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<mods:title id="DFB966A9A121A071E684801BDC62305B">Filling gaps in global myrmecology: ants of the Kingdom of Bahrain (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="35E06E96409881689F25B3FA1F32E3B1">Sharaf, Mostafa R.</mods:namePart>
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<mods:namePart id="379F21789A424868C37F994D1DE246E4">Mohamed, AbdulAziz M. A.</mods:namePart>
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<mods:namePart id="82A7191DAAA963C93CF9A7CCDC908298">Georgiadis, Christos</mods:namePart>
<mods:affiliation id="6DA67BE9581AD8952705FCDF1285790A">Section of Zoology-Marine Biology and Zoological Museum, Department of Biology, National and Kapodistrian University of Athens, Athens, Greece;</mods:affiliation>
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<mods:namePart id="CE4211F2A2B62834344984C379A0255B">Nasser, Mohamed G.</mods:namePart>
<mods:affiliation id="4E537EF0902B7E640062CFFDDFFE7E4A">Research Laboratory of Biogeography and Wildlife Parasitology (RLBWP), Department of Entomology, Faculty of Science, Ain Shams University, Abbassia, Egypt</mods:affiliation>
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<mods:date id="51BB643237839B631B9373F314F7E879">2024</mods:date>
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<paragraph id="8BCF4C2DFFB2FF85FE5EFF25AE5BFF32" blockId="41.[441,874,142,169]" box="[441,874,142,169]" pageId="41" pageNumber="1746">
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<taxonomicName id="4C7037AEFFB2FF85FE5EFF25AE5BFF32" ID-CoL="44829" authority="Andre, 1881" authorityName="Andre" authorityYear="1881" box="[441,874,142,169]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="1746" phylum="Arthropoda" rank="species" species="abeillei">
<emphasis id="B904903FFFB2FF85FE5EFF25AFFDFF32" bold="true" box="[441,716,142,169]" italics="true" pageId="41" pageNumber="1746">Monomorium abeillei</emphasis>
André, 1881
</taxonomicName>
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(
<figureCitation id="134B50A8FFB2FF85FDB0FF10AFFDFF4D" box="[599,716,187,214]" captionStart="Figure 26" captionStartId="42.[160,225,1753,1776]" captionTargetBox="[370,945,168,1719]" captionTargetId="figure-3@42.[367,948,142,1722]" captionTargetPageId="42" captionText="Figure 26. Monomorium abeillei. Lectotype worker. (A) Body in profile. (B) Head in full-face view, CASENT0915411, AntWeb.org (Will Ericson). (C) Distribution map." figureDoi="http://doi.org/10.5281/zenodo.14241377" httpUri="https://zenodo.org/record/14241377/files/figure.png" pageId="41" pageNumber="1746">Figure 26</figureCitation>
)
</paragraph>
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<paragraph id="8BCF4C2DFFB2FF85FF27FF47AEA7FE9E" blockId="41.[192,918,236,261]" box="[192,918,236,261]" pageId="41" pageNumber="1746">
<treatmentCitationGroup id="AB606B03FFB2FF85FF27FF47AEA7FE9E" box="[192,918,236,261]" pageId="41" pageNumber="1746">
<treatmentCitation id="0AD16A3CFFB2FF85FF27FF47AFA8FE9E" author="Andre E" box="[192,665,236,261]" page="531" pageId="41" pageNumber="1746" year="1881">
<taxonomicName id="4C7037AEFFB2FF85FF27FF47AFA8FE9E" ID-CoL="44829" authority="Andre, 1881 b, p. 531" authorityName="Andre" authorityPageNumber="531" authorityYear="1881" box="[192,665,236,261]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="1746" phylum="Arthropoda" rank="species" species="abeillei">
<emphasis id="B904903FFFB2FF85FF27FF47AC98FE9E" box="[192,425,236,261]" italics="true" pageId="41" pageNumber="1746">Monomorium abeillei</emphasis>
<bibRefCitation id="EFE131DCFFB2FF85FE57FF47AF76FE9E" author="Andre E" box="[432,583,236,261]" pageId="41" pageNumber="1746" pagination="525 - 535" refId="ref18443" refString="Andre E. 1881 b. [Untitled. Monomorium abeillei, n. sp.]. p. 531 In: emery, C. 1881. Viaggio ad Assab nel Mar Rosso dei Signori G. Doria ed O. Beccari con il R. Avviso &quot; Esploratore &quot; dal 16 novembre 1879 al 26 febbraio 1880. I. Formiche Ann Mus Civ St Nat. 16: 525 - 535." type="journal article" year="1881">André, 1881b</bibRefCitation>
, p. 531
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(w.)
