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<document id="8C528C0E7F5D175E158036EC154E34CF" ID-DOI="10.11646/zootaxa.3994.2.2" ID-GBIF-Dataset="69a42e8b-4d2a-4a8e-ad1a-da6d2fc952aa" ID-ISSN="1175-5326" ID-Zenodo-Dep="243281" ID-ZooBank="41A8AD52-D7E1-4242-87D1-C7819433FA22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461273516540" checkinUser="plazi" docAuthor="Batista, Abel, Ponce, Marcos, Vesely, Milan, Mebert, Konrad, Hertz, Andreas, Köhler, Gunther, Carrizo, Arcadio &amp; Lotzkat, Sebastian" docDate="2015" docId="03F087C8FFCDFF8261A2FF066A7F917B" docLanguage="en" docName="zt03994p221.pdf" docOrigin="Zootaxa 3994 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidoblepharis rufigularis Batista, Ponce, Vesely, Mebert, Hertz, Köhler, Carrizo &amp; Lotzkat, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="201" masterDocId="FFC9FFB0FFC7FF8C6135FF9169259747" masterDocTitle="Revision of the genus Lepidoblepharis (Reptilia: Squamata: Sphaerodactylidae) in Central America, with the description of three new species" masterLastPageNumber="221" masterPageNumber="187" pageNumber="197" updateTime="1698612224987" updateUser="plazi">
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<mods:title id="682422C1ABE81378EE9BB185AFFE8704">Revision of the genus Lepidoblepharis (Reptilia: Squamata: Sphaerodactylidae) in Central America, with the description of three new species</mods:title>
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<mods:namePart id="593925244FCDF056E1DCFEBA86BC12BB">Batista, Abel</mods:namePart>
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<mods:namePart id="F89407AFBAAC62A009CFC56CF4C2655E">Vesely, Milan</mods:namePart>
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<mods:namePart id="074DB3D62C6CDD0AD54BC6D9C75AFAED">Mebert, Konrad</mods:namePart>
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<mods:namePart id="D1539900584AC5B707351A11E5ED3AC1">Hertz, Andreas</mods:namePart>
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<mods:namePart id="DE37BDC77A3528F8B31975DFD01E8B0B">Köhler, Gunther</mods:namePart>
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<mods:namePart id="51B3A7F025E2853A315922EF12718F5E">Carrizo, Arcadio</mods:namePart>
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<mods:namePart id="389C5811F6C8547E10B5CF94BBE2973F">Lotzkat, Sebastian</mods:namePart>
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<heading id="D0AE81B2FFCDFF8661A2FF066B6C97F5" bold="true" box="[151,585,151,178]" fontSize="11" level="1" pageId="10" pageNumber="197" reason="1">
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<taxonomicName id="4C594D5DFFCDFF8661A2FF0668C197F6" box="[151,484,151,177]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="rufigularis" status="sp. nov.">
<emphasis id="B92DEACCFFCDFF8661A2FF0668C197F6" bold="true" box="[151,484,151,177]" italics="true" pageId="10" pageNumber="197">Lepidoblepharis rufigularis</emphasis>
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<paragraph id="8BE636DEFFCDFF8661A2FF2B680A9794" blockId="10.[151,585,151,211]" box="[151,303,186,211]" pageId="10" pageNumber="197">
<figureCitation id="13622A5BFFCDFF8661A2FF2B68249794" box="[151,257,186,211]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="11.[151,250,1918,1940]" captionStartId-1="12.[151,250,792,814]" captionStartId-2="13.[151,250,1562,1584]" captionTargetBox-0="[244,1340,193,1892]" captionTargetBox-1="[150,1436,193,770]" captionTargetBox-2="[151,1436,179,1539]" captionTargetId-0="figure@11.[244,1340,193,1893]" captionTargetId-1="figure@12.[150,1436,193,771]" captionTargetId-2="figure@13.[151,1436,179,1541]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." captionText-1="FIGURE 4. Feet and hands of Panamanian Lepidoblepharis. Lepidoblepharis emberawoundule: right foot of SMF 81954 and right hand of MHCH 2957; L. rufigularis (SMF 50659): right foot and right hand; L. sanctaemartae: left foot of SMF 97419 and right hand of MHCH 2948; L. victormartinezi: left foot and left hand of SMF 89963; L. xanthostigma: left foot of SMF 90190 and left hand of SMF 89576. Scale bars equal 1 mm, tip of fourth finger and toe marked with an asterisk." captionText-2="FIGURE 5. Escutcheon scales in males of Panamanian Lepidoblepharis. (A) L. emberawoundule: ventral escutcheon with 46 scales, 7 scales long x 11 wide, and subfemoral escutcheon with 4 scales per side, of SMF 81951 (SVL = 25 mm); (B) L. rufigularis: ventral escutcheon with 62 scales, 7 scales long x 13 wide, and subfemoral escutcheon with 3 and 4 scales per side, of SMF 50659 (SVL = 25 mm); (C) L. sanctaemartae: ventral escutcheon of SMF 97419 (SVL = 22 mm) with 92 scales, 10 scales long x 12 wide; (D) L. victormartinezi: ventral escutcheon of SMF 89963 (SVL = 27 mm) comprising 57 intact scales (an original total of 61 is assumed considering the symmetrical scale arrangement of the non-damaged portions), 7 scales long x 14 wide; (E F) L. xanthostigma: (E) ventral escutcheon with 75 scales, 12 scales long x 11 wide, and subfemoral escutcheon with 2 scales per side (no additional scales hidden under strings), of SMF 90189 (SVL = 35 mm), and (F) ventral escutcheon with 36 scales, 6 scales long x 8 wide, and no subfemoral escutcheon of AB 1250 (SVL = 24 mm). All scale bars equal 1 mm." httpUri-0="https://zenodo.org/record/243283/files/figure.png" httpUri-1="https://zenodo.org/record/243284/files/figure.png" httpUri-2="https://zenodo.org/record/243285/files/figure.png" pageId="10" pageNumber="197">Figs. 35</figureCitation>
,
<figureCitation id="13622A5BFFCDFF866038FF2A680E9794" box="[269,299,187,211]" captionStart="FIGURE 11" captionStartId="23.[151,250,852,874]" captionTargetBox="[188,1398,181,830]" captionTargetId="figure@23.[188,1398,181,831]" captionTargetPageId="23" captionText="FIGURE 11. Holotype of Lepidoblepharis rufigularis (male SMF 50659) in life. (A B) lateral view; (C) gular and chest region." httpUri="https://zenodo.org/record/243291/files/figure.png" pageId="10" pageNumber="197">11</figureCitation>
.
