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<document id="0E47853D1E60922E8EEBFBAFF186729F" ID-CLB-Dataset="305810" ID-DOI="10.1080/00222933.2024.2388791" ID-GBIF-Dataset="7328f914-b6fd-4df6-be55-0e3ae7a633fa" ID-ISSN="1464-5262" ID-Zenodo-Dep="14241324" ID-ZooBank="18D05DD2- 4B64-4A87-8389-582D5714411C" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1732836998606" checkinUser="plazi" docAuthor="Sharaf, Mostafa R., Wetterer, James K., Mohamed, AbdulAziz M. A., Georgiadis, Christos, Nasser, Mohamed G. &amp; Aldawood, Abdulrahman S." docDate="2024" docId="03D9FD3BFF94FFBEFE48FC64AC2AF92C" docLanguage="en" docName="JNATHIST.58.41-44.1705-1786.pdf" docOrigin="Journal of Natural History 58 (41 - 44)" docSource="http://dx.doi.org/10.1080/00222933.2024.2388791" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Lepisiota bahrainensis Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood 2024, sp. n." docType="treatment" docVersion="7" lastPageNumber="1723" masterDocId="FFE08543FF9BFFACFFE7FFABAD31FF9B" masterDocTitle="Filling gaps in global myrmecology: ants of the Kingdom of Bahrain (Hymenoptera: Formicidae)" masterLastPageNumber="1786" masterPageNumber="1705" pageNumber="1720" updateTime="1733171467126" updateUser="felipe" zenodo-license-document="CLOSED">
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<mods:title id="70FE1A477E7ADE0017CDED97FB64DCE4">Filling gaps in global myrmecology: ants of the Kingdom of Bahrain (Hymenoptera: Formicidae)</mods:title>
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<mods:namePart id="9A487440B3036A7C3529F46C42D829D4">Sharaf, Mostafa R.</mods:namePart>
<mods:affiliation id="257EB455BFF8A911EE31188EDA303220">Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Kingdom of Saudi Arabia; &amp; Division of Invertebrate Zoology, American Museum of Natural History, New York, NY, USA;</mods:affiliation>
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<mods:namePart id="EF8515B29527C7B184CCC389CE1F9864">Wetterer, James K.</mods:namePart>
<mods:affiliation id="241B0F513CD69AABD3B8EB05E98DF012">Wilkes Honors College, Florida Atlantic University, Jupiter, FL, USA;</mods:affiliation>
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<mods:namePart id="FD91290E556140E0CDC8582C2372350F">Mohamed, AbdulAziz M. A.</mods:namePart>
<mods:affiliation id="DA1CC721DCE7330F1A0EA0A576428DCA">Agriculture Affairs, Ministry of Municipalities Affairs and Agriculture, Kingdom of Bahrain;</mods:affiliation>
</mods:name>
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<mods:namePart id="FDB83F133F7E07F7B0D6748053EA7FA3">Georgiadis, Christos</mods:namePart>
<mods:affiliation id="031849F868F6868BBED16A192D85FD45">Section of Zoology-Marine Biology and Zoological Museum, Department of Biology, National and Kapodistrian University of Athens, Athens, Greece;</mods:affiliation>
</mods:name>
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<mods:namePart id="1D12BCEB2B47BE36870188EC9E155855">Nasser, Mohamed G.</mods:namePart>
<mods:affiliation id="81F12B5023931DDD1D6D2B6657C0E026">Research Laboratory of Biogeography and Wildlife Parasitology (RLBWP), Department of Entomology, Faculty of Science, Ain Shams University, Abbassia, Egypt</mods:affiliation>
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<mods:namePart id="58587B515F3842DF8B1E8E057F0DC141">Aldawood, Abdulrahman S.</mods:namePart>
<mods:affiliation id="4AC43B88C1F2B2880DAB0FBBC9384E95">Department of Plant Protection, College of Food and Agriculture Sciences, King Saud University, Riyadh, Kingdom of Saudi Arabia;</mods:affiliation>
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<mods:title id="CB85B7237474EBC51958454A051998A3">Journal of Natural History</mods:title>
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<mods:date id="1A4FC98F55AD3EBA583991F02BDC9A98">2024</mods:date>
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<mods:number id="ACAF6942B4DC7356789F79486CDF4A9B">2024-09-16</mods:number>
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<mods:number id="0874DB962FD90C055AD6537D3669A206">58</mods:number>
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<mods:number id="D6DF9FB91074024C8D548B5A49EB5C6B">41 - 44</mods:number>
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<treatment id="03D9FD3BFF94FFBEFE48FC64AC2AF92C" ID-DOI="http://doi.org/10.5281/zenodo.14261660" ID-GBIF-Taxon="248207636" ID-Zenodo-Dep="14261660" LSID="urn:lsid:plazi:treatment:03D9FD3BFF94FFBEFE48FC64AC2AF92C" httpUri="http://treatment.plazi.org/id/03D9FD3BFF94FFBEFE48FC64AC2AF92C" lastPageId="18" lastPageNumber="1723" pageId="15" pageNumber="1720" scope_family="Formicidae" scope_order="Hymenoptera" scope_subFamily="Formicinae">
<subSubSection id="C36A1FA6FF94FFA3FE48FC64AE45FC71" box="[431,884,975,1002]" pageId="15" pageNumber="1720" type="nomenclature">
<paragraph id="8BCF4C2DFF94FFA3FE48FC64AE45FC71" blockId="15.[431,884,975,1002]" box="[431,884,975,1002]" pageId="15" pageNumber="1720">
<heading id="D087FB41FF94FFA3FE48FC64AE45FC71" box="[431,884,975,1002]" centered="true" fontSize="11" level="2" pageId="15" pageNumber="1720" reason="5">
<taxonomicName id="4C7037AEFF94FFA3FE48FC64AE14FC71" authority="Sharaf" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" box="[431,805,975,1002]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="1720" phylum="Arthropoda" rank="species" species="bahrainensis" status="sp. nov.">
<emphasis id="B904903FFF94FFA3FE48FC64AFFEFC71" bold="true" box="[431,719,975,1002]" italics="true" pageId="15" pageNumber="1720">Lepisiota bahrainensis</emphasis>
Sharaf
</taxonomicName>
<emphasis id="B904903FFF94FFA3FCCAFC64AE45FC71" bold="true" box="[813,884,975,1002]" pageId="15" pageNumber="1720">
<taxonomicNameLabel id="A2372D44FF94FFA3FCCAFC64AE45FC71" box="[813,884,975,1002]" pageId="15" pageNumber="1720" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF94FFA3FDA8FC55AFE5FB82" box="[591,724,1022,1049]" pageId="15" pageNumber="1720" type="description">
<paragraph id="8BCF4C2DFF94FFA3FDA8FC55AFE5FB82" blockId="15.[591,724,1022,1049]" box="[591,724,1022,1049]" pageId="15" pageNumber="1720">
(
<figureCitation id="134B50A8FF94FFA3FDB0FC55AFFDFB82" box="[599,716,1022,1049]" captionStart="Figure 10" captionStartId="17.[160,225,1048,1071]" captionTargetBox="[165,1150,144,1014]" captionTargetId="figure-173@17.[163,1153,142,1017]" captionTargetPageId="17" captionText="Figure 10. Lepisiota bahrainensis sp. n. Holotype worker. (A) Body in profile. (B) Head in full-face view. (C) Propodeum and petiole in dorsal view, ANTWEB1041925, AntWeb.org (Michele Esposito). (D) Distribution map." figureDoi="http://doi.org/10.5281/zenodo.14241345" httpUri="https://zenodo.org/record/14241345/files/figure.png" pageId="15" pageNumber="1720">Figure 10</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF94FFA3FF47FBFDACF8FB08" pageId="15" pageNumber="1720" type="materials_examined">
<paragraph id="8BCF4C2DFF94FFA3FF47FBFDACF8FB08" blockId="15.[160,1155,1110,1171]" pageId="15" pageNumber="1720">
<materialsCitation id="3B184670FF94FFA3FF47FBFDAE11FBEB" box="[160,800,1110,1136]" collectionCode="KSMA" pageId="15" pageNumber="1720" specimenCode="ANTWEB1041925" specimenCount="1" specimenCount-worker="1" typeStatus="holotype">
<emphasis id="B904903FFF94FFA3FF47FBFDAC5DFBEB" bold="true" box="[160,364,1110,1136]" italics="true" pageId="15" pageNumber="1720">
<typeStatus id="54CBF28FFF94FFA3FF47FBFDAC3AFBEB" box="[160,267,1110,1136]" pageId="15" pageNumber="1720" type="holotype">Holotype</typeStatus>
<specimenCount id="9D7687A4FF94FFA3FEF4FBFDAC56FBEB" box="[275,359,1110,1136]" pageId="15" pageNumber="1720" type="worker">worker</specimenCount>
.