<collectingCountry id="F3670CBDFFB2FF85FD36FF47AE3FFE9E" box="[721,782,236,261]" name="Israel" pageId="41" pageNumber="1746">Israel</collectingCountry>
. Palaearctic.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FFB2FF85FF47FE9BAE5BFD81" pageId="41" pageNumber="1746" type="diagnosis">
<paragraph id="8BCF4C2DFFB2FF85FF47FE9BAC25FED1" blockId="41.[160,1156,304,538]" box="[160,276,304,330]" pageId="41" pageNumber="1746">
<emphasis id="B904903FFFB2FF85FF47FE9BAC25FED1" bold="true" box="[160,276,304,330]" italics="true" pageId="41" pageNumber="1746">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BCF4C2DFFB2FF85FF47FEF8AE5BFD81" blockId="41.[160,1156,304,538]" pageId="41" pageNumber="1746">
<emphasis id="B904903FFFB2FF85FF47FEF8ADCCFEF6" bold="true" box="[160,253,339,365]" italics="true" pageId="41" pageNumber="1746">Worker.</emphasis>
Colour uniform brown-black to black; with 910 ommatidia in longest row; mesosoma in profile with promesonotal dorsum meeting flat propodeum at oblique angle; metanotal groove feebly impressed; propodeum with well-defined, longitudinal furrow; petiole in profile with high, rounded triangular dorsum; pronotum and petiole each with one pair of setae, postpetiole with two pairs; head, pronotum, and nodes superficially reticulate; gaster shining with superficial sculpture.
</paragraph>
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<paragraph id="8BCF4C2DFFB2FF85FF47FDEDAC4EFDFB" blockId="41.[160,821,582,642]" box="[160,383,582,608]" pageId="41" pageNumber="1746">
<emphasis id="B904903FFFB2FF85FF47FDEDAC4EFDFB" bold="true" box="[160,383,582,608]" italics="true" pageId="41" pageNumber="1746">Material examined</emphasis>
</paragraph>
<paragraph id="8BCF4C2DFFB2FF85FF47FDC2AE04FD19" blockId="41.[160,821,582,642]" box="[160,821,617,642]" pageId="41" pageNumber="1746">Fourteen sites: 2; 3; 4; 8; 9; 10; 11; 13; 15; 21; 24; 27; 29; 30.</paragraph>
<paragraph id="8BCF4C2DFFB2FF85FF47FD05AF8DFCEE" blockId="41.[160,1156,686,885]" pageId="41" pageNumber="1746">
<emphasis id="B904903FFFB2FF85FF47FD05AC4BFD53" bold="true" box="[160,378,686,712]" italics="true" pageId="41" pageNumber="1746">Geographic range.</emphasis>
A palaearctic species originally described from
<collectingCountry id="F3670CBDFFB2FF85FC76FD04AEFAFD53" box="[913,971,687,712]" name="Israel" pageId="41" pageNumber="1746">Israel</collectingCountry>
and widespread in countries of the Middle East including
<collectingCountry id="F3670CBDFFB2FF85FD99FD7AAFF0FD71" box="[638,705,721,746]" name="Egypt" pageId="41" pageNumber="1746">Egypt</collectingCountry>
,
<collectingCountry id="F3670CBDFFB2FF85FD29FD7AAE3AFD71" box="[718,779,721,746]" name="Israel" pageId="41" pageNumber="1746">Israel</collectingCountry>
,
<collectingCountry id="F3670CBDFFB2FF85FCF0FD7AAE52FD71" box="[791,867,721,746]" name="Jordan" pageId="41" pageNumber="1746">Jordan</collectingCountry>