</paragraph>
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<paragraph id="8BE636DEFFCDFF8661A2FE936AB49670" blockId="10.[151,1436,258,311]" pageId="10" pageNumber="197">
<emphasis id="B92DEACCFFCDFF8661A2FE9368F3965F" box="[151,470,258,280]" italics="true" pageId="10" pageNumber="197">
<taxonomicName id="4C594D5DFFCDFF8661A2FE9368F5965F" box="[151,464,258,280]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="xanthostigma">Lepidoblepharis xanthostigma</taxonomicName>
:
</emphasis>
all in part. (referring to populations in extreme southeastern
<collectingCountry id="F34E764EFFCDFF86657EFE926D84965F" box="[1099,1185,259,280]" name="Panama" pageId="10" pageNumber="197">Panama</collectingCountry>
):
<bibRefCitation id="EFC84B2FFFCDFF866586FE936C11965F" author="Auth" box="[1203,1332,258,280]" pageId="10" pageNumber="197" refString="Auth, D. L. (1994) Checklist and bibliography of the amphibians and reptiles of Panama. Smithsonian Herpetological Information Service, 98, 1 - 59. http: // dx. doi. org / 10.5479 / si. 23317515.98.1" type="journal article" year="1994">Auth (1994)</bibRefCitation>
;
<bibRefCitation id="EFC84B2FFFCDFF866475FE92680B9670" pageId="10" pageNumber="197" refString="Young, B. E., Sedaghatkish, G., Roca, E. &amp; Fuenmayor, Q. D. (1999) El estatus de la conservacion de la herpetofauna de Panama. Resumen del primer taller internacional sobre la herpetofauna de Panama. The Nature Conservancy y Asociacion Nacional para la Conservacion de la Naturaleza (ANCON), Arlington, Virginia, 40 pp." type="book">
Young
<emphasis id="B92DEACCFFCDFF8664BFFE9269C79670" italics="true" pageId="10" pageNumber="197">et al.</emphasis>
(1999)
</bibRefCitation>
; Ibáñez
<emphasis id="B92DEACCFFCDFF8660B7FEB368909670" box="[386,437,289,311]" italics="true" pageId="10" pageNumber="197">et al.</emphasis>
(2001);
<bibRefCitation id="EFC84B2FFFCDFF866339FEB06BBB9670" author="Kohler" box="[524,670,289,311]" pageId="10" pageNumber="197" refString="Kohler, G. (2008) Reptiles of Central America. Second edition. Herpeton, Offenbach, 400 pp." type="book" year="2008">Köhler (2008)</bibRefCitation>
;
<bibRefCitation id="EFC84B2FFFCDFF86639CFEB06AA89670" box="[681,909,289,311]" pageId="10" pageNumber="197" refString="Jaramillo A., C. A., Wilson, L. D., Ibanez D., R. &amp; Jaramillo, F. E. (2010) The herpetofauna of Panama: distribution and conservation status. In: Wilson, L. D., Townsend, J. H., Johnson, J. D. &amp; Murphy, J. B. (Eds.) Conservation of Mesoamerican amphibians and reptiles. Eagle Mountain Press, Eagle Mountain, Utah, pp. 604 - 671." type="book chapter">
Jaramillo
<emphasis id="B92DEACCFFCDFF866238FEB36A649670" box="[781,833,289,311]" italics="true" pageId="10" pageNumber="197">et al.</emphasis>
(2010)
</bibRefCitation>
.
</paragraph>
</subSubSection>
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<emphasis id="B92DEACCFFCDFF8661A2FEF5682E963A" bold="true" box="[151,267,356,381]" pageId="10" pageNumber="197">
<typeStatus id="54E2887CFFCDFF8661A2FEF56823963A" box="[151,262,356,381]" pageId="10" pageNumber="197" type="holotype">Holotype</typeStatus>
.
</emphasis>
Adult male
<collectionCode id="ED48AE1BFFCDFF8660A2FEF568EA963B" LSID="urn:lsid:biocol.org:col:34838" box="[407,463,356,380]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="10" pageNumber="197">SMF</collectionCode>
50659, original field number AB 527 (
<figureCitation id="13622A5BFFCDFF8662BDFEF56AD1963A" box="[904,1012,356,381]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStart-2="FIGURE 5" captionStartId-0="11.[151,250,1918,1940]" captionStartId-1="12.[151,250,792,814]" captionStartId-2="13.[151,250,1562,1584]" captionTargetBox-0="[244,1340,193,1892]" captionTargetBox-1="[150,1436,193,770]" captionTargetBox-2="[151,1436,179,1539]" captionTargetId-0="figure@11.[244,1340,193,1893]" captionTargetId-1="figure@12.[150,1436,193,771]" captionTargetId-2="figure@13.[151,1436,179,1541]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." captionText-1="FIGURE 4. Feet and hands of Panamanian Lepidoblepharis. Lepidoblepharis emberawoundule: right foot of SMF 81954 and right hand of MHCH 2957; L. rufigularis (SMF 50659): right foot and right hand; L. sanctaemartae: left foot of SMF 97419 and right hand of MHCH 2948; L. victormartinezi: left foot and left hand of SMF 89963; L. xanthostigma: left foot of SMF 90190 and left hand of SMF 89576. Scale bars equal 1 mm, tip of fourth finger and toe marked with an asterisk." captionText-2="FIGURE 5. Escutcheon scales in males of Panamanian Lepidoblepharis. (A) L. emberawoundule: ventral escutcheon with 46 scales, 7 scales long x 11 wide, and subfemoral escutcheon with 4 scales per side, of SMF 81951 (SVL = 25 mm); (B) L. rufigularis: ventral escutcheon with 62 scales, 7 scales long x 13 wide, and subfemoral escutcheon with 3 and 4 scales per side, of SMF 50659 (SVL = 25 mm); (C) L. sanctaemartae: ventral escutcheon of SMF 97419 (SVL = 22 mm) with 92 scales, 10 scales long x 12 wide; (D) L. victormartinezi: ventral escutcheon of SMF 89963 (SVL = 27 mm) comprising 57 intact scales (an original total of 61 is assumed considering the symmetrical scale arrangement of the non-damaged portions), 7 scales long x 14 wide; (E F) L. xanthostigma: (E) ventral escutcheon with 75 scales, 12 scales long x 11 wide, and subfemoral escutcheon with 2 scales per side (no additional scales hidden under strings), of SMF 90189 (SVL = 35 mm), and (F) ventral escutcheon with 36 scales, 6 scales long x 8 wide, and no subfemoral escutcheon of AB 1250 (SVL = 24 mm). All scale bars equal 1 mm." httpUri-0="https://zenodo.org/record/243283/files/figure.png" httpUri-1="https://zenodo.org/record/243284/files/figure.png" httpUri-2="https://zenodo.org/record/243285/files/figure.png" pageId="10" pageNumber="197">Figs. 35</figureCitation>
,
<figureCitation id="13622A5BFFCDFF866535FEF56D38963B" box="[1024,1053,356,380]" captionStart="FIGURE 11" captionStartId="23.[151,250,852,874]" captionTargetBox="[188,1398,181,830]" captionTargetId="figure@23.[188,1398,181,831]" captionTargetPageId="23" captionText="FIGURE 11. Holotype of Lepidoblepharis rufigularis (male SMF 50659) in life. (A B) lateral view; (C) gular and chest region." httpUri="https://zenodo.org/record/243291/files/figure.png" pageId="10" pageNumber="197">11</figureCitation>
), collected on a hill
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north of Río Púcuro (
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,
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,
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elev.), Pinogana, Darién,
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, on 0
<date id="FFE7101EFFCDFF86655AFE196DD596E7" box="[1135,1264,391,416]" pageId="10" pageNumber="197" value="2012-07-08">8 July 2012</date>
at 22:40 hrs by Abel Batista.
</paragraph>
</subSubSection>
<subSubSection id="C3436555FFCDFF8661F2FE5E6DDB91CA" pageId="10" pageNumber="197" type="diagnosis">
<paragraph id="8BE636DEFFCDFF8661F2FE5E6DA99440" blockId="10.[151,1437,356,2037]" pageId="10" pageNumber="197">
<emphasis id="B92DEACCFFCDFF8661F2FE5E686496AF" bold="true" box="[199,321,463,488]" pageId="10" pageNumber="197">Diagnosis.</emphasis>
<taxonomicName id="4C594D5DFFCDFF866078FE416BAD96AF" box="[333,648,464,488]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCDFF866078FE416BAD96AF" box="[333,648,464,488]" italics="true" pageId="10" pageNumber="197">Lepidoblepharis rufigularis</emphasis>
</taxonomicName>
(our
<emphasis id="B92DEACCFFCDFF8663FAFE5E6A0B96AF" bold="true" box="[719,814,463,488]" pageId="10" pageNumber="197">
sp.