</emphasis>
One site: 2 (
<specimenCode id="DBD6E456FF94FFA3FDE7FBFCAFF7FBEB" box="[512,710,1111,1136]" collectionCode="ANTWEB" pageId="15" pageNumber="1720">ANTWEB1041925</specimenCode>
,
<collectionCode id="ED61D4E8FF94FFA3FD37FBFCAE27FBEB" box="[720,790,1111,1136]" collectionName="KSMA" pageId="15" pageNumber="1720">KSMA</collectionCode>
).
</materialsCitation>
The new species is known only from the
<typeStatus id="54CBF28FFF94FFA3FEEEFBD1AC5CFB08" box="[265,365,1146,1171]" pageId="15" pageNumber="1720" type="holotype">holotype</typeStatus>
worker.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF94FFA3FF47FB14AC22FA85" pageId="15" pageNumber="1720" type="description">
<paragraph id="8BCF4C2DFF94FFA3FF47FB14AC7CFB42" blockId="15.[160,1155,1215,1310]" box="[160,333,1215,1241]" pageId="15" pageNumber="1720">
<emphasis id="B904903FFF94FFA3FF47FB14AC7CFB42" bold="true" box="[160,333,1215,1241]" italics="true" pageId="15" pageNumber="1720">Measurements</emphasis>
</paragraph>
<paragraph id="8BCF4C2DFF94FFA3FF47FB49AC22FA85" blockId="15.[160,1155,1215,1310]" pageId="15" pageNumber="1720">
<emphasis id="B904903FFF94FFA3FF47FB49AC23FB67" bold="true" box="[160,274,1250,1276]" italics="true" pageId="15" pageNumber="1720">
<typeStatus id="54CBF28FFF94FFA3FF47FB49AC3CFB67" box="[160,269,1250,1276]" pageId="15" pageNumber="1720" type="holotype">Holotype</typeStatus>
.
</emphasis>
EL 0.17, HL 0.60, HW 0.49,
<collectionCode id="ED61D4E8FF94FFA3FDAEFB48AF5FFB67" box="[585,622,1251,1276]" country="Thailand" name="Paleontological Collections" pageId="15" pageNumber="1720">PW</collectionCode>
0.40, SL 0.94, TL 2.56, WL 1.01. Indices: CI 82, EI 35, SI 191.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF94FFA3FF47FAE0A9B5F989" pageId="15" pageNumber="1720" type="diagnosis">
<paragraph id="8BCF4C2DFF94FFA3FF47FAE0AC25FAFE" blockId="15.[160,1156,1355,1554]" box="[160,276,1355,1381]" pageId="15" pageNumber="1720">
<emphasis id="B904903FFF94FFA3FF47FAE0AC25FAFE" bold="true" box="[160,276,1355,1381]" italics="true" pageId="15" pageNumber="1720">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BCF4C2DFF94FFA3FF47FAC5A9B5F989" blockId="15.[160,1156,1355,1554]" pageId="15" pageNumber="1720">
This new species most closely resembles
<taxonomicName id="4C7037AEFF94FFA3FD88FAC5AEADFA1C" authority="(Forel, 1892)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[623,924,1390,1415]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="1720" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B904903FFF94FFA3FD88FAC5AE3BFA1C" box="[623,778,1390,1415]" italics="true" pageId="15" pageNumber="1720">L. gracilicornis</emphasis>
(Forel, 1892)
</taxonomicName>
in all morphological aspects, but can be distinguished by its petiole, which has an acute dorsal angle that lacks dorsal spines in profile. In contrast, the petiole of
<taxonomicName id="4C7037AEFF94FFA3FD21FA1FAE79FA56" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[710,840,1460,1485]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="1720" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B904903FFF94FFA3FD21FA1FAE79FA56" box="[710,840,1460,1485]" italics="true" pageId="15" pageNumber="1720">gracilicornis</emphasis>
</taxonomicName>
has an upright anterior face and a downward-sloping posterior margin, and well-developed petiolar spines.
<taxonomicName id="4C7037AEFF94FFA3FBC4FA7DAC18F989" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="15" pageNumber="1720" phylum="Arthropoda" rank="species" species="bahrainensis">
<emphasis id="B904903FFF94FFA3FBC4FA7DAC18F989" italics="true" pageId="15" pageNumber="1720">Lepisiota bahrainensis</emphasis>
</taxonomicName>
can also be distinguished from the Arabian species by the petiole structure.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF94FFBDFF47F995A909FA13" lastPageId="17" lastPageNumber="1722" pageId="15" pageNumber="1720" type="description">
<paragraph id="8BCF4C2DFF94FFBDFF47F995A909FA13" blockId="15.[160,1156,1598,1728]" lastBlockId="17.[160,1156,1218,1416]" lastPageId="17" lastPageNumber="1722" pageId="15" pageNumber="1720">
<emphasis id="B904903FFF94FFA3FF47F995AC95F9C3" bold="true" box="[160,420,1598,1624]" italics="true" pageId="15" pageNumber="1720">Description of worker.</emphasis>
<emphasis id="B904903FFF94FFA3FE53F994ACC3F9C3" bold="true" box="[436,498,1599,1624]" pageId="15" pageNumber="1720">Head</emphasis>
. Distinctly longer than broad in full-face view, with straight posterior margin and shallowly convex lateral margins; eyes relatively large (EI 35), situated posteriorly behind midlength of head; antennal scape when laid back from its insertion surpassing the posterior margin of head by about a third of its length.