and
<collectingCountry id="F3670CBDFFB2FF85FC46FD7AAEE6FD71" box="[929,983,721,746]" name="Syria" pageId="41" pageNumber="1746">Syria</collectingCountry>
(
<bibRefCitation id="EFE131DCFFB2FF85FC01FD7AAC13FC96" author="Borowiec L &amp; Salata S" pageId="41" pageNumber="1746" pagination="1 - 26" refId="ref19104" refString="Borowiec L, Salata S. 2020. Review of ants (Hymenoptera: Formicidae) from Jordan. Ann Up Sil Mus Bytom Entom. 29: 1 - 26. doi: 10.5281 / zenodo. 3733156." type="journal article" year="2020">Borowiec and Salata 2020</bibRefCitation>
), and all countries of the Arabian Peninsula: the
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,
<collectingCountry id="F3670CBDFFB2FF85FC6FFD5FAEE6FC96" box="[904,983,756,781]" name="Kuwait" pageId="41" pageNumber="1746">Kuwait</collectingCountry>
,
<collectingCountry id="F3670CBDFFB2FF85FC04FD5FA91BFC96" box="[995,1066,756,781]" name="Oman" pageId="41" pageNumber="1746">Oman</collectingCountry>
,
<collectingCountry id="F3670CBDFFB2FF85FBD2FD5FA9B2FC96" box="[1077,1155,756,781]" name="Yemen" pageId="41" pageNumber="1746">Yemen</collectingCountry>
(
<bibRefCitation id="EFE131DCFFB2FF85FF41FCBCAC44FCAB" author="Collingwood CA" box="[166,373,791,816]" pageId="41" pageNumber="1746" pagination="230 - 302" refId="ref19269" refString="Collingwood CA. 1985. Hymenoptera: fam. Formicidae of Saudi Arabia. Fauna Saudi Arabia. 7: 230 - 302. doi: 10.5281 / zenodo. 26692." type="journal article" year="1985">Collingwood 1985</bibRefCitation>
;
<bibRefCitation id="EFE131DCFFB2FF85FE65FCBCAFE1FCAB" author="Collingwood CA &amp; Agosti D" box="[386,720,791,816]" pageId="41" pageNumber="1746" pagination="300 - 385" refId="ref19301" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
;
<bibRefCitation id="EFE131DCFFB2FF85FD3AFCBCAEAEFCAB" author="Sharaf MR &amp; Mohamed AA &amp; Boudinot BE &amp; Wetterer JK &amp; Hita Garcia F &amp; Al Dhafer HM &amp; Aldawood AS" box="[733,927,791,816]" pageId="41" pageNumber="1746" refId="ref24396" refString="Sharaf MR, Mohamed AA, Boudinot BE, Wetterer JK, Hita Garcia F, Al Dhafer HM, Aldawood AS. 2021. Monomorium (Hymenoptera: Formicidae) of the Arabian Peninsula with description of two new species, M. heggyi sp. n. and M. khalidi sp. n. PeerJ. 9: e 10726. doi: 10.7717 / peerj. 10726." type="journal volume" year="2021">
Sharaf
<emphasis id="B904903FFFB2FF85FCCCFCBCAE6AFCAB" box="[811,859,791,816]" italics="true" pageId="41" pageNumber="1746">et al</emphasis>
. 2021
</bibRefCitation>
),
<collectingCountry id="F3670CBDFFB2FF85FC54FCBCAEC0FCAB" box="[947,1009,791,816]" name="Qatar" pageId="41" pageNumber="1746">Qatar</collectingCountry>
(
