treatments-xml/data/D6/5A/87/D65A87DF33077450FF10DB6BFDD3FDF5.xml

<document id="73C4B9D188AE994B26AE5C83816954A6" ID-DOI="10.11646/phytotaxa.207.3.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13637261" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725336952098" checkinUser="felipe" docAuthor="Wisitrassameewong, Komsit, Nuytinck, Jorinde, Le, Huyen Thanh, Crop, Eske De, Hampe, Felix &amp; Verbeken, Annemieke" docDate="2015" docId="D65A87DF33077450FF10DB6BFDD3FDF5" docLanguage="en" docName="phytotaxa.207.3.1.pdf" docOrigin="Phytotaxa 207 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.207.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Lactarius fuscomaculatus Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken 2015, sp. nov." docType="treatment" docVersion="2" lastPageNumber="228" masterDocId="2A63FFA7330C745DFF98D941FFF5FFD4" masterDocTitle="Lactarius subgenus Russularia (Russulaceae) in South-East Asia: 3. new diversity in Thailand and Vietnam" masterLastPageNumber="241" masterPageNumber="215" pageNumber="226" updateTime="1725341186633" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="9A0C91A9CFCB03C5A961DB3B5B2A6DA1">Lactarius subgenus Russularia (Russulaceae) in South-East Asia: 3. new diversity in Thailand and Vietnam</mods:title>
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<mods:namePart id="229C034044B8076A493C43285FDBA856">Wisitrassameewong, Komsit</mods:namePart>
<mods:affiliation id="B4994D4DE98F7443A945D96B2C5FFA1D">School of Science, Mae Fah Luang University, 333 Moo 1, Thasud sub-district, Muang district, Chiang Rai 57100, Thailand &amp; Research Group Mycology, Department of Biology, Gent University, K. L. Ledeganckstraat 35, 9000 Gent, Belgium</mods:affiliation>
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<mods:namePart id="4BC0990607358E0B5D1A5945C0D7CB68">Nuytinck, Jorinde</mods:namePart>
<mods:affiliation id="69928F6E490790D36190ED76C385B1DF">Naturalis Biodiversity Center, Section National Herbarium of the Netherlands, P. O. Box 9517, 2300 RA Leiden, The Netherlands</mods:affiliation>
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<mods:namePart id="1AC35132E05BF1D1BDC57CC96C92F8E6">Le, Huyen Thanh</mods:namePart>
<mods:affiliation id="2056241F56FBBD8E6618F66B5F2A61DA">Faculty of Environment, Hanoi University of Natural Resources and Environment, 41 A Street K 1, Cau Dien, Tu Liem, Hanoi, Vietnam</mods:affiliation>
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<mods:namePart id="61D32E05859FC2F5BECD725BE26E81D8">Crop, Eske De</mods:namePart>
<mods:affiliation id="89861B121BA729B8AD4356C3D7E4F1C0">Research Group Mycology, Department of Biology, Gent University, K. L. Ledeganckstraat 35, 9000 Gent, Belgium</mods:affiliation>
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<mods:namePart id="BD638C4415874E1DA92AAB497511F3E7">Hampe, Felix</mods:namePart>
<mods:affiliation id="129040F732EF5A9649B719FE600A7852">Research Group Mycology, Department of Biology, Gent University, K. L. Ledeganckstraat 35, 9000 Gent, Belgium &amp; School of Science, Mae Fah Luang University, 333 Moo 1, Thasud sub-district, Muang district, Chiang Rai 57100, Thailand</mods:affiliation>
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<mods:namePart id="EC95096F7A4A53340434A36AEA1CADBA">Verbeken, Annemieke</mods:namePart>
<mods:affiliation id="F881355B5E6D235061DFE7623B4C288C">Research Group Mycology, Department of Biology, Gent University, K. L. Ledeganckstraat 35, 9000 Gent, Belgium</mods:affiliation>
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<mods:title id="D3A64D9FA656E8B40BC29DBC064E9A0F">Phytotaxa</mods:title>
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<paragraph id="5E4C36C933077456FF10DB6BFB97FD91" blockId="11.[136,1122,554,581]" box="[136,1122,554,581]" pageId="11" pageNumber="226">