<emphasis id="B904903FFF8AFFBDFF47FB69AC11FB40" bold="true" box="[160,288,1218,1243]" pageId="17" pageNumber="1722">Mesosoma</emphasis>
. Propodeal spines well developed and sharp.
<emphasis id="B904903FFF8AFFBDFCCAFB69AEB1FB40" bold="true" box="[813,896,1218,1243]" pageId="17" pageNumber="1722">Petiole</emphasis>
. In profile the anterior face meets the posterior face at a characteristically acute angle and lacks dorsal spines.
<emphasis id="B904903FFF8AFFBDFF47FAACADC9FABB" bold="true" box="[160,248,1287,1312]" pageId="17" pageNumber="1722">Pilosity</emphasis>
. Anterior clypeal margin with several pairs of long yellow setae; cephalic surface with scatted appressed pubescence; pronotum with a single pair of setae; rest of mesosoma without setae; gastral tergites with several pairs of setae.
<emphasis id="B904903FFF8AFFBDFCBCFAE6AEFFFAFD" bold="true" box="[859,974,1357,1382]" pageId="17" pageNumber="1722">Sculpture</emphasis>
. Overall smooth and shining.
<emphasis id="B904903FFF8AFFBDFED5FAC4ACB3FA13" bold="true" box="[306,386,1391,1416]" pageId="17" pageNumber="1722">Colour</emphasis>
. Uniform black; antennae, legs, and mandibles yellow-brown.
</paragraph>
</subSubSection>
<caption id="DF0F1CA5FF8BFFBCFF47F971AF9DF895" ID-DOI="http://doi.org/10.5281/zenodo.14241342" ID-Zenodo-Dep="14241342" httpUri="https://zenodo.org/record/14241342/files/figure.png" pageId="16" pageNumber="1721" startId="16.[160,225,1754,1777]" targetBox="[388,927,144,1719]" targetPageId="16" targetType="figure">
<paragraph id="8BCF4C2DFF8BFFBCFF47F971AF9DF895" blockId="16.[160,1157,1754,1806]" pageId="16" pageNumber="1721">
<emphasis id="B904903FFF8BFFBCFF47F971ADCAF96A" bold="true" box="[160,251,1754,1777]" pageId="16" pageNumber="1721">Figure 9.</emphasis>
<taxonomicName id="4C7037AEFF8BFFBCFEE4F971AC9DF96A" baseAuthorityName="Andre" baseAuthorityYear="1881" box="[259,428,1754,1777]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="16" pageNumber="1721" phylum="Arthropoda" rank="species" species="livida">
<emphasis id="B904903FFF8BFFBCFEE4F971AC9DF96A" box="[259,428,1754,1777]" italics="true" pageId="16" pageNumber="1721">Cataglyphis livida</emphasis>
</taxonomicName>
. Worker. (A) Body in profile. (B) Head in full-face view, ANTWEB1041918, AntWeb.org (Michele Esposito). (C) Distribution map.
</paragraph>
</caption>
<caption id="DF0F1CA5FF8AFFBDFF47FBB3AC7CFBF1" ID-DOI="http://doi.org/10.5281/zenodo.14241345" ID-Zenodo-Dep="14241345" httpUri="https://zenodo.org/record/14241345/files/figure.png" pageId="17" pageNumber="1722" startId="17.[160,225,1048,1071]" targetBox="[165,1150,144,1014]" targetPageId="17" targetType="figure">
<paragraph id="8BCF4C2DFF8AFFBDFF47FBB3AC7CFBF1" blockId="17.[160,1157,1048,1130]" pageId="17" pageNumber="1722">
<emphasis id="B904903FFF8AFFBDFF47FBB3AC37FBB4" bold="true" box="[160,262,1048,1071]" pageId="17" pageNumber="1722">Figure 10.</emphasis>
<taxonomicName id="4C7037AEFF8AFFBDFEEBFBB3ACEFFBB4" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" box="[268,478,1048,1071]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="1722" phylum="Arthropoda" rank="species" species="bahrainensis" status="sp. nov.">
<emphasis id="B904903FFF8AFFBDFEEBFBB3ACEFFBB4" box="[268,478,1048,1071]" italics="true" pageId="17" pageNumber="1722">Lepisiota bahrainensis</emphasis>
</taxonomicName>
<emphasis id="B904903FFF8AFFBDFE03FBB3AF2DFBB4" bold="true" box="[484,540,1048,1071]" pageId="17" pageNumber="1722">
<taxonomicNameLabel id="A2372D44FF8AFFBDFE03FBB3AF2DFBB4" box="[484,540,1048,1071]" pageId="17" pageNumber="1722" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
Holotype worker. (A) Body in profile. (B) Head in full-face view. (C) Propodeum and petiole in dorsal view, ANTWEB1041925, AntWeb.org (Michele Esposito). (D) Distribution map.
</paragraph>
</caption>
<subSubSection id="C36A1FA6FF8AFFBDFF47FA1FAE1AFA55" box="[160,811,1460,1486]" pageId="17" pageNumber="1722" type="etymology">
<paragraph id="8BCF4C2DFF8AFFBDFF47FA1FAE1AFA55" blockId="17.[160,811,1460,1486]" box="[160,811,1460,1486]" pageId="17" pageNumber="1722">
<emphasis id="B904903FFF8AFFBDFF47FA1FAC12FA55" bold="true" box="[160,291,1460,1486]" italics="true" pageId="17" pageNumber="1722">Etymology.</emphasis>
Named after the country of the
<typeStatus id="54CBF28FFF8AFFBDFD7BFA1EAFFFFA55" box="[668,718,1461,1486]" pageId="17" pageNumber="1722">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF8AFFBEFF47FA52AF85FEA9" lastPageId="18" lastPageNumber="1723" pageId="17" pageNumber="1722" type="diagnosis">
<paragraph id="8BCF4C2DFF8AFFBEFF47FA52AF85FEA9" blockId="17.[160,1156,1529,1728]" lastBlockId="18.[160,1156,142,306]" lastPageId="18" lastPageNumber="1723" pageId="17" pageNumber="1722">
<emphasis id="B904903FFF8AFFBDFF47FA52AC97F988" bold="true" box="[160,422,1529,1555]" italics="true" pageId="17" pageNumber="1722">Differential diagnosis.</emphasis>
This new species cannot be identified with Sharaf
<emphasis id="B904903FFF8AFFBDFBD2FA51A95CF988" box="[1077,1133,1530,1555]" italics="true" pageId="17" pageNumber="1722">et al</emphasis>
.s (2020b) key to
<taxonomicName id="4C7037AEFF8AFFBDFE83F9B6AF1AF9AD" box="[356,555,1565,1590]" pageId="17" pageNumber="1722">
<emphasis id="B904903FFF8AFFBDFE83F9B6ACF9F9AD" box="[356,456,1565,1590]" italics="true" pageId="17" pageNumber="1722">Lepisiota</emphasis>
species
</taxonomicName>
of the Arabian Peninsula.