<bibRefCitation id="EFE131DCFFB2FF85FC19FCBCADD5FCC9" author="Sharaf MR &amp; Abdel-Dayem MS &amp; Mohamed AA &amp; Fisher BL &amp; Aldawood AS" pageId="41" pageNumber="1746" pagination="533 - 560" refId="ref23380" refString="Sharaf MR, Abdel-Dayem MS, Mohamed AA, Fisher BL, Aldawood AS. 2020 a. A preliminary synopsis of the ant fauna (Hymenoptera: Formicidae) of Qatar with remarks on the zoogeography. Ann Zool. 70 (4): 533 - 560. doi: 10.3161 / 00034541 ANZ 2020.70.4.005." type="journal article" year="2020">
Sharaf
<emphasis id="B904903FFFB2FF85FBA9FCBCA94FFCAB" box="[1102,1150,791,816]" italics="true" pageId="41" pageNumber="1746">et al</emphasis>
. 2020a
</bibRefCitation>
), and the
<collectingCountry id="F3670CBDFFB2FF85FEB6FC92ACB1FCC9" box="[337,384,825,850]" name="United Arab Emirates" pageId="41" pageNumber="1746">UAE</collectingCountry>
(
<bibRefCitation id="EFE131DCFFB2FF85FE6CFC92AFF7FCC9" author="Tigar BJ &amp; Collingwood CA" box="[395,710,825,850]" pageId="41" pageNumber="1746" pagination="13 - 14" refId="ref25007" refString="Tigar BJ, Collingwood CA. 1993. A preliminary list of ant records from Abu Dhabi Emirate, U. A. E. Tribulus. 3 (2): 13 - 14." type="journal article" year="1993">Tigar and Collingwood 1993</bibRefCitation>
;
<bibRefCitation id="EFE131DCFFB2FF85FD37FC92AEE4FCC9" author="Collingwood CA &amp; Agosti D &amp; Sharaf MR &amp; van Harten A" box="[720,981,825,850]" pageId="41" pageNumber="1746" pagination="405 - 474" refId="ref19333" refString="Collingwood CA, Agosti D, Sharaf MR, van Harten A. 2011. Order Hymenoptera, family Formicidae. Arthropod Fauna UAE. 4: 405 - 474. doi: 10.5281 / zenodo. 1168586." type="journal article" year="2011">
Collingwood
<emphasis id="B904903FFFB2FF85FC82FC92AEA3FCC9" box="[869,914,825,850]" italics="true" pageId="41" pageNumber="1746">et al</emphasis>
. 2011
</bibRefCitation>
). It is one of the most common
<taxonomicName id="4C7037AEFFB2FF85FEABFCF7AF08FCEE" box="[332,569,860,885]" pageId="41" pageNumber="1746">
<emphasis id="B904903FFFB2FF85FEABFCF7ACD0FCEE" box="[332,481,860,885]" italics="true" pageId="41" pageNumber="1746">Monomorium</emphasis>
species
</taxonomicName>
in the
<collectingCountry id="F3670CBDFFB2FF85FD6EFCF7AF89FCEE" box="[649,696,860,885]" name="Saudi Arabia" pageId="41" pageNumber="1746">KSA</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FFB2FF85FF47FC0BAEABFBDE" pageId="41" pageNumber="1746" type="biology_ecology">
<paragraph id="8BCF4C2DFFB2FF85FF47FC0BAEABFBDE" blockId="41.[160,1156,928,1093]" pageId="41" pageNumber="1746">
<emphasis id="B904903FFFB2FF85FF47FC0BAC98FC21" bold="true" box="[160,425,928,954]" italics="true" pageId="41" pageNumber="1746">Ecology and biology.</emphasis>
The habitat preference of
<taxonomicName id="4C7037AEFFB2FF85FD1CFC0AAE43FC21" authorityName="Andre" authorityYear="1881" box="[763,882,929,954]" class="Insecta" family="Formicidae" genus="Monomorium" kingdom="Animalia" order="Hymenoptera" pageId="41" pageNumber="1746" phylum="Arthropoda" rank="species" species="abeillei">