<date id="FFE7101EFFCDFF8663C8FE5E6A0B96AF" box="[765,814,463,488]" pageId="10" pageNumber="197">nov.</date>
</emphasis>
2) is characterized by the following combination of characters:
<emphasis id="B92DEACCFFCDFF86602CFE65681F954B" box="[281,314,500,524]" italics="true" pageId="10" pageNumber="197">(1)</emphasis>
dorsal scales small, granular, and juxtaposed, ventral scales large, cycloid, flat, and imbricate;
<emphasis id="B92DEACCFFCDFF86644EFE656CB9954B" box="[1403,1436,500,524]" italics="true" pageId="10" pageNumber="197">(2)</emphasis>
scales on head small and granular;
<emphasis id="B92DEACCFFCDFF86632AFD896B1A9577" box="[543,575,536,560]" italics="true" pageId="10" pageNumber="197">(3)</emphasis>
four postrostral scales;
<emphasis id="B92DEACCFFCDFF86627EFD896A499577" box="[843,876,536,560]" italics="true" pageId="10" pageNumber="197">(4)</emphasis>
two short, barely discernible paramedian clefts in the more or less U-shaped posterior mental border;
<emphasis id="B92DEACCFFCDFF8663CCFDAD6A3F9514" box="[761,794,572,595]" italics="true" pageId="10" pageNumber="197">(5)</emphasis>
five postmentals, the two median ones larger than the posteriorly adjacent chin scales;
<emphasis id="B92DEACCFFCDFF866321FDF16B10953F" box="[532,565,608,632]" italics="true" pageId="10" pageNumber="197">(6)</emphasis>
13 lamellae under fourth toe, 11 lamellae under fourth finger;
<emphasis id="B92DEACCFFCDFF86642BFDF16C1A9530" box="[1310,1343,608,631]" italics="true" pageId="10" pageNumber="197">(7)</emphasis>
median subcaudals conspicuously wider than long, almost twice as wide as the laterally adjacent scales, with straight posterior margins arranged in a regular tail sequence of 1'1'';
<emphasis id="B92DEACCFFCDFF866270FD396A439587" box="[837,870,680,704]" italics="true" pageId="10" pageNumber="197">(8)</emphasis>
ventral escutcheon consisting of 62 scales, almost twice as wide (13 scales) as long (7 scales);
<emphasis id="B92DEACCFFCDFF8663B8FD5D6B8A95A3" box="[653,687,716,740]" italics="true" pageId="10" pageNumber="197">(9)</emphasis>
subfemoral escutcheon consisting of 34 scales per thigh;
<emphasis id="B92DEACCFFCDFF86647CFD5D6C5C95A3" box="[1353,1401,716,740]" italics="true" pageId="10" pageNumber="197">(10)</emphasis>
17 longitudinal rows of ventral scales at midbody;
<emphasis id="B92DEACCFFCDFF866398FD616BFF944F" box="[685,730,752,776]" italics="true" pageId="10" pageNumber="197">(11)</emphasis>
bilobate hemipenis;
<emphasis id="B92DEACCFFCDFF8662F1FD616AD6944F" box="[964,1011,752,776]" italics="true" pageId="10" pageNumber="197">(12)</emphasis>
SVL
<quantity id="4CA19B3BFFCDFF866500FD7E6DA29440" box="[1077,1159,751,776]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="197" unit="mm" value="25.0">25 mm</quantity>
.
</paragraph>
<paragraph id="8BE636DEFFCDFF8661F2FC856DDB91CA" blockId="10.[151,1437,356,2037]" pageId="10" pageNumber="197">
<emphasis id="B92DEACCFFCDFF8661F2FC856BD5946A" bold="true" box="[199,752,788,813]" pageId="10" pageNumber="197">Comparison with other species of the genus.</emphasis>
<taxonomicName id="4C594D5DFFCDFF8663C9FC856D12946B" box="[764,1079,788,812]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCDFF8663C9FC856D12946B" box="[764,1079,788,812]" italics="true" pageId="10" pageNumber="197">Lepidoblepharis rufigularis</emphasis>
</taxonomicName>
can be differentiated from all species in the genus by its small size, number of lamellae under the fourth toe and finger, the reddish throat in males (
<figureCitation id="13622A5BFFCDFF8661DCFCCD68189433" box="[233,317,860,885]" captionStart="FIGURE 11" captionStartId="23.[151,250,852,874]" captionTargetBox="[188,1398,181,830]" captionTargetId="figure@23.[188,1398,181,831]" captionTargetPageId="23" captionText="FIGURE 11. Holotype of Lepidoblepharis rufigularis (male SMF 50659) in life. (A B) lateral view; (C) gular and chest region." httpUri="https://zenodo.org/record/243291/files/figure.png" pageId="10" pageNumber="197">Fig. 11</figureCitation>
), and the configuration of the ventral escutcheon. In the following, we present comparisons to all other species within the genus, with the characteristics for
<taxonomicName id="4C594D5DFFCDFF866242FC106D3094DF" box="[887,1045,896,920]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCDFF866242FC106D3094DF" box="[887,1045,896,920]" italics="true" pageId="10" pageNumber="197">L. rufigularis</emphasis>
</taxonomicName>
in parentheses.
<taxonomicName id="4C594D5DFFCDFF8665D1FC11682A94FB" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCDFF8665D1FC11682A94FB" italics="true" pageId="10" pageNumber="197">Lepidoblepharis rufigularis</emphasis>
</taxonomicName>
can be distinguished from the Panamanian species
<taxonomicName id="4C594D5DFFCDFF866289FC346D5B94FB" box="[956,1150,932,956]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="xanthostigma">
<emphasis id="B92DEACCFFCDFF866289FC346D5B94FB" box="[956,1150,932,956]" italics="true" pageId="10" pageNumber="197">L. xanthostigma</emphasis>
</taxonomicName>
,
<emphasis id="B92DEACCFFCDFF8665AFFC34687A94A7" italics="true" pageId="10" pageNumber="197">
<taxonomicName id="4C594D5DFFCDFF8665AFFC346C4894FB" box="[1178,1389,933,956]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="sanctaemartae">L. sanctaemartae</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF8664B0FC34687E94A7" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="emberawoundule">L. emberawoundule</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C594D5DFFCDFF8660ACFC596B7494A7" box="[409,593,968,992]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="genus" status="sp. nov.">
<emphasis id="B92DEACCFFCDFF8660ACFC596B7494A7" box="[409,593,968,992]" italics="true" pageId="10" pageNumber="197">Lepidoblepharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E57B7FFCDFF86636CFC566B9094A7" box="[601,693,967,992]" pageId="10" pageNumber="197" rank="species">
<emphasis id="B92DEACCFFCDFF86636CFC566B9094A7" bold="true" box="[601,693,967,992]" pageId="10" pageNumber="197">
sp.
<date id="FFE7101EFFCDFF8663B1FC566B9094A7" box="[644,693,967,992]" pageId="10" pageNumber="197">nov.</date>
</emphasis>
</taxonomicNameLabel>
3 (described below) by a genetic p-distance of 1423% between individuals in 16S mtDNA.
<taxonomicName id="4C594D5DFFCDFF8660E1FC7D6A0F9343" box="[468,810,1004,1028]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="xanthostigma">
<emphasis id="B92DEACCFFCDFF8660E1FC7D6A0F9343" box="[468,810,1004,1028]" italics="true" pageId="10" pageNumber="197">Lepidoblepharis xanthostigma</emphasis>
</taxonomicName>
is the most similar species, but has a different chin and throat coloration with dark reticulations on a pale background (orange background), and greatly enlarged median subcaudal scales which are more than two times as wide as the laterally adjacent subcaudal scales (enlarged but less than two times the width of laterally adjacent subcaudals,
<figureCitation id="13622A5BFFCDFF866251FBC66A8C9337" box="[868,937,1111,1136]" captionStart="FIGURE 3" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[244,1340,193,1892]" captionTargetId="figure@11.[244,1340,193,1893]" captionTargetPageId="11" captionText="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." httpUri="https://zenodo.org/record/243283/files/figure.png" pageId="10" pageNumber="197">Fig. 3</figureCitation>
), usually 21 or more, very rarely 18, scales across snout (19), usually 18 or more, rarely 16 or 17, ventral scales a midbody (17), and an escutcheon long/wide ratio of 67120% (54%).