<taxonomicName id="4C7037AEFF8AFFBDFC65F9B6A9B5F9AD" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" box="[898,1156,1565,1590]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="1722" phylum="Arthropoda" rank="species" species="bahrainensis">
<emphasis id="B904903FFF8AFFBDFC65F9B6A9B5F9AD" box="[898,1156,1565,1590]" italics="true" pageId="17" pageNumber="1722">Lepisiota bahrainensis</emphasis>
</taxonomicName>
appears to be most similar to
<taxonomicName id="4C7037AEFF8AFFBDFDC7F994AE5BF9C3" authority="(Forel, 1892)" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[544,874,1599,1624]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="1722" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B904903FFF8AFFBDFDC7F994AFF6F9C3" box="[544,711,1599,1624]" italics="true" pageId="17" pageNumber="1722">L. gracilicornis</emphasis>
(Forel, 1892)
</taxonomicName>
from
<collectingCountry id="F3670CBDFF8AFFBDFC5BF994A93EF9C3" box="[956,1039,1599,1624]" name="Yemen" pageId="17" pageNumber="1722">Yemen</collectingCountry>
. The two species are uniform black with paler appendages, with exceptionally long scapes (SI 190200), short and acute propodeal spines, and pronotum with a single pair of setae. However,
<taxonomicName id="4C7037AEFF8AFFBDFE8CF90CAF2BF95B" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" box="[363,538,1703,1728]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="17" pageNumber="1722" phylum="Arthropoda" rank="species" species="bahrainensis">
<emphasis id="B904903FFF8AFFBDFE8CF90CAF2BF95B" box="[363,538,1703,1728]" italics="true" pageId="17" pageNumber="1722">L. bahrainensis</emphasis>
</taxonomicName>
can be easily distinguished by the lack of dorsal petiole spines. In the key to Arabian
<taxonomicName id="4C7037AEFF89FFBEFD7FFF25A9B5FF3C" authority="(Sharaf et al. 2020 b, p. 131)" baseAuthorityName="Sharaf" baseAuthorityPageNumber="131" baseAuthorityYear="2020" box="[664,1156,142,167]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFF89FFBEFD7FFF25AFCEFF3C" box="[664,767,142,167]" italics="true" pageId="18" pageNumber="1723">Lepisiota</emphasis>
(
<bibRefCitation id="EFE131DCFF89FFBEFCF0FF25A923FF3C" author="Sharaf MR &amp; Aldawood AS &amp; Mohamed AA &amp; Hita Garcia F" box="[791,1042,142,167]" pageId="18" pageNumber="1723" pagination="127 - 152" refId="ref23777" refString="Sharaf MR, Aldawood AS, Mohamed AA, Hita Garcia F. 2020 b. The genus Lepisiota Santschi, 1926 of the Arabian Peninsula with the description of a new species, Lepisiota elbazi sp. nov. from Oman, an updated species identification key, and assessment of zoogeographic affinities. J Hym Res. 76: 127 - 152. doi: 10.3897 / jhr. 76.50193." type="journal article" year="2020">
Sharaf
<emphasis id="B904903FFF89FFBEFC92FF25AE80FF3C" box="[885,945,142,167]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2020b
</bibRefCitation>
, p. 131)
</taxonomicName>
<taxonomicName id="4C7037AEFF89FFBEFF46FF1AAC6BFF51" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" box="[161,346,177,202]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="bahrainensis">
<emphasis id="B904903FFF89FFBEFF46FF1AAC6BFF51" box="[161,346,177,202]" italics="true" pageId="18" pageNumber="1723">L. bahrainensis</emphasis>
</taxonomicName>
will key to couplet 12 along with
<taxonomicName id="4C7037AEFF89FFBEFCA8FF1AA933FF51" baseAuthorityName="Forel" baseAuthorityYear="1892" box="[847,1026,177,202]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B904903FFF89FFBEFCA8FF1AA933FF51" box="[847,1026,177,202]" italics="true" pageId="18" pageNumber="1723">L. gracilicornis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7037AEFF89FFBEFBFCFF1AA94CFF51" baseAuthorityName="Dalla Torre" baseAuthorityYear="1893" box="[1051,1149,177,202]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="nigra">
<emphasis id="B904903FFF89FFBEFBFCFF1AA94CFF51" box="[1051,1149,177,202]" italics="true" pageId="18" pageNumber="1723">L. nigra</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7037AEFF89FFBEFF46FF7FAC06FF76" authorityName="Sharaf and Monks" authorityYear="2016" box="[161,311,212,237]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="omanensis">
<emphasis id="B904903FFF89FFBEFF46FF7FAC06FF76" box="[161,311,212,237]" italics="true" pageId="18" pageNumber="1723">L. omanensis</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7037AEFF89FFBEFEADFF7FACC0FF76" baseAuthorityName="Finzi" baseAuthorityYear="1936" box="[330,497,212,237]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="opaciventris">
<emphasis id="B904903FFF89FFBEFEADFF7FACC0FF76" box="[330,497,212,237]" italics="true" pageId="18" pageNumber="1723">L. opaciventris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7037AEFF89FFBEFDD0FF7FAFD4FF76" baseAuthorityName="Kuznetsov-Ugamsky" baseAuthorityYear="1929" box="[567,741,212,237]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="spinisquama">
<emphasis id="B904903FFF89FFBEFDD0FF7FAFD4FF76" box="[567,741,212,237]" italics="true" pageId="18" pageNumber="1723">L. spinisquama</emphasis>
</taxonomicName>
, and the key is modified here to separate the species. Some variations in the petiole profile might likely be identified with the collection of more individuals.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF89FFBEFF27FEF0A9B2FE21" pageId="18" pageNumber="1723" type="key">
<key id="FFE11B1CFF89FFBEFF27FEF0A9B2FE21" pageId="18" pageNumber="1723">
<keyStep id="30845488FF89FFBEFF27FEF0A9B2FE21" pageId="18" pageNumber="1723">
<paragraph id="8BCF4C2DFF89FFBEFF27FEF0AC84FE0C" blockId="18.[192,1155,347,442]" pageId="18" pageNumber="1723">