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<taxonomicName id="99F34D4A33077456FF10DB6BFCDEFD91" ID-CoL="6NSNL" authority="Wisitrassameewong &amp; Verbeken" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[136,811,554,581]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="11" pageNumber="226" phylum="Basidiomycota" rank="species" species="fuscomaculatus" status="sp. nov.">
<emphasis id="6C87EADB33077456FF10DB6BFE41FD91" bold="true" box="[136,436,554,581]" italics="true" pageId="11" pageNumber="226">Lactarius fuscomaculatus</emphasis>
Wisitrassameewong &amp; Verbeken
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, 
<emphasis id="6C87EADB33077456FCAEDB6DFC7EFD91" box="[822,907,556,581]" italics="true" pageId="11" pageNumber="226">
<taxonomicNameLabel id="77B457A033077456FCAEDB6DFC7EFD91" box="[822,907,556,581]" pageId="11" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="C6C82A4C33077456FC01DB6DFC0CFD91" box="[921,1017,556,581]" captionStart="FIGURE 5" captionStartId="12.[136,229,2030,2051]" captionTargetBox="[230,1367,344,1978]" captionTargetId="figure-59@12.[200,1388,317,2004]" captionTargetPageId="12" captionText="FIGURE 5. L. fuscomaculatus: a. basidiospore, b. cheilocystidia, c. marginal cell, d. basidia, e. pseudocystidia, f. pleuromacrocystidia, g. pileipellis (a and c−f: KW365, b and g: KW373, holotype) (scale bar = 10 μm)." figureDoi="http://doi.org/10.5281/zenodo.13637275" httpUri="https://zenodo.org/record/13637275/files/figure.png" pageId="11" pageNumber="226">Figure 5</figureCitation>
and 
<figureCitation id="C6C82A4C33077456FBB7DB6DFBACFD91" box="[1071,1113,556,581]" captionStart="FIGURE 11" captionStartId="22.[136,229,1862,1883]" captionTargetBox="[151,1435,190,1837]" captionTargetId="figure-17@22.[151,1436,190,1837]" captionTargetPageId="22" captionText="FIGURE 11. basidiocarps a. L. aquosus (KW231), b. L.fuscomaculatus (KW373), c. L.rubrobrunneus (AV12-044, photo byA.Verbeken), d. L. inconspicuus (KW339, photo by F. Hampe), e–f. L. tangerinus (EDC14-475, photo by E. De Crop), g–h. L. kesiyae (g: KW032, h: KW353), i. L. rubrocorrugatus (KW384), j. L. chichuensis (KW271)." figureDoi="http://doi.org/10.5281/zenodo.13637287" httpUri="https://zenodo.org/record/13637287/files/figure.png" pageId="11" pageNumber="226">11b</figureCitation>
)
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<emphasis id="6C87EADB33077456FF10DB35FE90FD59" box="[136,357,628,653]" italics="true" pageId="11" pageNumber="226">MycoBank number:</emphasis>
MB811743; 
<emphasis id="6C87EADB33077456FE62DB35FD1DFD59" box="[506,744,628,653]" italics="true" pageId="11" pageNumber="226">Facesoffungi number</emphasis>
: FoF00636
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</subSubSection>
<subSubSection id="16E9654233077450FF25DBD9FDD3FDF5" lastPageId="13" lastPageNumber="228" pageId="11" pageNumber="226" type="diagnosis">
<paragraph id="5E4C36C933077456FF25DBD9FCD1FD2D" blockId="11.[136,1452,628,2093]" pageId="11" pageNumber="226">
<emphasis id="6C87EADB33077456FF25DBD9FECDFD65" box="[189,312,664,689]" italics="true" pageId="11" pageNumber="226">Diagnosis:</emphasis>
a medium sized species with a brown to orange-brown pileus and remarkably dark brown spots or tinges, white latex, a strong odor of 
<taxonomicName id="99F34D4A33077456FDFFDBFDFD0AFD01" authorityName="W.E.Leach" authorityYear="1815" box="[615,767,700,725]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="226" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
bugs, incompletely reticulate basidiospore ornamentation, pleuromacrocystidia rare and pileipellis a hyphoepithelium.
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<emphasis id="6C87EADB33077456FF25DA45FECBFCC9" box="[189,318,772,797]" italics="true" pageId="11" pageNumber="226">Etymology:</emphasis>
—‘fuscomaculatus’ refers to the dark, brownish spots on the pileus.