<emphasis id="B92DEACCFFCDFF8660F1FB316D0393FF" box="[452,1062,1184,1208]" italics="true" pageId="10" pageNumber="197">
<taxonomicName id="4C594D5DFFCDFF8660F1FB316A6F93FF" box="[452,842,1184,1208]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="emberawoundule">Lepidoblepharis emberawoundule</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF86626CFB306D0793FF" box="[857,1058,1185,1208]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="sanctaemartae">L. sanctaemartae</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C594D5DFFCDFF866551FB316C3993FF" box="[1124,1308,1184,1208]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="genus" status="sp. nov.">
<emphasis id="B92DEACCFFCDFF866551FB316C3993FF" box="[1124,1308,1184,1208]" italics="true" pageId="10" pageNumber="197">Lepidoblepharis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21E57B7FFCDFF866413FB0E6CA693FF" box="[1318,1411,1183,1208]" pageId="10" pageNumber="197" rank="species">
<emphasis id="B92DEACCFFCDFF866413FB0E6CA693FF" bold="true" box="[1318,1411,1183,1208]" pageId="10" pageNumber="197">
sp.
<date id="FFE7101EFFCDFF866467FB0E6CA693FF" box="[1362,1411,1183,1208]" pageId="10" pageNumber="197">nov.</date>
</emphasis>
</taxonomicNameLabel>
3 (described below) have fewer than 10 lamellae under the fourth toe (13) and under the fourth finger (11). Additionally,
<taxonomicName id="4C594D5DFFCDFF866006FB7868D29247" box="[307,503,1257,1280]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="sanctaemartae">
<emphasis id="B92DEACCFFCDFF866006FB7868D29247" box="[307,503,1257,1280]" italics="true" pageId="10" pageNumber="197">L. sanctaemartae</emphasis>
</taxonomicName>
has large, flat, imbricate dorsal body scales (small, granular, and juxtaposed). To date, seven species of the genus
<taxonomicName id="4C594D5DFFCDFF866324FA9D6BED9263" box="[529,712,1292,1316]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="genus">
<emphasis id="B92DEACCFFCDFF866324FA9D6BED9263" box="[529,712,1292,1316]" italics="true" pageId="10" pageNumber="197">Lepidoblepharis</emphasis>
</taxonomicName>
have been reported to possess ten or fewer lamellae under the fourth toe,
<emphasis id="B92DEACCFFCDFF866022FAA168659200" box="[279,320,1328,1351]" italics="true" pageId="10" pageNumber="197">i.e.,</emphasis>
to be short-toed (
<taxonomicName id="4C594D5DFFCDFF86633AFAA06BA69200" box="[527,643,1328,1352]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="miyatai">
<emphasis id="B92DEACCFFCDFF86633AFAA06BA69200" box="[527,643,1328,1352]" italics="true" pageId="10" pageNumber="197">L. miyatai</emphasis>
</taxonomicName>
,
<emphasis id="B92DEACCFFCDFF8663A6FAA06821922B" italics="true" pageId="10" pageNumber="197">
<taxonomicName id="4C594D5DFFCDFF8663A6FAA06A7E920F" box="[659,859,1329,1352]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="sanctaemartae">L. sanctaemartae</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF86625DFAA06ADA920F" box="[872,1023,1328,1352]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="buchwaldi">L. buchwaldi</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF866539FAA06DC79200" box="[1036,1250,1328,1352]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="montecanoensis">L. montecanoensis</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF8665DAFAA06C5D920F" box="[1263,1400,1328,1352]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="williamsi">L. williamsi</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF8664B0FAA06824922B" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="peraccae">L. peraccae</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C594D5DFFCDFF86600EFAC468E8922B" box="[315,461,1364,1388]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="microlepis">
<emphasis id="B92DEACCFFCDFF86600EFAC468E8922B" box="[315,461,1364,1388]" italics="true" pageId="10" pageNumber="197">L. microlepis</emphasis>
</taxonomicName>
), and are therefore readily differentiable from the long-toed
<taxonomicName id="4C594D5DFFCDFF866541FAC46C2F922B" box="[1140,1290,1364,1388]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCDFF866541FAC46C2F922B" box="[1140,1290,1364,1388]" italics="true" pageId="10" pageNumber="197">L. rufigularis</emphasis>
</taxonomicName>
(13 lamellae under the fourth toe). Of the remaining members of the genus,
<emphasis id="B92DEACCFFCDFF86626CFAE86A2A92F3" italics="true" pageId="10" pageNumber="197">
<taxonomicName id="4C594D5DFFCDFF86626CFAE86D2D92D7" box="[857,1032,1400,1424]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="colombianus">L. colombianus</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF866521FAE86D8592D7" box="[1044,1184,1400,1424]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="conolepis">L. conolepis</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF86659EFAE86C0E92D7" box="[1195,1323,1400,1424]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="duolepis">L. duolepis</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF866403FAE86CBD92D7" box="[1334,1432,1400,1424]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="festae">L. festae</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF8661A2FA0C682892F3" box="[151,269,1436,1460]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="grandis">L. grandis</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF86602CFA0C689792F3" box="[281,434,1436,1460]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="heyerorum">L. heyerorum</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF86608BFA0C6B7F92F3" box="[446,602,1436,1460]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="hoogmoedi">L. hoogmoedi</taxonomicName>
,
<taxonomicName id="4C594D5DFFCDFF866353FA0C6A2E92F3" box="[614,779,1436,1460]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="intermedius">L. intermedius</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C594D5DFFCDFF866272FA0C6AE492F3" box="[839,961,1436,1460]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="ruthveni">
<emphasis id="B92DEACCFFCDFF866272FA0C6AE492F3" box="[839,961,1436,1460]" italics="true" pageId="10" pageNumber="197">L. ruthveni</emphasis>
</taxonomicName>
are relatively to very large lizards for this genus with adult SVLs between 33 and
<quantity id="4CA19B3BFFCDFF866353FA2E6B9D9290" box="[614,696,1471,1496]" metricMagnitude="-2" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="197" unit="mm" value="56.0">56 mm</quantity>
(
<quantity id="4CA19B3BFFCDFF8663FCFA2E6A05929F" box="[713,800,1471,1496]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="197" unit="mm" value="25.0">25 mm</quantity>
). Additionally,
<taxonomicName id="4C594D5DFFCDFF8662E4FA506D78929F" box="[977,1117,1472,1496]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="conolepis">
<emphasis id="B92DEACCFFCDFF8662E4FA506D78929F" box="[977,1117,1472,1496]" italics="true" pageId="10" pageNumber="197">L. conolepis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C594D5DFFCDFF8665ADFA506C2B929F" box="[1176,1294,1472,1496]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B92DEACCFFCDFF8665ADFA506C2B929F" box="[1176,1294,1472,1496]" italics="true" pageId="10" pageNumber="197">L. grandis</emphasis>
</taxonomicName>
have 1420 lamellae under the fourth toe (13). The dorsal ground color in males of
<taxonomicName id="4C594D5DFFCDFF8662FCFA746D4592BB" box="[969,1120,1508,1532]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="heyerorum">
<emphasis id="B92DEACCFFCDFF8662FCFA746D4592BB" box="[969,1120,1508,1532]" italics="true" pageId="10" pageNumber="197">L. heyerorum</emphasis>
</taxonomicName>
is black with yellow dorsal markings (no yellow dorsal markings). The two long-toed specimens with granular dorsals reported as
<taxonomicName id="4C594D5DFFCDFF8664B0F998680B9103" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="xanthostigma">
<emphasis id="B92DEACCFFCDFF8664B0F998680B9103" italics="true" pageId="10" pageNumber="197">L. xanthostigma</emphasis>
</taxonomicName>
from
<collectingCountry id="F34E764EFFCDFF866044F9BD68FA9102" box="[369,479,1580,1605]" name="Colombia" pageId="10" pageNumber="197">Colombia</collectingCountry>
by
<bibRefCitation id="EFC84B2FFFCDFF86633DF9BD6A209103" author="Ayala" box="[520,773,1580,1605]" pageId="10" pageNumber="197" refString="Ayala, S. C. &amp; Castro, F. (1983) Dos nuevos gecos (Sauria: Gekkonidae, Sphaerodactylinae) para Colombia: Lepidoblepharis xanthostigma (Noble) y descripcion de una nueva especie. Caldasia, 13, 743 - 753." type="journal article" year="1983">Ayala &amp; Castro (1983)</bibRefCitation>
are similar to
<taxonomicName id="4C594D5DFFCDFF866291F9BC6D1C9103" box="[932,1081,1580,1604]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="10" pageNumber="197" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCDFF866291F9BC6D1C9103" box="[932,1081,1580,1604]" italics="true" pageId="10" pageNumber="197">L. rufigularis</emphasis>
</taxonomicName>
in the number of ventral scales, but they have 2225 scales across snout (19), an escutcheon with only 25 scales (62), the gular region with blotches (gular region with longitudinal bars), and an occipital pale W-shaped mark (no occipital mark at all).