<keyLead id="3081EF18FF89FFBEFF27FEF0AC84FE0C" pageId="18" pageNumber="1723">
12 Petiole dorsum rounded, without armature or spines of any description.
<taxonomicName id="4C7037AEFF89FFBEFBC4FEF0AC5CFE0C" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="bahrainensis" status="sp. nov.">
<emphasis id="B904903FFF89FFBEFBC4FEF0AC5CFE0C" italics="true" pageId="18" pageNumber="1723">Lepisiota bahrainensis</emphasis>
</taxonomicName>
<emphasis id="B904903FFF89FFBEFE92FED5AC84FE0C" bold="true" box="[373,437,382,407]" pageId="18" pageNumber="1723">
<taxonomicNameLabel id="A2372D44FF89FFBEFE92FED5AC84FE0C" box="[373,437,382,407]" pageId="18" pageNumber="1723" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BCF4C2DFF89FFBEFF27FE0AA9B2FE21" blockId="18.[192,1155,347,442]" box="[192,1155,417,442]" pageId="18" pageNumber="1723">
<keyLead id="3081EF18FF89FFBEFF27FE0AA9B2FE21" box="[192,1155,417,442]" pageId="18" pageNumber="1723"> Petiole dorsum with well-developed spines .......................................................................... 13</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
<subSubSection id="C36A1FA6FF89FFBEFF27FE48AC3AFD84" pageId="18" pageNumber="1723" type="discussion">
<paragraph id="8BCF4C2DFF89FFBEFF27FE48AC3AFD84" blockId="18.[160,1155,483,543]" pageId="18" pageNumber="1723">
However,
<taxonomicName id="4C7037AEFF89FFBEFED4FE48ACE7FE67" authorityName="Sharaf &amp; Wetterer &amp; Mohamed &amp; Georgiadis &amp; Nasser &amp; Aldawood" authorityYear="2024" box="[307,470,483,508]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="bahrainensis">
<emphasis id="B904903FFF89FFBEFED4FE48ACE7FE67" box="[307,470,483,508]" italics="true" pageId="18" pageNumber="1723">L. bahrainensis</emphasis>
</taxonomicName>
is readily separated from all Arabian species by the petiole structure.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF89FFBEFF47FDF0AF5EFDEE" box="[160,623,603,629]" pageId="18" pageNumber="1723" type="distribution">
<paragraph id="8BCF4C2DFF89FFBEFF47FDF0AF5EFDEE" blockId="18.[160,623,603,629]" box="[160,623,603,629]" pageId="18" pageNumber="1723">
<emphasis id="B904903FFF89FFBEFF47FDF0AC4DFDEE" bold="true" box="[160,380,603,629]" italics="true" pageId="18" pageNumber="1723">Geographic range.</emphasis>
Endemic to
<collectingCountry id="F3670CBDFF89FFBEFDF4FDF7AF5BFDEE" box="[531,618,604,629]" name="Bahrain" pageId="18" pageNumber="1723">Bahrain</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF89FFBEFF47FD1BAE59FC94" pageId="18" pageNumber="1723" type="biology_ecology">
<paragraph id="8BCF4C2DFF89FFBEFF47FD1BAE59FC94" blockId="18.[160,1157,688,1719]" pageId="18" pageNumber="1723">
<emphasis id="B904903FFF89FFBEFF47FD1BACAAFD51" bold="true" box="[160,411,688,714]" italics="true" pageId="18" pageNumber="1723">Ecology and biology.</emphasis>
The
<typeStatus id="54CBF28FFF89FFBEFE3AFD1AAF3EFD51" box="[477,527,689,714]" pageId="18" pageNumber="1723">type</typeStatus>
locality is an open area of wet, sandy soil with spread construction rubble and adjacent to an urban habitat. The dominant plant cover is the grey mangrove
<taxonomicName id="4C7037AEFF89FFBEFEB4FD5DAF8DFC94" authority="(Forssk.) Vierh." authorityName="Vierh." baseAuthorityName="Forssk." box="[339,700,758,783]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="18" pageNumber="1723" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="B904903FFF89FFBEFEB4FD5DAF23FC94" box="[339,530,758,783]" italics="true" pageId="18" pageNumber="1723">Avicennia marina</emphasis>
(Forssk.) Vierh.
</taxonomicName>
(
<taxonomicName id="4C7037AEFF89FFBEFD2FFD5DAE6FFC94" box="[712,862,758,783]" class="Magnoliopsida" family="Acanthaceae" kingdom="Plantae" order="Lamiales" pageId="18" pageNumber="1723" phylum="Tracheophyta" rank="family">Acanthaceae</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A1FA6FF89FFBEFF27FCB2AC2AF92C" pageId="18" pageNumber="1723" type="discussion">
<paragraph id="8BCF4C2DFF89FFBEFF27FCB2AFB2FB49" blockId="18.[160,1157,688,1719]" pageId="18" pageNumber="1723">
The formicine ant genus
<taxonomicName id="4C7037AEFF89FFBEFE12FCB2AE3EFCA9" authority="Santschi, 1926" authorityName="Santschi" authorityYear="1926" box="[501,783,793,818]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFF89FFBEFE12FCB2AF68FCA9" box="[501,601,793,818]" italics="true" pageId="18" pageNumber="1723">Lepisiota</emphasis>
<bibRefCitation id="EFE131DCFF89FFBEFD82FCB2AE3EFCA9" author="Santschi F" box="[613,783,793,818]" pageId="18" pageNumber="1723" pagination="13 - 28" refId="ref23043" refString="Santschi F. 1926. Trois notes myrmecologiques. Ann Soc Entomol Fr. 95 (1): 13 - 28. doi: 10.1080 / 21686351.1926. 12280094." type="journal article" year="1926">Santschi, 1926</bibRefCitation>
</taxonomicName>
was treated as a subgenus of
<taxonomicName id="4C7037AEFF89FFBEFF47FC90ACF3FCCF" authority="Mayr, 1861" authorityName="Mayr" authorityYear="1861" box="[160,450,827,852]" class="Insecta" family="Formicidae" genus="Acantholepis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFF89FFBEFF47FC90AC00FCCF" box="[160,305,827,852]" italics="true" pageId="18" pageNumber="1723">Acantholepis</emphasis>
