="570">
<treatmentCitationGroup id="AB4911CEFFCC4F0C6FFDFAEC570E4855" box="[151,802,1284,1306]" pageId="1" pageNumber="570">
<taxonomicName id="4C594D63FFCC4F0C6FFDFAEC56A74855" ID-CoL="53YRQ" authority="Verhoeff 1939: 113" authorityName="Verhoeff" authorityPageNumber="113" authorityYear="1939" box="[151,651,1284,1306]" class="Diplopoda" family="Diplomaragnidae" genus="Syntelopodeuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="species" species="hokkaidense">
<emphasis id="B92DEAF2FFCC4F0C6FFDFAEC55ED4855" box="[151,449,1284,1306]" italics="true" pageId="1" pageNumber="570">Syntelopodeuma hokkaidense</emphasis>
<treatmentCitation id="0AF810F1FFCC4F0C6EA2FAEC56A74855" author="Verhoeff" box="[456,651,1284,1306]" page="113" pageId="1" pageNumber="570" year="1939">
<bibRefCitation id="EFC84B11FFCC4F0C6EA2FAEC56704855" author="Verhoeff" box="[456,604,1284,1306]" pageId="1" pageNumber="570" refString="Verhoeff, K. W. (1939) Zur Kenntnis ostasiatischer Diplopoden. III. Zoologischer Anzeiger, 127, 113 - 125." type="journal article" year="1939">Verhoeff 1939</bibRefCitation>
: 113
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116, figs 14.
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</paragraph>
<paragraph id="8BE636E0FFCC4F0C6FFDFACD56424815" blockId="1.[151,1333,1284,1400]" pageId="1" pageNumber="570">
<treatmentCitationGroup id="AB4911CEFFCC4F0C6FFDFACD56424815" pageId="1" pageNumber="570">
<emphasis id="B92DEAF2FFCC4F0C6FFDFACD55FB4874" box="[151,471,1317,1339]" italics="true" pageId="1" pageNumber="570">
<taxonomicName id="4C594D63FFCC4F0C6FFDFACD55124874" ID-CoL="7Q3KB" box="[151,318,1317,1339]" class="Diplopoda" family="Diplomaragnidae" genus="Syntelopodeuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="genus">Syntelopodeuma</taxonomicName>
hokkaidense—
</emphasis>
<treatmentCitation id="0AF810F1FFCC4F0C6EB2FACD56804874" author="Takakuwa" box="[472,684,1317,1339]" page="127" pageId="1" pageNumber="570" year="1954">
<bibRefCitation id="EFC84B11FFCC4F0C6EB2FACD56504874" author="Takakuwa" box="[472,636,1317,1339]" pageId="1" pageNumber="570" refString="Takakuwa, Y. (1954) Diplopoden aus Japan und ihn angrenzenden Gebieten. Japan Society for the Promotion of Science, Tokyo, 241 pp." type="book" year="1954">Takakuwa 1954</bibRefCitation>
: 127
</treatmentCitation>
128, fig. 145;
<treatmentCitation id="0AF810F1FFCC4F0C6C2DFACD57DC4874" author="Miyosi" box="[839,1008,1317,1339]" page="53" pageId="1" pageNumber="570" year="1959">
<bibRefCitation id="EFC84B11FFCC4F0C6C2DFACD57E74874" author="Miyosi" box="[839,971,1317,1339]" pageId="1" pageNumber="570" refString="Miyosi, Y. (1959) Uber japanische Diplopoden. Arachnological Society of East Asia, Osaka, 223 pp. [in Japanese, with German summary]" type="book" year="1959">Miyosi 1959</bibRefCitation>
: 53
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, 126, 219, pl. 12: figs 174, 174'.