</paragraph>
</subSubSection>
<subSubSection id="C3436555FFCDFF8261F2F9066C0293BF" lastPageId="14" lastPageNumber="201" pageId="10" pageNumber="197" type="description">
<paragraph id="8BE636DEFFCDFF8061F2F90668D69041" blockId="10.[151,1437,356,2037]" lastBlockId="12.[151,1437,945,2014]" lastPageId="12" lastPageNumber="199" pageId="10" pageNumber="197">
<emphasis id="B92DEACCFFCDFF8661F2F9066B0191F7" bold="true" box="[199,548,1687,1712]" pageId="10" pageNumber="197">
Description of the
<typeStatus id="54E2887CFFCDFF866083F9066B3A91F7" box="[438,543,1687,1712]" pageId="10" pageNumber="197" type="holotype">holotype</typeStatus>
.
</emphasis>
Adult male as judged by everted hemipenes; SVL
<quantity id="4CA19B3BFFCDFF8665ACF9066DD591E8" box="[1177,1264,1687,1712]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="10" pageNumber="197" unit="mm" value="25.0">25 mm</quantity>
, TL 33.0 mm (measured while the now broken tail was still intact), HL
<quantity id="4CA19B3BFFCDFF86622AF92D6A529193" box="[799,887,1724,1749]" metricMagnitude="-3" metricUnit="m" metricValue="5.6" pageId="10" pageNumber="197" unit="mm" value="5.6">5.6 mm</quantity>
, HW
<quantity id="4CA19B3BFFCDFF86628CF92D6D349193" box="[953,1041,1724,1748]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" pageId="10" pageNumber="197" unit="mm" value="4.1">4.1 mm</quantity>
, forelimbs 5.0 mm, hind limbs 9.3, shank
<quantity id="4CA19B3BFFCDFF8661EAF94E681D91B0" box="[223,312,1759,1784]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="10" pageNumber="197" unit="mm" value="3.6">3.6 mm</quantity>
; rostral large, clearly visible from above, with a shallow, horseshoe-shaped posterior depression and a long median cleft; postrostrals four, including supranasals, one median postrostral slightly larger than posteriorly adjacent scales on snout and indenting the rostral; postnasals two, both about the same size as posteriorly adjacent loreal scales; scales on snout rounded and pointed backward; loreal scales subimbricate, elevated toward posterior and dorsal directions, six loreal scales on a longitudinal line between postnasals and orbit; 19 scales across snout between anterior sutures of second SPLs; scales on top of head small, granular, juxtaposed, generally pointing upward, about half the size of those on the middle area of snout; superciliary flap with two enlarged scales on anterior border, of which the first is slightly longer than the second, followed by 25 small and globular scales; supralabials three, posteriormost one below center of eye; ear-opening small, oval, in oblique orientation; mental large, posterior margin somewhat U-shaped, with two barely discernible, very short paramedian clefts; five enlarged postmentals, slightly larger than the posteriorly adjacent chin scales (
<figureCitation id="13622A5BFFCBFF80653CFC686D699355" box="[1033,1100,1017,1042]" captionStart="FIGURE 3" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[244,1340,193,1892]" captionTargetId="figure@11.[244,1340,193,1893]" captionTargetPageId="11" captionText="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." httpUri="https://zenodo.org/record/243283/files/figure.png" pageId="12" pageNumber="199">Fig. 3</figureCitation>
B); scales on chin small, flat, rounded, and juxtaposed, on posterior region flat to flat-granular, slightly inclined backward; scales near posterior infralabials flat, subimbricate, and larger than scales in median area of chin; infralabials four, first largest and almost reaching anterior level of orbit, fourth below center of eye; throat with small granular scales, pointed and directed upward; dorsal scales on neck and body small, granular, and juxtaposed, at midbody and on posterior trunk pointed, granular, or, in frontal view, triangular, mostly directed posteriorly, some scales on flanks and in lumbar region pointed; dorsal skin on body too damaged to conduct longitudinal counts of middorsal scales; ventral scales flat, smooth, imbricate, with an ovoid posterior margin, increasing moderately in size from gular region to belly, posterior ventral scales longer than wide; 13 midventral scales in one HL, 23 between levels of axilla and groin, 28 to border of cloaca; ventral escutcheon patch with 62 scales, 7 long and 13 wide, escutcheon long/wide ratio 54%; subfemoral escutcheon with 3 scales in a single row under each thigh, and a barely discernible fourth scale in a second row under the right thigh; transition between ventrals and lateral scales abrupt; 17 longitudinal rows of ventrals at midbody; scales on precloacal plate similar to ventrals, except for those on border of cloaca, which are smaller; tail dorsally and laterally with flattened, smooth, and imbricate scales that are less elongate than ventrals (with a transitional zone at base of tail); underside of tail with a median row of moderately enlarged scales (usually no more than two times wider than the laterally adjacent subcaudal scales;
<figureCitation id="13622A5BFFCBFF8062D7F9846D019169" box="[994,1060,1557,1582]" captionStart="FIGURE 3" captionStartId="11.[151,250,1918,1940]" captionTargetBox="[244,1340,193,1892]" captionTargetId="figure@11.[244,1340,193,1893]" captionTargetPageId="11" captionText="FIGURE 3. Morphological distinction among Lepidoblepharis from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A) L. emberawoundule; (B) L. rufigularis; (C) L. sanctaemartae; (D) L victormartinezi; (E) L. xanthostigma." httpUri="https://zenodo.org/record/243283/files/figure.png" pageId="12" pageNumber="199">Fig. 3</figureCitation>
B), mostly with a repeated series of one median scale bordered laterodistally by one scale, followed by a slightly larger median scale in contact laterodistally with two scales, constituting a regular tail sequence of 1'1''; dorsal scales on forelimbs granular to conical; scales on hind limbs flat, smooth, and imbricate on anteroventral femoral and shank surfaces, granular elsewhere; fingers, from longest to shortest, IV-III-II-V-I; toes IV-III-II-V-I, fourth and third toes about the same length; 11 lamellae under fourth finger, 13 under fourth toe; claws enclosed by an ungual sheath composed of six scales, as typical for the genus.