<bibRefCitation id="EFE131DCFF89FFBEFED9FC90ACF3FCCF" author="Mayr G" box="[318,450,827,852]" pageId="18" pageNumber="1723" refId="ref21841" refString="Mayr G. 1861. Die europaischen Formiciden. Nach der analytischen Methode bearbeitet. Wien: C. Gerolds Sohn; p. 80. doi: 10.5962 / bhl. title. 14089." type="journal volume" year="1861">Mayr, 1861</bibRefCitation>
</taxonomicName>
for many years before it was considered a junior homonym of the latter genus, and the first available replacement name was
<taxonomicName id="4C7037AEFF89FFBEFBC7FCF5AC62FC01" authority="Santschi, 1926" authorityName="Santschi" authorityYear="1926" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFF89FFBEFBC7FCF5A9B5FCEC" box="[1056,1156,862,887]" italics="true" pageId="18" pageNumber="1723">Lepisiota</emphasis>
<bibRefCitation id="EFE131DCFF89FFBEFF47FC2AAC62FC01" author="Santschi F" box="[160,339,897,922]" pageId="18" pageNumber="1723" pagination="13 - 28" refId="ref23043" refString="Santschi F. 1926. Trois notes myrmecologiques. Ann Soc Entomol Fr. 95 (1): 13 - 28. doi: 10.1080 / 21686351.1926. 12280094." type="journal article" year="1926">Santschi, 1926</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFE131DCFF89FFBEFE8EFC2AAF33FC01" author="Bolton B" box="[361,514,897,922]" pageId="18" pageNumber="1723" refId="ref18966" refString="Bolton B. 1995. A new general catalogue of the ants of the world. Cambridge (MA): Harvard University Press; p. 504." type="journal volume" year="1995">
<collectingRegion id="49B482CFFF89FFBEFE8EFC2AAC89FC01" box="[361,440,897,922]" country="United Kingdom" name="Bolton" pageId="18" pageNumber="1723">Bolton</collectingRegion>
1995
</bibRefCitation>
). The genus is one of the diverse genera of the subfamily
<taxonomicName id="4C7037AEFF89FFBEFEFAFC08ACACFC27" authorityName="Latreille" authorityYear="1802" box="[285,413,931,956]" class="Insecta" family="Formicidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="subFamily" subFamily="Formicinae">Formicinae</taxonomicName>
with 102 valid species and 45 described subspecies (
<bibRefCitation id="EFE131DCFF89FFBEFBD1FC08ADE8FC44" author="Bolton B" pageId="18" pageNumber="1723" refId="ref18995" refString="Bolton B 2023. An online catalog of the ants of the world. [accessed on 2023 Nov 3]. http: // antcat. org /." type="url" year="2023">
<collectingRegion id="49B482CFFF89FFBEFBD1FC08A9B5FC27" box="[1078,1156,931,956]" country="United Kingdom" name="Bolton" pageId="18" pageNumber="1723">Bolton</collectingRegion>
2023
</bibRefCitation>
) that is widely distributed throughout the Afrotropical,
<collectingRegion id="49B482CFFF89FFBEFC3AFC6DA971FC44" box="[989,1088,966,991]" country="Saudi Arabia" name="Ash Sharqiyah" pageId="18" pageNumber="1723">Oriental</collectingRegion>
, and Palaearctic regions (
<bibRefCitation id="EFE131DCFF89FFBEFE47FC42AF06FB99" author="Brown WL Jr." box="[416,567,1001,1026]" pageId="18" pageNumber="1723" pagination="45 - 79" refId="ref19178" refString="Brown WL Jr. 2000. Diversity of ants. In: Agosti D, Majer JD, Alonso LE, Schultz TR, editors. Ants. Standard methods for measuring and monitoring biodiversity. Biological diversity Handbook Series. Washington (D. C): Smithsonian Institution Press; p. 45 - 79." type="book chapter" year="2000">Brown 2000</bibRefCitation>
), where nests are built directly in the earth, under stones and other objects, and in rotten wood (
<bibRefCitation id="EFE131DCFF89FFBEFC9EFBA0A944FBBF" author="Sharaf MR &amp; Gotzek D &amp; Guenard B &amp; Fisher BL &amp; Aldawood AS &amp; Al Dhafer HM &amp; Mohamed AA" box="[889,1141,1035,1060]" pageId="18" pageNumber="1723" refId="ref24330" refString="Sharaf MR, Gotzek D, Guenard B, Fisher BL, Aldawood AS, Al Dhafer HM, Mohamed AA. 2020 c. Molecular phylogenetic analysis and morphological reassessments of thief ants identify a new potential case of biological invasions. Sc Rep. 10 (1): 12040. doi: 10.1038 / s 41598 - 020 - 69029 - 4." type="journal volume" year="2020">
Sharaf
<emphasis id="B904903FFF89FFBEFC3EFBA0A927FBBF" box="[985,1046,1035,1060]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2020c
</bibRefCitation>
). Member species can be diagnosed readily by the combination of the following characters in the worker caste: (
<bibRefCitation id="EFE131DCFF89FFBEFDDEFBFAAFFFFBF1" author="Bolton B" box="[569,718,1105,1130]" pageId="18" pageNumber="1723" refId="ref18943" refString="Bolton B. 1994. Identification guide to the Ant Genera of the world. Cambridge: Mass; p. 222." type="journal volume" year="1994">
<collectingRegion id="49B482CFFF89FFBEFDDEFBFAAFB6FBF1" box="[569,647,1105,1130]" country="United Kingdom" name="Bolton" pageId="18" pageNumber="1723">Bolton</collectingRegion>
1994
</bibRefCitation>
): eyes prominent, ocelli frequently present but sometimes reduced; antennae 11-segmented; propodeum with a pair of spines, teeth, or tubercles; petiole with the dorsal margin bispinose, bidentate or emarginated; acidopore well developed.