<emphasis id="B904903FFFB2FF85FD1CFC0AAE43FC21" box="[763,882,929,954]" italics="true" pageId="41" pageNumber="1746">M. abeillei</emphasis>
</taxonomicName>
includes sandy desert (
<bibRefCitation id="EFE131DCFFB2FF85FF41FC6FACCBFC46" author="Collingwood CA &amp; Agosti D" box="[166,506,964,989]" pageId="41" pageNumber="1746" pagination="300 - 385" refId="ref19301" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
), leaf litter (Sharaf, unpublished data), coastal areas, and high ground in the
<collectingRegion id="49B482CFFFB2FF85FE9BFC4CAC97FB9B" box="[380,422,999,1024]" country="Saudi Arabia" name="'Asir" pageId="41" pageNumber="1746">Asir</collectingRegion>
mountains above
<quantity id="4C88E1C8FFB2FF85FD9FFC4CAFFFFB9B" box="[632,718,999,1024]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="41" pageNumber="1746" unit="m" value="2200.0">2200 m</quantity>
.a.s.l. (
<bibRefCitation id="EFE131DCFFB2FF85FCEAFC4CAEEAFB9B" author="Collingwood CA" box="[781,987,999,1024]" pageId="41" pageNumber="1746" pagination="230 - 302" refId="ref19269" refString="Collingwood CA. 1985. Hymenoptera: fam. Formicidae of Saudi Arabia. Fauna Saudi Arabia. 7: 230 - 302. doi: 10.5281 / zenodo. 26692." type="journal article" year="1985">Collingwood 1985</bibRefCitation>
). In the
<collectingRegion id="49B482CFFFB2FF85FBD0FC4CAC33FBB9" country="Saudi Arabia" name="Ar Riyad" pageId="41" pageNumber="1746">Riyadh Province</collectingRegion>
(
<collectingCountry id="F3670CBDFFB2FF85FEF4FBA2AC77FBB9" box="[275,326,1033,1058]" name="Saudi Arabia" pageId="41" pageNumber="1746">KSA</collectingCountry>
), it nests in sandy habitats and forages next to
<taxonomicName id="4C7037AEFFB2FF85FC6FFBA2A9B5FBB9" authority="Decnet" authorityName="Decnet" box="[904,1156,1033,1058]" class="Magnoliopsida" family="Apocynaceae" genus="Rhazya" kingdom="Plantae" order="Gentianales" pageId="41" pageNumber="1746" phylum="Tracheophyta" rank="species" species="stricta">
<emphasis id="B904903FFFB2FF85FC6FFBA2A916FBB9" box="[904,1063,1033,1058]" italics="true" pageId="41" pageNumber="1746">Rhazya stricta</emphasis>
Decnet
</taxonomicName>
(
<taxonomicName id="4C7037AEFFB2FF85FF41FB87AC71FBDE" box="[166,320,1068,1093]" class="Magnoliopsida" family="Apocynaceae" kingdom="Plantae" order="Gentianales" pageId="41" pageNumber="1746" phylum="Tracheophyta" rank="family">Apocynaceae</taxonomicName>
) in the Rawdhat Khorim National Park (
<bibRefCitation id="EFE131DCFFB2FF85FD1CFB87AEBCFBDE" author="Salman S" box="[763,909,1068,1093]" pageId="41" pageNumber="1746" refId="ref22919" refString="Salman S 2018. Ant fauna (Hymenoptera: Formicidae) of Riyadh Province, Kingdom of Saudi Arabia [M. Sc. thesis (unpublished)]. King Saud University, Riyadh (Kingdom of Saudi Arabia). 297 pp." type="book" year="2018">Salman 2018</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>