<taxonomicName id="4C594D63FFCC4F0C6FFDFAAC55994815" ID-CoL="36LVZ" box="[151,437,1348,1370]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="species" species="hokkaidensis">
<emphasis id="B92DEAF2FFCC4F0C6FFDFAAC55994815" box="[151,437,1348,1370]" italics="true" pageId="1" pageNumber="570">Diplomaragna hokkaidensis</emphasis>
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<treatmentCitation id="0AF810F1FFCC4F0C6EA5FAAC56464815" author="Shear" box="[463,618,1348,1370]" page="32" pageId="1" pageNumber="570" year="1990">
<bibRefCitation id="EFC84B11FFCC4F0C6EA5FAAC56694815" author="Shear" box="[463,581,1348,1370]" pageId="1" pageNumber="570" refString="Shear, W. A. (1990) On the Central and East Asian milliped family Diplomaragnidae (Diplopoda, Chordeumatida, Diplomaragnoidea). American Museum Novitates, 2977, 1 - 40." type="journal article" year="1990">Shear 1990</bibRefCitation>
: 32
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.
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</paragraph>
<paragraph id="8BE636E0FFCC4F0C6FFDFA8B56814837" blockId="1.[151,1333,1284,1400]" box="[151,685,1378,1400]" pageId="1" pageNumber="570">
<treatmentCitationGroup id="AB4911CEFFCC4F0C6FFDFA8B56814837" box="[151,685,1378,1400]" pageId="1" pageNumber="570">
<emphasis id="B92DEAF2FFCC4F0C6FFDFA8B55EA4837" box="[151,454,1378,1400]" italics="true" pageId="1" pageNumber="570">
<taxonomicName id="4C594D63FFCC4F0C6FFDFA8B55004837" ID-CoL="7PNHH" box="[151,300,1379,1400]" class="Diplopoda" family="Diplomaragnidae" genus="Maritimosoma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="genus">Maritimosoma</taxonomicName>
hokkaidense—
</emphasis>
<treatmentCitation id="0AF810F1FFCC4F0C6EACFA8A56864837" author="Mikhaljova" box="[454,682,1378,1400]" page="177" pageId="1" pageNumber="570" year="2000">
<bibRefCitation id="EFC84B11FFCC4F0C6EACFA8A56544837" author="Mikhaljova" box="[454,632,1378,1400]" pageId="1" pageNumber="570" refString="Mikhaljova, E. V. (2000) (for 1999) Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida). Arthropoda Selecta, 8 (3), 153 - 181." type="journal article" year="2000">Mikhaljova 2000</bibRefCitation>
: 177
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.
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</paragraph>
</subSubSection>
<subSubSection id="C343656BFFCC4F0C6FFDFA4C577648F2" box="[151,858,1444,1469]" pageId="1" pageNumber="570" type="distribution">
<paragraph id="8BE636E0FFCC4F0C6FFDFA4C577648F2" blockId="1.[151,1436,1444,1686]" box="[151,858,1444,1469]" pageId="1" pageNumber="570">
<emphasis id="B92DEAF2FFCC4F0C6FFDFA4C551C48F2" bold="true" box="[151,304,1444,1469]" pageId="1" pageNumber="570">Distribution.</emphasis>
<collectingCountry id="F34E7670FFCC4F0C6E5DFA4E555B48F2" box="[311,375,1446,1469]" name="Japan" pageId="1" pageNumber="570">Japan</collectingCountry>
(Hokkaido Island: Rumoi Sub-prefecture).