</paragraph>
<paragraph id="8BCF4C2DFF89FFBEFF27FB70AC2AF92C" blockId="18.[160,1157,688,1719]" pageId="18" pageNumber="1723">
<taxonomicName id="4C7037AEFF89FFBEFF27FB70AC10FB6F" authorityName="Santschi" authorityYear="1926" box="[192,289,1243,1268]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFF89FFBEFF27FB70AC10FB6F" box="[192,289,1243,1268]" italics="true" pageId="18" pageNumber="1723">Lepisiota</emphasis>
</taxonomicName>
is considered moderately diverse compared with other formicine genera in the Arabian Peninsula (eg
<taxonomicName id="4C7037AEFF89FFBEFE28FB55AFC8FA8C" authority="Foerster, 1850" authorityName="Foerster" authorityYear="1850" box="[463,761,1278,1303]" class="Insecta" family="Formicidae" genus="Cataglyphis" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFF89FFBEFE28FB55AF61FA8C" box="[463,592,1278,1303]" italics="true" pageId="18" pageNumber="1723">Cataglyphis</emphasis>
Foerster, 1850
</taxonomicName>
(28 spp.),
<taxonomicName id="4C7037AEFF89FFBEFC95FB55A9B2FA8C" authority="Mayr, 1861" authorityName="Mayr" authorityYear="1861" box="[882,1155,1278,1303]" class="Insecta" family="Formicidae" genus="Camponotus" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="genus">
<emphasis id="B904903FFF89FFBEFC95FB55AECFFA8C" box="[882,1022,1278,1303]" italics="true" pageId="18" pageNumber="1723">Camponotus</emphasis>
<bibRefCitation id="EFE131DCFF89FFBEFBE1FB55A9B2FA8C" author="Mayr G" box="[1030,1155,1278,1303]" pageId="18" pageNumber="1723" refId="ref21841" refString="Mayr G. 1861. Die europaischen Formiciden. Nach der analytischen Methode bearbeitet. Wien: C. Gerolds Sohn; p. 80. doi: 10.5962 / bhl. title. 14089." type="journal volume" year="1861">Mayr, 1861</bibRefCitation>
</taxonomicName>
(25 spp.)) with 21 species and subspecies; all of them are reported from the
<collectingCountry id="F3670CBDFF89FFBEFBB0FA8AA9B2FAA1" box="[1111,1155,1313,1338]" name="Saudi Arabia" pageId="18" pageNumber="1723">KSA</collectingCountry>
(
<bibRefCitation id="EFE131DCFF89FFBEFF41FAE8AC4CFAC7" author="Collingwood CA" box="[166,381,1347,1372]" pageId="18" pageNumber="1723" pagination="230 - 302" refId="ref19269" refString="Collingwood CA. 1985. Hymenoptera: fam. Formicidae of Saudi Arabia. Fauna Saudi Arabia. 7: 230 - 302. doi: 10.5281 / zenodo. 26692." type="journal article" year="1985">Collingwood 1985</bibRefCitation>
;
<bibRefCitation id="EFE131DCFF89FFBEFE77FAE8AFC2FAC7" author="Collingwood CA &amp; Agosti D" box="[400,755,1347,1372]" pageId="18" pageNumber="1723" pagination="300 - 385" refId="ref19301" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
;
<bibRefCitation id="EFE131DCFF89FFBEFCE0FAE8AEEDFAC7" author="Sharaf MR &amp; Salman S &amp; Aldawood AS" box="[775,988,1347,1372]" pageId="18" pageNumber="1723" pagination="195 - 214" refId="ref24535" refString="Sharaf MR, Salman S, Aldawood AS. 2023. Additions to the ant fauna of the Kingdom of Saudi Arabia (Hymenoptera: Formicidae), with an updated list of the Saudi species. Ann Zool. 73: 195 - 214. doi: 10.3161 / 00034541 ANZ 2023.73.2.005." type="journal article" year="2023">
Sharaf
<emphasis id="B904903FFF89FFBEFCBCFAE8AEA0FAC7" box="[859,913,1347,1372]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2023
</bibRefCitation>
), followed by
<collectingCountry id="F3670CBDFF89FFBEFF47FACDADD4FAE4" box="[160,229,1382,1407]" name="Oman" pageId="18" pageNumber="1723">Oman</collectingCountry>
(11 spp.) (
<bibRefCitation id="EFE131DCFF89FFBEFE84FACDAF0FFAE4" author="Sharaf MR &amp; Al Dhafer HM &amp; Aldawood AS &amp; Hita Garcia F" box="[355,574,1382,1407]" pageId="18" pageNumber="1723" refId="ref24023" refString="Sharaf MR, Al Dhafer HM, Aldawood AS, Hita Garcia F. 2018 a. Ants of the Monomorium monomorium species-group (Hymenoptera: Formicidae) in the Arabian Peninsula with description of a new species from southwestern Saudi Arabia. PeerJ. 6: 31. e 4277. doi: 10.7717 / peerj. 4277." type="journal volume" year="2018">
Sharaf
<emphasis id="B904903FFF89FFBEFE51FACDACD8FAE4" box="[438,489,1382,1407]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2018
<bibRefCitation id="EFE131DCFF89FFBEFDD6FACDAF0FFAE4" author="Bar A &amp; Marom C &amp; Zorin N &amp; Gilad T &amp; Subach A &amp; Foitzik S &amp; Scharf I" box="[561,574,1382,1407]" pageId="18" pageNumber="1723" refId="ref18589" refString="Bar A, Marom C, Zorin N, Gilad T, Subach A, Foitzik S, Scharf I. 2022. Desert ants learn to avoid pitfall traps while foraging. Biology. 11 (6): 897. doi: 10.3390 / biology 11060897." type="journal volume" year="2022">a</bibRefCitation>
</bibRefCitation>
, 2022),
<collectingCountry id="F3670CBDFF89FFBEFD7BFACDAFDAFAE4" box="[668,747,1382,1407]" name="Yemen" pageId="18" pageNumber="1723">Yemen</collectingCountry>
(10 spp.) (
<bibRefCitation id="EFE131DCFF89FFBEFC8DFACDADE6FA39" author="Collingwood CA &amp; Agosti D" pageId="18" pageNumber="1723" pagination="300 - 385" refId="ref19301" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
), the
<collectingCountry id="F3670CBDFF89FFBEFEF8FA22AC7FFA39" box="[287,334,1417,1442]" name="United Arab Emirates" pageId="18" pageNumber="1723">UAE</collectingCountry>
(8 spp.) (
<bibRefCitation id="EFE131DCFF89FFBEFE58FA22AE2AFA39" author="Collingwood CA &amp; Agosti D" box="[447,795,1417,1442]" pageId="18" pageNumber="1723" pagination="300 - 385" refId="ref19301" refString="Collingwood CA, Agosti D. 1996. Formicidae (Insecta: Hymenoptera) of Saudi Arabia (part 2). Fauna Saudi Arabia. 15: 300 - 385." type="journal article" year="1996">Collingwood and Agosti 1996</bibRefCitation>
;
<bibRefCitation id="EFE131DCFF89FFBEFCCCFA22A971FA39" author="Collingwood CA &amp; Agosti D &amp; Sharaf MR &amp; van Harten A" box="[811,1088,1417,1442]" pageId="18" pageNumber="1723" pagination="405 - 474" refId="ref19333" refString="Collingwood CA, Agosti D, Sharaf MR, van Harten A. 2011. Order Hymenoptera, family Formicidae. Arthropod Fauna UAE. 4: 405 - 474. doi: 10.5281 / zenodo. 1168586." type="journal article" year="2011">
Collingwood
<emphasis id="B904903FFF89FFBEFC21FA22AEC8FA39" box="[966,1017,1417,1442]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2011
</bibRefCitation>
), and finally
<collectingCountry id="F3670CBDFF89FFBEFF0BFA00AC1BFA5F" box="[236,298,1451,1476]" name="Qatar" pageId="18" pageNumber="1723">Qatar</collectingCountry>