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<paragraph id="5E4C36C933077456FF25DA69FD67FA89" blockId="11.[136,1452,628,2093]" pageId="11" pageNumber="226">
<emphasis id="6C87EADB33077456FF25DA69FEF2FC95" box="[189,263,808,833]" italics="true" pageId="11" pageNumber="226">Typus:</emphasis>
— 
<collectingCountry id="26E4765933077456FEBCDA69FE42FC95" box="[292,439,808,833]" name="Thailand" pageId="11" pageNumber="226">THAILAND</collectingCountry>
, 
<collectingRegion id="9C37F82B33077456FE5EDA69FD4CFC95" box="[454,697,808,833]" country="Thailand" name="Chiang Rai" pageId="11" pageNumber="226">Chiang Rai province</collectingRegion>
, Muang district, Thasai sub-district, forest near Doi Pui Reverse Signal Station, Doi Pui, 
<geoCoordinate id="3BC7500E33077456FE00DA0DFDE5FCB1" box="[408,528,844,869]" degrees="19" direction="north" minutes="49.26" orientation="latitude" pageId="11" pageNumber="226" precision="9" value="19.821">N19º49.26</geoCoordinate>
<geoCoordinate id="3BC7500E33077456FD8FDA0DFD7AFCB1" box="[535,655,844,869]" degrees="99" direction="east" minutes="52.19" orientation="longitude" pageId="11" pageNumber="226" precision="9" value="99.869835">E99º52.19</geoCoordinate>
, alt. 
<quantity id="990B9B2C33077456FD5CDA0DFCF8FCB1" box="[708,781,844,869]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="11" pageNumber="226" unit="m" value="655.0">655 m</quantity>
, 
<date id="2A4D100933077456FC81DA0DFC6EFCB1" box="[793,923,844,869]" pageId="11" pageNumber="226" value="2012-07-31">31/07/2012</date>
, KW373 (
<typeStatus id="8148886B33077456FB96DA0DFB81FCB1" box="[1038,1140,844,869]" pageId="11" pageNumber="226" type="holotype">holotype</typeStatus>
, GENT!, 
<typeStatus id="8148886B33077456FB79DA0DFAC0FCB1" box="[1249,1333,844,869]" pageId="11" pageNumber="226" type="isotype">isotype</typeStatus>
, MFLU!). 
<emphasis id="6C87EADB33077456FF10DA31FEE9FC5D" box="[136,284,880,905]" italics="true" pageId="11" pageNumber="226">Basidiocarps</emphasis>
medium sized. 
<emphasis id="6C87EADB33077456FE4FDA31FDE8FC5D" box="[471,541,880,905]" italics="true" pageId="11" pageNumber="226">Pileus</emphasis>
<quantity id="990B9B2C33077456FDBFDA31FD50FC5D" box="[551,677,880,905]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" metricValueMax="6.2" metricValueMin="2.5" pageId="11" pageNumber="226" unit="mm" value="43.5" valueMax="62.0" valueMin="25.0">25−62 mm</quantity>
diam., broadly convex initially, turning slightly infundibuliform to deeply infundibuliform in age, with a more or less distinct papilla; surface dry, more rugose in the center in age, brown (7D8 to 7E7) to dark brown (8F7) in the center, towards the margin, paler, brownish orange (5C5) to brown (6D7) to pale yellow (3A3), typically becoming uneven in color in age, with dark brown (8F7) discolorations, varying from spotted to brushed over the whole surface except the margin; margin indistinctly striate, slightly crenulate. 
<emphasis id="6C87EADB33077456FADBDD41FA5EFBCD" box="[1347,1451,1024,1049]" italics="true" pageId="11" pageNumber="226">Lamellae</emphasis>
subdecurrent to decurrent, 
<quantity id="990B9B2C33077456FE28DD65FDFBFBE9" box="[432,526,1060,1085]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="3.0" metricValueMin="1.0" pageId="11" pageNumber="226" unit="mm" value="2.0" valueMax="3.0" valueMin="1.0">1−3 mm</quantity>
broad, crowded, with 3−4 series of lamellulae, pale yellow (4A3) to cream-colored, discoloring light brown (6D6) to brown (7E6), spotted in age. 
<emphasis id="6C87EADB33077456FCB7DD09FC92FBB5" box="[815,871,1096,1121]" italics="true" pageId="11" pageNumber="226">Stipe</emphasis>
34−76 x 
<quantity id="990B9B2C33077456FC54DD09FBDEFBB5" box="[972,1067,1096,1121]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="11" pageNumber="226" unit="mm" value="5.5" valueMax="7.0" valueMin="4.0">4−7 mm</quantity>
, cylindrical, central, rarely slightly eccentric; surface dry, smooth, whitish pruinose, particularly in young specimens, dull yellow (3B3) to olive brown (4D6) at the apex, yellowish brown (5E4) to dark brown (7F5) towards the base, sometimes brownish orange (7C7), with 
<quantity id="990B9B2C33077456FF5ADDF5FED4FB19" box="[194,289,1204,1229]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="11" pageNumber="226" unit="mm" value="1.5" valueMax="2.0" valueMin="1.0">1−2 mm</quantity>
long hairs at the base. 