</paragraph>
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<subSubSection id="C343656BFFCC4F0C6FADFA21511C4BDA" pageId="1" pageNumber="570" type="discussion">
<paragraph id="8BE636E0FFCC4F0C6FADFA21511C4BDA" blockId="1.[151,1436,1444,1686]" pageId="1" pageNumber="570">
<emphasis id="B92DEAF2FFCC4F0C6FADFA21551748AD" bold="true" box="[199,315,1481,1506]" pageId="1" pageNumber="570">Remarks.</emphasis>
The species was originally described in
<taxonomicName id="4C594D63FFCC4F0C6C77FA2157FA48AD" box="[797,982,1481,1506]" class="Diplopoda" family="Diplomaragnidae" genus="Syntelopodeuma" higherTaxonomySource="CoL" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF2FFCC4F0C6C77FA2157FA48AD" box="[797,982,1481,1506]" italics="true" pageId="1" pageNumber="570">Syntelopodeuma</emphasis>
</taxonomicName>
from Hokkaido Island (Rumoi),
<collectingCountry id="F34E7670FFCC4F0C6A36FA2251B048AE" box="[1372,1436,1482,1505]" name="Japan" pageId="1" pageNumber="570">Japan</collectingCountry>
(
<bibRefCitation id="EFC84B11FFCC4F0C6FF4FA04556B4B4A" author="Verhoeff" box="[158,327,1516,1541]" pageId="1" pageNumber="570" refString="Verhoeff, K. W. (1939) Zur Kenntnis ostasiatischer Diplopoden. III. Zoologischer Anzeiger, 127, 113 - 125." type="journal article" year="1939">Verhoeff 1939</bibRefCitation>
), and later transferred to the genus
<taxonomicName id="4C594D63FFCC4F0C6D88FA04500B4B4A" authority="Shear 1990" authorityName="Shear" authorityYear="1990" box="[738,1063,1516,1541]" class="Diplopoda" family="Diplomaragnidae" genus="Diplomaragna" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF2FFCC4F0C6D88FA0457AB4B4A" box="[738,903,1516,1541]" italics="true" pageId="1" pageNumber="570">Diplomaragna</emphasis>
(
<bibRefCitation id="EFC84B11FFCC4F0C6CF2FA0450334B4A" author="Shear" box="[920,1055,1516,1541]" pageId="1" pageNumber="570" refString="Shear, W. A. (1990) On the Central and East Asian milliped family Diplomaragnidae (Diplopoda, Chordeumatida, Diplomaragnoidea). American Museum Novitates, 2977, 1 - 40." type="journal article" year="1990">Shear 1990</bibRefCitation>
)
</taxonomicName>
.
<bibRefCitation id="EFC84B11FFCC4F0C6B5EFA0451264B4A" author="Mikhaljova" box="[1076,1290,1516,1541]" pageId="1" pageNumber="570" refString="Mikhaljova, E. V. (2000) (for 1999) Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida). Arthropoda Selecta, 8 (3), 153 - 181." type="journal article" year="2000">Mikhaljova (2000)</bibRefCitation>
put it in her newly erected genus
<taxonomicName id="4C594D63FFCC4F0C6EEFF9FA56D24B66" authority="Mikhaljova, 2000" authorityName="Mikhaljova" authorityYear="2000" box="[389,766,1553,1578]" class="Diplopoda" family="Diplomaragnidae" genus="Maritimosoma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF2FFCC4F0C6EEFF9FA56064B66" box="[389,554,1554,1577]" italics="true" pageId="1" pageNumber="570">Maritimosoma</emphasis>
<bibRefCitation id="EFC84B11FFCC4F0C6D58F9F956D24B66" author="Mikhaljova" box="[562,766,1553,1578]" pageId="1" pageNumber="570" refString="Mikhaljova, E. V. (2000) (for 1999) Review of the millipede family Diplomaragnidae (Diplopoda: Chordeumatida). Arthropoda Selecta, 8 (3), 153 - 181." type="journal article" year="2000">Mikhaljova, 2000</bibRefCitation>
</taxonomicName>
. However, taking into account the structure of gonopods of this species (especially the lack of posterior gonopod mesal processes, the presence of which is one of the main distinguishing characters of species of
<taxonomicName id="4C594D63FFCC4F0C6D24F9B256DF4B3E" box="[590,755,1626,1649]" class="Diplopoda" family="Diplomaragnidae" genus="Maritimosoma" kingdom="Animalia" order="Chordeumatida" pageId="1" pageNumber="570" phylum="Arthropoda" rank="genus">
<emphasis id="B92DEAF2FFCC4F0C6D24F9B256DF4B3E" box="[590,755,1626,1649]" italics="true" pageId="1" pageNumber="570">Maritimosoma</emphasis>
</taxonomicName>
), morphological similarities with other Hokkaido species of
<emphasis id="B92DEAF2FFCC4F0C6FFDF99555664BD9" box="[151,330,1661,1686]" italics="true" pageId="1" pageNumber="570">Diplopmaragna</emphasis>
and the geographical evidence, here we refer to it again as belonging to
<emphasis id="B92DEAF2FFCC4F0C6B1CF99551054BD9" box="[1142,1321,1661,1686]" italics="true" pageId="1" pageNumber="570">Diplopmaragna</emphasis>
.
</paragraph>
</subSubSection>
</treatment>
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