(2 spp.) (
<bibRefCitation id="EFE131DCFF89FFBEFE72FA00AF5CFA5F" author="Sharaf MR &amp; Aldawood AS &amp; Mohamed AA &amp; Hita Garcia F" box="[405,621,1451,1476]" pageId="18" pageNumber="1723" pagination="127 - 152" refId="ref23777" refString="Sharaf MR, Aldawood AS, Mohamed AA, Hita Garcia F. 2020 b. The genus Lepisiota Santschi, 1926 of the Arabian Peninsula with the description of a new species, Lepisiota elbazi sp. nov. from Oman, an updated species identification key, and assessment of zoogeographic affinities. J Hym Res. 76: 127 - 152. doi: 10.3897 / jhr. 76.50193." type="journal article" year="2020">
Sharaf
<emphasis id="B904903FFF89FFBEFE00FA00AF29FA5F" box="[487,536,1451,1476]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2020b
</bibRefCitation>
). Recently, a couple of species were described from the region,
<taxonomicName id="4C7037AEFF89FFBEFE85FA65AE3BFA7C" authority="Sharaf and Monks, 2016" authorityName="Sharaf and Monks" authorityYear="2016" box="[354,778,1486,1511]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="omanensis">
<emphasis id="B904903FFF89FFBEFE85FA65ACC0FA7C" box="[354,497,1486,1511]" italics="true" pageId="18" pageNumber="1723">L. omanensis</emphasis>
<bibRefCitation id="EFE131DCFF89FFBEFE1EFA65AE3BFA7C" author="Sharaf MR &amp; Monks J &amp; Polaszek A &amp; Aldawood AS" box="[505,778,1486,1511]" pageId="18" pageNumber="1723" pagination="1875 - 1887" refId="ref24468" refString="Sharaf MR, Monks J, Polaszek A, Aldawood AS. 2016 a. A remarkable new species of the genus Lepisiota Santschi (Hymenoptera: Formicidae) from Oman and the United Arab Emirates with a key to the Arabian species. J Nat Hist. 50 (29 - 30): 1875 - 1887. doi: 10.1080 / 00222933.2016. 1180722." type="journal article" year="2016">Sharaf and Monks, 2016</bibRefCitation>
</taxonomicName>
(
<bibRefCitation id="EFE131DCFF89FFBEFCFFFA65AEDDFA7C" author="Sharaf MR &amp; Monks J &amp; Polaszek A &amp; Aldawood AS" box="[792,1004,1486,1511]" pageId="18" pageNumber="1723" pagination="1875 - 1887" refId="ref24468" refString="Sharaf MR, Monks J, Polaszek A, Aldawood AS. 2016 a. A remarkable new species of the genus Lepisiota Santschi (Hymenoptera: Formicidae) from Oman and the United Arab Emirates with a key to the Arabian species. J Nat Hist. 50 (29 - 30): 1875 - 1887. doi: 10.1080 / 00222933.2016. 1180722." type="journal article" year="2016">
Sharaf
<emphasis id="B904903FFF89FFBEFC8EFA65AEA8FA7C" box="[873,921,1486,1511]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2016a
</bibRefCitation>
), from
<collectingCountry id="F3670CBDFF89FFBEFBD8FA65A9B5FA7C" box="[1087,1156,1486,1511]" name="Oman" pageId="18" pageNumber="1723">Oman</collectingCountry>
and the
<collectingCountry id="F3670CBDFF89FFBEFEE3FA5AAC07F991" box="[260,310,1521,1546]" name="United Arab Emirates" pageId="18" pageNumber="1723">UAE</collectingCountry>
, and
<taxonomicName id="4C7037AEFF89FFBEFE90FA5AAE17F991" authority="Sharaf and Hita Garcia, 2020" authorityName="Sharaf and Hita Garcia" authorityYear="2020" box="[375,806,1521,1546]" class="Insecta" family="Formicidae" genus="Lepisiota" kingdom="Animalia" order="Hymenoptera" pageId="18" pageNumber="1723" phylum="Arthropoda" rank="species" species="elbazi">
<emphasis id="B904903FFF89FFBEFE90FA5AACE3F991" box="[375,466,1521,1546]" italics="true" pageId="18" pageNumber="1723">L. elbazi</emphasis>
<bibRefCitation id="EFE131DCFF89FFBEFE3BFA5AAE17F991" author="Sharaf MR &amp; Aldawood AS &amp; Mohamed AA &amp; Hita Garcia F" box="[476,806,1521,1546]" pageId="18" pageNumber="1723" pagination="127 - 152" refId="ref23777" refString="Sharaf MR, Aldawood AS, Mohamed AA, Hita Garcia F. 2020 b. The genus Lepisiota Santschi, 1926 of the Arabian Peninsula with the description of a new species, Lepisiota elbazi sp. nov. from Oman, an updated species identification key, and assessment of zoogeographic affinities. J Hym Res. 76: 127 - 152. doi: 10.3897 / jhr. 76.50193." type="journal article" year="2020">Sharaf and Hita Garcia, 2020</bibRefCitation>
</taxonomicName>
from the
<collectingRegion id="49B482CFFF89FFBEFC78FA5AA9B5F991" box="[927,1156,1521,1546]" country="Oman" name="Zufar" pageId="18" pageNumber="1723">Dhofar Governorate</collectingRegion>
(
<collectingCountry id="F3670CBDFF89FFBEFF41F9BFADDFF9B6" box="[166,238,1556,1581]" name="Oman" pageId="18" pageNumber="1723">Oman</collectingCountry>
) (
<bibRefCitation id="EFE131DCFF89FFBEFEE4F9BFACECF9B6" author="Sharaf MR &amp; Aldawood AS &amp; Mohamed AA &amp; Hita Garcia F" box="[259,477,1556,1581]" pageId="18" pageNumber="1723" pagination="127 - 152" refId="ref23777" refString="Sharaf MR, Aldawood AS, Mohamed AA, Hita Garcia F. 2020 b. The genus Lepisiota Santschi, 1926 of the Arabian Peninsula with the description of a new species, Lepisiota elbazi sp. nov. from Oman, an updated species identification key, and assessment of zoogeographic affinities. J Hym Res. 76: 127 - 152. doi: 10.3897 / jhr. 76.50193." type="journal article" year="2020">
Sharaf
<emphasis id="B904903FFF89FFBEFEB2F9BFACB6F9B6" box="[341,391,1556,1581]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2020b
</bibRefCitation>
). The most remarkable contributions on the genus (
<bibRefCitation id="EFE131DCFF89FFBEFBDDF9BFAC17F9D4" author="Sharaf MR &amp; Aldawood AS &amp; Mohamed AA &amp; Hita Garcia F" pageId="18" pageNumber="1723" pagination="127 - 152" refId="ref23777" refString="Sharaf MR, Aldawood AS, Mohamed AA, Hita Garcia F. 2020 b. The genus Lepisiota Santschi, 1926 of the Arabian Peninsula with the description of a new species, Lepisiota elbazi sp. nov. from Oman, an updated species identification key, and assessment of zoogeographic affinities. J Hym Res. 76: 127 - 152. doi: 10.3897 / jhr. 76.50193." type="journal article" year="2020">
Sharaf
<emphasis id="B904903FFF89FFBEFF47F99DADE0F9D4" box="[160,209,1590,1615]" italics="true" pageId="18" pageNumber="1723">et al</emphasis>
. 2020b
</bibRefCitation>
) reviewed and keyed the fauna in the Arabian Peninsula and presented an assessment of the zoogeographical affinities dividing the fauna into two categories, an Afrotropical category with 11 species (52%) and a Palaearctic category with 10 species (47%).
</paragraph>
</subSubSection>
</treatment>
</document>