</paragraph>
<caption id="DF266656FFCCFF8761A2F8EF6BCA9095" httpUri="https://zenodo.org/record/243283/files/figure.png" pageId="11" pageNumber="198" targetBox="[244,1340,193,1892]" targetPageId="11">
<paragraph id="8BE636DEFFCCFF8761A2F8EF6BCA9095" blockId="11.[151,1437,1918,2002]" pageId="11" pageNumber="198">
<emphasis id="B92DEACCFFCCFF8761A2F8EF683D90D3" bold="true" box="[151,280,1918,1940]" pageId="11" pageNumber="198">FIGURE 3.</emphasis>
Morphological distinction among
<taxonomicName id="4C594D5DFFCCFF8763B9F8EF6A1790D3" box="[652,818,1918,1940]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="198" phylum="Chordata" rank="genus">
<emphasis id="B92DEACCFFCCFF8763B9F8EF6A1790D3" box="[652,818,1918,1940]" italics="true" pageId="11" pageNumber="198">Lepidoblepharis</emphasis>
</taxonomicName>
from Lower Central America; images from left to right: dorsals, ventral views of foot, tail, and mental region; scale bars equal 1 mm. (A)
<taxonomicName id="4C594D5DFFCCFF8762FBF80F6DB290F4" box="[974,1175,1949,1971]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="198" phylum="Chordata" rank="species" species="emberawoundule">
<emphasis id="B92DEACCFFCCFF8762FBF80F6DB290F4" box="[974,1175,1949,1971]" italics="true" pageId="11" pageNumber="198">L. emberawoundule</emphasis>
</taxonomicName>
; (B)
<taxonomicName id="4C594D5DFFCCFF8765F9F80F6C7790F4" box="[1228,1362,1949,1971]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="198" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCCFF8765F9F80F6C7790F4" box="[1228,1362,1949,1971]" italics="true" pageId="11" pageNumber="198">L. rufigularis</emphasis>
</taxonomicName>
; (C)
<taxonomicName id="4C594D5DFFCCFF8764B2F80F68099095" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="198" phylum="Chordata" rank="species" species="sanctaemartae">
<emphasis id="B92DEACCFFCCFF8764B2F80F68099095" italics="true" pageId="11" pageNumber="198">L. sanctaemartae</emphasis>
</taxonomicName>
; (D)
<taxonomicName id="4C594D5DFFCCFF876056F82C6B379095" box="[355,530,1981,2002]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="198" phylum="Chordata" rank="species" species="victormartinezi">
<emphasis id="B92DEACCFFCCFF876056F82C6B379095" box="[355,530,1981,2002]" italics="true" pageId="11" pageNumber="198">L victormartinezi</emphasis>
</taxonomicName>
; (E)
<taxonomicName id="4C594D5DFFCCFF876373F82C6BCD9095" box="[582,744,1980,2002]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="11" pageNumber="198" phylum="Chordata" rank="species" species="xanthostigma">
<emphasis id="B92DEACCFFCCFF876373F82C6BCD9095" box="[582,744,1980,2002]" italics="true" pageId="11" pageNumber="198">L. xanthostigma</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF266656FFCBFF8061A2FC896C2094CD" httpUri="https://zenodo.org/record/243284/files/figure.png" pageId="12" pageNumber="199" targetBox="[150,1436,193,770]" targetPageId="12">
<paragraph id="8BE636DEFFCBFF8061A2FC896C2094CD" blockId="12.[151,1436,792,906]" pageId="12" pageNumber="199">
<emphasis id="B92DEACCFFCBFF8061A2FC8968369469" bold="true" box="[151,275,792,814]" pageId="12" pageNumber="199">FIGURE 4.</emphasis>
Feet and hands of Panamanian
<emphasis id="B92DEACCFFCBFF80636DFC896D4B9469" box="[600,1134,792,814]" italics="true" pageId="12" pageNumber="199">
<taxonomicName id="4C594D5DFFCBFF80636DFC896A279469" box="[600,770,792,814]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="199" phylum="Chordata" rank="genus">Lepidoblepharis</taxonomicName>
.
<taxonomicName id="4C594D5DFFCBFF80623EFC896D4C9469" box="[779,1129,792,814]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="199" phylum="Chordata" rank="species" species="emberawoundule">Lepidoblepharis emberawoundule</taxonomicName>
:
</emphasis>
right foot of SMF 81954 and right hand of MHCH 2957;
<taxonomicName id="4C594D5DFFCBFF80608DFCA96B64940B" box="[440,577,822,845]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="199" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCBFF80608DFCA96B64940B" box="[440,577,822,845]" italics="true" pageId="12" pageNumber="199">L. rufigularis</emphasis>
</taxonomicName>
(SMF 50659): right foot and right hand;
<emphasis id="B92DEACCFFCBFF8062DEFCA96D80940A" box="[1003,1189,824,845]" italics="true" pageId="12" pageNumber="199">
<taxonomicName id="4C594D5DFFCBFF8062DEFCA96DBA940A" box="[1003,1183,824,845]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="199" phylum="Chordata" rank="species" species="sanctaemartae">L. sanctaemartae</taxonomicName>
:
</emphasis>
left foot of SMF 97419 and right hand of MHCH 2948;
<taxonomicName id="4C594D5DFFCBFF8060D2FCC76BBB942C" box="[487,670,854,875]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="199" phylum="Chordata" rank="species" species="victormartinezi">
<emphasis id="B92DEACCFFCBFF8060D2FCC76BBB942C" box="[487,670,854,875]" italics="true" pageId="12" pageNumber="199">L. victormartinezi</emphasis>
</taxonomicName>
: left foot and left hand of SMF 89963;
<emphasis id="B92DEACCFFCBFF806509FCC76DCF942C" box="[1084,1258,853,875]" italics="true" pageId="12" pageNumber="199">
<taxonomicName id="4C594D5DFFCBFF806509FCC76DC1942C" box="[1084,1252,853,875]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="199" phylum="Chordata" rank="species" species="xanthostigma">L. xanthostigma</taxonomicName>
:
</emphasis>
left foot of SMF 90190 and left hand of SMF 89576. Scale bars equal 1 mm, tip of fourth finger and toe marked with an asterisk.
</paragraph>
</caption>
<paragraph id="8BE636DEFFCBFF8061F2F8806D7D9099" blockId="12.[151,1437,945,2014]" pageId="12" pageNumber="199">
<emphasis id="B92DEACCFFCBFF8061F2F88068C3906D" bold="true" box="[199,486,1809,1834]" pageId="12" pageNumber="199">Hemipenis morphology.</emphasis>
The partially everted hemipenis of SMF 50659 (
<figureCitation id="13622A5BFFCBFF80652EF8806D45906D" box="[1051,1120,1809,1834]" captionStart="FIGURE 5" captionStartId="13.[151,250,1562,1584]" captionTargetBox="[151,1436,179,1539]" captionTargetId="figure@13.[151,1436,179,1541]" captionTargetPageId="13" captionText="FIGURE 5. Escutcheon scales in males of Panamanian Lepidoblepharis. (A) L. emberawoundule: ventral escutcheon with 46 scales, 7 scales long x 11 wide, and subfemoral escutcheon with 4 scales per side, of SMF 81951 (SVL = 25 mm); (B) L. rufigularis: ventral escutcheon with 62 scales, 7 scales long x 13 wide, and subfemoral escutcheon with 3 and 4 scales per side, of SMF 50659 (SVL = 25 mm); (C) L. sanctaemartae: ventral escutcheon of SMF 97419 (SVL = 22 mm) with 92 scales, 10 scales long x 12 wide; (D) L. victormartinezi: ventral escutcheon of SMF 89963 (SVL = 27 mm) comprising 57 intact scales (an original total of 61 is assumed considering the symmetrical scale arrangement of the non-damaged portions), 7 scales long x 14 wide; (E F) L. xanthostigma: (E) ventral escutcheon with 75 scales, 12 scales long x 11 wide, and subfemoral escutcheon with 2 scales per side (no additional scales hidden under strings), of SMF 90189 (SVL = 35 mm), and (F) ventral escutcheon with 36 scales, 6 scales long x 8 wide, and no subfemoral escutcheon of AB 1250 (SVL = 24 mm). All scale bars equal 1 mm." httpUri="https://zenodo.org/record/243285/files/figure.png" pageId="12" pageNumber="199">Fig. 5</figureCitation>
B) is a small organ; sulcus spermaticus bordered by well-developed and smooth sulcal lips; asulcate area of the truncus covered by papillate calyces; proximal portion of apex covered by small spinulate calyces. Due to its incomplete eversion, it is not possible to determinate whether the hemipenis is as bilobate as those of other species of the genus described and/or pictured herein (
<figureCitation id="13622A5BFFCBFF80607BF83068B790FD" box="[334,402,1953,1978]" captionStart="FIGURE 6" captionStartId="14.[151,250,944,966]" captionTargetBox="[216,1371,176,923]" captionTargetId="figure@14.[216,1371,169,923]" captionTargetPageId="14" captionText="FIGURE 6. Hemipenis morphology of four Lepidoblepharis species from Panama. Sulcate-asulcate view, (A B) L. emberawoundule SMF 50968; (C D) L. sanctaemartae SMF 97419 (E F) L. victormartinezi SMF 50950; (G H) L. xanthostigma SMF 91558. All scale bars equal 1 mm." httpUri="https://zenodo.org/record/243286/files/figure.png" pageId="12" pageNumber="199">Fig. 6</figureCitation>
). Nevertheless, it is apparent that the hemipenis of
<taxonomicName id="4C594D5DFFCBFF8062FEF8306C2490FE" box="[971,1281,1953,1977]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="12" pageNumber="199" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFCBFF8062FEF8306C2490FE" box="[971,1281,1953,1977]" italics="true" pageId="12" pageNumber="199">Lepidoblepharis rufigularis</emphasis>
</taxonomicName>
does not bear a conspicuous basal third lobule, in contrast to the other two species described herein.