<emphasis id="6C87EADB33077456FDBFDDF5FD8BFB19" box="[551,638,1204,1229]" italics="true" pageId="11" pageNumber="226">Context</emphasis>
<quantity id="990B9B2C33077456FD1EDDF5FD0EFB19" box="[646,763,1204,1229]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" metricValueMax="4.0" metricValueMin="0.5" pageId="11" pageNumber="226" unit="mm" value="2.25" valueMax="4.0" valueMin="0.5">0.5−4 mm</quantity>
broad in the pileus, partially hollow to completely hollow in stipe, pale yellow (3A3); odor strong, reminiscent of 
<taxonomicName id="99F34D4A33077456FD47DD99FCBBFB25" box="[735,846,1240,1265]" class="Agaricomycetes" family="Russulaceae" genus="Lactifluus" kingdom="Fungi" order="Russulales" pageId="11" pageNumber="226" phylum="Basidiomycota" rank="species" species="quietus">
<emphasis id="6C87EADB33077456FD47DD99FCBBFB25" box="[735,846,1240,1265]" italics="true" pageId="11" pageNumber="226">L. quietus</emphasis>
</taxonomicName>
or 
<taxonomicName id="99F34D4A33077456FCECDD99FBF9FB25" authorityName="W.E.Leach" authorityYear="1815" box="[884,1036,1240,1265]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="11" pageNumber="226" phylum="Arthropoda" rank="family">Pentatomidae</taxonomicName>
bugs; taste mild. 
<emphasis id="6C87EADB33077456FB4CDD99FAE7FB25" box="[1236,1298,1240,1265]" italics="true" pageId="11" pageNumber="226">Latex</emphasis>
watery white to white, abundant, unchanging on exposure, unchanging with 10% KOH, unchanging on white tissue paper and on white cotton handkerchief; taste slightly astringent and becoming faintly peppery. 
<emphasis id="6C87EADB33077456FB80DC61FADCFAED" box="[1048,1321,1312,1337]" italics="true" pageId="11" pageNumber="226">Macrochemical reaction</emphasis>
no reaction on the context with 10% KOH, or with FeSO 
<subScript id="C277348C33077456FD1ADC14FD7FFAB7" attach="left" box="[642,650,1365,1379]" fontSize="6" pageId="11" pageNumber="226">4</subScript>
.
</paragraph>
<paragraph id="5E4C36C933077456FF25DC29FBCDF93D" blockId="11.[136,1452,628,2093]" pageId="11" pageNumber="226">
<emphasis id="6C87EADB33077456FF25DC29FEAEFA55" box="[189,347,1384,1409]" italics="true" pageId="11" pageNumber="226">Basidiospores</emphasis>
globose to broadly ellipsoid, 6.0− 
<emphasis id="6C87EADB33077456FD42DC29FCC5FA55" box="[730,816,1384,1409]" italics="true" pageId="11" pageNumber="226">6.6−7.2</emphasis>
−7.7(−7.8) × 5.5− 
<emphasis id="6C87EADB33077456FC60DC29FBBBFA55" box="[1016,1102,1384,1409]" italics="true" pageId="11" pageNumber="226">6.1−6.6</emphasis>
.−7.0(−7.2) μm, Q = 1.02− 
<emphasis id="6C87EADB33077456FAE2DC29FF3CFA71" italics="true" pageId="11" pageNumber="226">1.08 −1.10</emphasis>
−1.21 (n=100); ornamentation amyloid, composed of irregular ridges up to 1 μm high, forming an incomplete reticulum, warts and ridges connected by fine lines; isolated warts present; plage inamyloid to distally amyloid. 
<emphasis id="6C87EADB33077456FACEDCF1FA59FA1D" box="[1366,1452,1456,1481]" italics="true" pageId="11" pageNumber="226">Basidia</emphasis>
42−53 × 13−17 μm, 4-spored, some 2-spored, subclavate to clavate; with guttate contents. 
<emphasis id="6C87EADB33077456FB11DC95FA87FA39" box="[1161,1394,1492,1517]" italics="true" pageId="11" pageNumber="226">Pleuromacrocystidia</emphasis>
rare, protruding up to10μm,55−68×13−17μm,fusiform,with a mucronate,occasionally ramified apex. 