</paragraph>
<caption id="DF266656FFCAFF8161A2F98B6CB09062" httpUri="https://zenodo.org/record/243285/files/figure.png" pageId="13" pageNumber="200" targetBox="[151,1436,179,1539]" targetPageId="13">
<paragraph id="8BE636DEFFCAFF8161A2F98B6CB09062" blockId="13.[151,1437,1562,1829]" pageId="13" pageNumber="200">
<emphasis id="B92DEACCFFCAFF8161A2F98B68369177" bold="true" box="[151,275,1562,1584]" pageId="13" pageNumber="200">FIGURE 5.</emphasis>
Escutcheon scales in males of Panamanian
<emphasis id="B92DEACCFFCAFF8163E1F98B6AA49177" box="[724,897,1562,1584]" italics="true" pageId="13" pageNumber="200">
<taxonomicName id="4C594D5DFFCAFF8163E1F98B6A589177" box="[724,893,1562,1584]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="200" phylum="Chordata" rank="genus">Lepidoblepharis</taxonomicName>
.
</emphasis>
(A)
<emphasis id="B92DEACCFFCAFF816284F98A6DA69177" box="[945,1155,1562,1584]" italics="true" pageId="13" pageNumber="200">
<taxonomicName id="4C594D5DFFCAFF816284F98A6D589177" box="[945,1149,1562,1584]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="200" phylum="Chordata" rank="species" species="emberawoundule">L. emberawoundule</taxonomicName>
:
</emphasis>
ventral escutcheon with 46 scales, 7 scales long x 11 wide, and subfemoral escutcheon with 4 scales per side, of SMF 81951 (SVL = 25 mm); (B)
<emphasis id="B92DEACCFFCAFF8164B2F9AB6829912A" italics="true" pageId="13" pageNumber="200">
<taxonomicName id="4C594D5DFFCAFF8164B2F9AB6823912A" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="200" phylum="Chordata" rank="species" species="rufigularis">L. rufigularis</taxonomicName>
:
</emphasis>
ventral escutcheon with 62 scales, 7 scales long x 13 wide, and subfemoral escutcheon with 3 and 4 scales per side, of SMF 50659 (SVL = 25 mm); (C)
<emphasis id="B92DEACCFFCAFF816327F9E66BE991CB" box="[530,716,1655,1676]" italics="true" pageId="13" pageNumber="200">
<taxonomicName id="4C594D5DFFCAFF816327F9E66BE391CB" box="[530,710,1655,1676]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="200" phylum="Chordata" rank="species" species="sanctaemartae">L. sanctaemartae</taxonomicName>
:
</emphasis>
ventral escutcheon of SMF 97419 (SVL = 22 mm) with 92 scales, 10 scales long x 12 wide; (D)
<emphasis id="B92DEACCFFCAFF81609BF9046B4B91ED" box="[430,622,1685,1706]" italics="true" pageId="13" pageNumber="200">
<taxonomicName id="4C594D5DFFCAFF81609BF9046B4D91ED" box="[430,616,1685,1706]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="200" phylum="Chordata" rank="species" species="victormartinezi">L. victormartinezi</taxonomicName>
:
</emphasis>
ventral escutcheon of SMF 89963 (SVL = 27 mm) comprising 57 intact scales (an original total of 61 is assumed considering the symmetrical scale arrangement of the non-damaged portions), 7 scales long x 14 wide; (EF)
<emphasis id="B92DEACCFFCAFF81607FF94268D391AF" box="[330,502,1746,1768]" italics="true" pageId="13" pageNumber="200">
<taxonomicName id="4C594D5DFFCAFF81607FF94268CA91AF" box="[330,495,1746,1768]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="13" pageNumber="200" phylum="Chordata" rank="species" species="xanthostigma">L. xanthostigma</taxonomicName>
:
</emphasis>
(E) ventral escutcheon with 75 scales, 12 scales long x 11 wide, and subfemoral escutcheon with 2 scales per side (no additional scales hidden under strings), of SMF 90189 (SVL = 35 mm), and (F) ventral escutcheon with 36 scales, 6 scales long x 8 wide, and no subfemoral escutcheon of AB 1250 (SVL = 24 mm). All scale bars equal 1 mm.
</paragraph>
</caption>
<paragraph id="8BE636DEFFCAFF8161F2F8DD69D690B2" blockId="13.[151,1436,1868,2037]" pageId="13" pageNumber="200">
<emphasis id="B92DEACCFFCAFF8161F2F8DD6B269022" bold="true" box="[199,515,1868,1893]" pageId="13" pageNumber="200">Coloration in preservative</emphasis>
(alcohol 70%). Dorsal ground color Hair Brown (277); posterior margin of orbit with a pale Beige (254) line; dorsum of head suffused with Beige (254); infra- and supralabials with alternating Sepia (279) and white bars; chin and throat with Sepia (286) reticulations on a dirty white background; venter pigmented with Vandyke Brown (281); escutcheon scales unpigmented in the center, with Olive Brown (278) borders.
</paragraph>
<caption id="DF266656FFC9FF8261A2FC216B9C9343" httpUri="https://zenodo.org/record/243286/files/figure.png" pageId="14" pageNumber="201" targetBox="[216,1371,176,923]" targetPageId="14">
<paragraph id="8BE636DEFFC9FF8261A2FC216B9C9343" blockId="14.[151,1437,944,1028]" pageId="14" pageNumber="201">
<emphasis id="B92DEACCFFC9FF8261A2FC21683E9481" bold="true" box="[151,283,944,966]" pageId="14" pageNumber="201">FIGURE 6.</emphasis>
Hemipenis morphology of four
<taxonomicName id="4C594D5DFFC9FF8263BBFC216A119481" box="[654,820,944,966]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="genus">
<emphasis id="B92DEACCFFC9FF8263BBFC216A119481" box="[654,820,944,966]" italics="true" pageId="14" pageNumber="201">Lepidoblepharis</emphasis>
</taxonomicName>
species from Panama. Sulcate-asulcate view, (AB)
<taxonomicName id="4C594D5DFFC9FF8264B2FC23686094A2" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="emberawoundule">
<emphasis id="B92DEACCFFC9FF8264B2FC23686094A2" italics="true" pageId="14" pageNumber="201">L. emberawoundule</emphasis>
</taxonomicName>
SMF 50968; (CD)
<taxonomicName id="4C594D5DFFC9FF82630DFC416BD594A2" box="[568,752,976,997]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="sanctaemartae">
<emphasis id="B92DEACCFFC9FF82630DFC416BD594A2" box="[568,752,976,997]" italics="true" pageId="14" pageNumber="201">L. sanctaemartae</emphasis>
</taxonomicName>
SMF 97419 (EF)
<taxonomicName id="4C594D5DFFC9FF8262E3FC416DB194A2" box="[982,1172,976,997]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="victormartinezi">
<emphasis id="B92DEACCFFC9FF8262E3FC416DB194A2" box="[982,1172,976,997]" italics="true" pageId="14" pageNumber="201">L. victormartinezi</emphasis>
</taxonomicName>
SMF 50950; (GH)
<taxonomicName id="4C594D5DFFC9FF8264B2FC4168059343" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="xanthostigma">
<emphasis id="B92DEACCFFC9FF8264B2FC4168059343" italics="true" pageId="14" pageNumber="201">L. xanthostigma</emphasis>
</taxonomicName>
SMF 91558. All scale bars equal 1 mm.
</paragraph>
</caption>
<paragraph id="8BE636DEFFC9FF8261F2FBBA6C0293BF" blockId="14.[151,1437,1067,1596]" pageId="14" pageNumber="201">
<emphasis id="B92DEACCFFC9FF8261F2FBBA68B39303" bold="true" box="[199,406,1067,1092]" pageId="14" pageNumber="201">Coloration in life</emphasis>
(
<figureCitation id="13622A5BFFC9FF82609DFBBA68DE9303" box="[424,507,1067,1092]" captionStart="FIGURE 11" captionStartId="23.[151,250,852,874]" captionTargetBox="[188,1398,181,830]" captionTargetId="figure@23.[188,1398,181,831]" captionTargetPageId="23" captionText="FIGURE 11. Holotype of Lepidoblepharis rufigularis (male SMF 50659) in life. (A B) lateral view; (C) gular and chest region." httpUri="https://zenodo.org/record/243291/files/figure.png" pageId="14" pageNumber="201">Fig. 11</figureCitation>
). Dorsal ground color Grayish Horn Color (268), with patches of Lavender Blue (195) and Vandyke Brown (281) scales, same pattern on neck and head; a straight Light Sky Blue (191) postorbital line, running up to level of ear; infra- and supralabials with alternating Sepia (279) and Burnt Orange (10) bars; chin and throat with Sepia (279) reticulations on a Burnt Orange (10) background; ventral parts suffused with Dusky Brown (285) and Light Sky Blue (191); escutcheon scales Smoke Gray (267), with Dusky Brown (285) scale tips; tail with a dorsolateral Salmon Color (59) line; toes and fingers suffused with Brick Red (36).
</paragraph>
</subSubSection>
<subSubSection id="C3436555FFC9FF8261F2FA9268A592B3" pageId="14" pageNumber="201" type="distribution">
<paragraph id="8BE636DEFFC9FF8261F2FA9268A592B3" blockId="14.[151,1437,1067,1596]" pageId="14" pageNumber="201">
<emphasis id="B92DEACCFFC9FF8261F2FA9268D3925B" bold="true" box="[199,502,1283,1308]" pageId="14" pageNumber="201">Distribution and habitat.</emphasis>
<taxonomicName id="4C594D5DFFC9FF8260CAFA926A1D925C" box="[511,824,1283,1307]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFC9FF8260CAFA926A1D925C" box="[511,824,1283,1307]" italics="true" pageId="14" pageNumber="201">Lepidoblepharis rufigularis</emphasis>
</taxonomicName>
is known only from the
<typeStatus id="54E2887CFFC9FF82656FFA956DAF925B" box="[1114,1162,1284,1308]" pageId="14" pageNumber="201">type</typeStatus>
locality in southeastern Darién province. Given the close proximity of the
<typeStatus id="54E2887CFFC9FF8263D4FAB96A349207" box="[737,785,1320,1344]" pageId="14" pageNumber="201">type</typeStatus>
locality to
<collectingCountry id="F34E764EFFC9FF8262ACFAB66D2F9207" box="[921,1034,1319,1344]" name="Colombia" pageId="14" pageNumber="201">Colombia</collectingCountry>
, it likely occurs there as well. The habitat at the
<typeStatus id="54E2887CFFC9FF826007FADD68479223" box="[306,354,1356,1380]" pageId="14" pageNumber="201">type</typeStatus>
locality of
<taxonomicName id="4C594D5DFFC9FF8260D3FADD6B589224" box="[486,637,1355,1379]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFC9FF8260D3FADD6B589224" box="[486,637,1355,1379]" italics="true" pageId="14" pageNumber="201">L. rufigularis</emphasis>
</taxonomicName>
is part of the eastern Panamanian montane forests (
<bibRefCitation id="EFC84B2FFFC9FF8265FCFADA6C629223" author="Fund" box="[1225,1351,1355,1380]" pageId="14" pageNumber="201" refString="Fund, W. (2011) Isthmian-Atlantic moist forests. Available from http: // www. eoearth. org / view / article / 153927 (accessed 24 October 2013)" type="journal article" year="2011">Fund 2011</bibRefCitation>
), in the Darién mountain range, around
<quantity id="4CA19B3BFFC9FF8260CEFAE16B6A92C0" box="[507,591,1392,1416]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="14" pageNumber="201" unit="m" value="1000.0">1000 m</quantity>
elev. Most probably,
<taxonomicName id="4C594D5DFFC9FF826275FAE16AF092C0" box="[832,981,1391,1415]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFC9FF826275FAE16AF092C0" box="[832,981,1391,1415]" italics="true" pageId="14" pageNumber="201">L. rufigularis</emphasis>
</taxonomicName>
lives in the leaf-litter and feeds on small invertebrates like other
<taxonomicName id="4C594D5DFFC9FF826087FA026B4C92EC" box="[434,617,1427,1451]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="genus">
<emphasis id="B92DEACCFFC9FF826087FA026B4C92EC" box="[434,617,1427,1451]" italics="true" pageId="14" pageNumber="201">Lepidoblepharis</emphasis>
</taxonomicName>
spp. (
<bibRefCitation id="EFC84B2FFFC9FF82638CFA026A5C92EB" author="Vitt" box="[697,889,1427,1452]" pageId="14" pageNumber="201" refString="Vitt, L. J., Sartorius, S. S., Avila-Pires, T. C., Zani, P. A. &amp; Esposito M. C. (2005) Small in a big world: Ecology of leaf-litter geckos in new world tropical forests. Herpetological Monographs, 19, 137 - 152. http: // dx. doi. org / 10.1655 / 0733 - 1347 (2005) 019 [0137: SIABWE] 2.0. CO; 2" type="journal article" year="2005">
Vitt
<emphasis id="B92DEACCFFC9FF8263C4FA056A0D92EC" box="[753,808,1427,1451]" italics="true" pageId="14" pageNumber="201">et al</emphasis>
. 2005
</bibRefCitation>
). The specimen probably resided between the leaf-litter on a large fallen log, and was uncovered and collected after one of the local supporters in the group stepped over the log.
</paragraph>
</subSubSection>
<subSubSection id="C3436555FFC9FF8261F2FA6E6A7F917B" pageId="14" pageNumber="201" type="etymology">
<paragraph id="8BE636DEFFC9FF8261F2FA6E6A7F917B" blockId="14.[151,1437,1067,1596]" pageId="14" pageNumber="201">
<emphasis id="B92DEACCFFC9FF8261F2FA6E686B915F" bold="true" box="[199,334,1535,1560]" pageId="14" pageNumber="201">Etymology.</emphasis>
The name
<taxonomicName id="4C594D5DFFC9FF8260F9FA6E6B619150" box="[460,580,1535,1559]" class="Reptilia" family="Sphaerodactylidae" genus="Lepidoblepharis" kingdom="Animalia" order="Squamata" pageId="14" pageNumber="201" phylum="Chordata" rank="species" species="rufigularis">
<emphasis id="B92DEACCFFC9FF8260F9FA6E6B619150" box="[460,580,1535,1559]" italics="true" pageId="14" pageNumber="201">rufigularis</emphasis>
</taxonomicName>
is a compound word that comes from the Latin
<emphasis id="B92DEACCFFC9FF826555F9916DBC915F" box="[1120,1177,1536,1560]" italics="true" pageId="14" pageNumber="201">rufus</emphasis>
(red) and
<emphasis id="B92DEACCFFC9FF82643AFA6E6C649150" box="[1295,1345,1535,1559]" italics="true" pageId="14" pageNumber="201">gula</emphasis>
(throat) referring to the bright orange throat color in this species in life.
</paragraph>
</subSubSection>
</treatment>
</document>