<emphasis id="6C87EADB33077456FB22DCB9FA59F9C5" box="[1210,1452,1528,1553]" italics="true" pageId="11" pageNumber="226">Pleuropseudocystidia</emphasis>
scarce to abundant, 3−6 μm diam., not emergent to slightly emergent, cylindrical to tortuous with an obtuse apex, often broadened at the apex. 
<emphasis id="6C87EADB33077456FE08DF01FDC1F98D" box="[400,564,1600,1625]" italics="true" pageId="11" pageNumber="226">Lamellar edge</emphasis>
heterogeneous, composed of basidia, abundant cylindrical to subclavate, thin-walled marginal cells 13−26 × 5−13 μm; cheilocystidia absent to rare, 34−38 ×11−14 μm, not emergent, fusiform with a mucronate apex. 
<emphasis id="6C87EADB33077456FEC4DFC9FDF8F975" box="[348,525,1672,1697]" italics="true" pageId="11" pageNumber="226">Lamellar trama</emphasis>
consisting of lactifers and sphaerocysts. 
<emphasis id="6C87EADB33077456FC42DFC9FBB9F975" box="[986,1100,1672,1697]" italics="true" pageId="11" pageNumber="226">Pileipellis</emphasis>
a hyphoepithelium, with a thin upper layer of repent or oblique hyphae; suprapellis composed of cylindrical hyphae, 10−20 μm thick; subpellis a layer of large globose cells, up to 30 μm diam., mixed with cylindrical to inflated hyphae.
</paragraph>
<paragraph id="5E4C36C933077456FF25DFB5FB53F8D9" blockId="11.[136,1452,628,2093]" box="[189,1190,1780,1805]" pageId="11" pageNumber="226">
<emphasis id="6C87EADB33077456FF25DFB5FEE9F8D9" box="[189,284,1780,1805]" italics="true" pageId="11" pageNumber="226">Habitat:</emphasis>
gregarious to scattered on the ground in montane tropical forests with 
<taxonomicName id="99F34D4A33077456FBAEDFB5FB57F8D9" authorityName="Dumortier" authorityYear="1829" box="[1078,1186,1780,1805]" class="Magnoliopsida" family="Fagaceae" kingdom="Plantae" order="Fagales" pageId="11" pageNumber="226" phylum="Tracheophyta" rank="family">Fagaceae</taxonomicName>
.
</paragraph>
<paragraph id="5E4C36C933077451FF25DE59FF05FEF1" blockId="11.[136,1452,628,2093]" lastBlockId="12.[136,1451,160,293]" lastPageId="12" lastPageNumber="227" pageId="11" pageNumber="226">
<emphasis id="6C87EADB33077456FF25DE59FE4EF8E5" box="[189,443,1816,1841]" italics="true" pageId="11" pageNumber="226">Collections examined:</emphasis>
<materialsCitation id="EE9B3C9433077456FE5DDE59FEBCF8AD" collectingDate="2012-07-31" collectionCode="GENT, MFLU" country="Thailand" county="Muang district" elevation="655" latitude="19.821" location="Doi Pui Reverse Signal Station" longLatPrecision="12" longitude="99.869835" municipality="Thasai" pageId="11" pageNumber="226" specimenCode="KW373" specimenCount="2" stateProvince="Chiang Rai" typeStatus="holotype">
<collectingCountry id="26E4765933077456FE5DDE59FDACF8E5" box="[453,601,1816,1841]" name="Thailand" pageId="11" pageNumber="226">THAILAND</collectingCountry>
, 
<collectingRegion id="9C37F82B33077456FDFEDE59FCA3F8E5" box="[614,854,1816,1841]" country="Thailand" name="Chiang Rai" pageId="11" pageNumber="226">Chiang Rai province</collectingRegion>
, 
<collectingCounty id="B72D4E4533077456FCFCDE59FBFBF8E5" box="[868,1038,1816,1841]" pageId="11" pageNumber="226">Muang district</collectingCounty>
, 
<collectingMunicipality id="BE28ACB333077456FB83DE59FB93F8E5" box="[1051,1126,1816,1841]" pageId="11" pageNumber="226">Thasai</collectingMunicipality>
sub-district, forest near 
<location id="5B2C601233077456FA19DE59FE30F881" LSID="urn:lsid:plazi:treatment:D65A87DF33077450FF10DB6BFDD3FDF5:5B2C601233077456FA19DE59FE30F881" country="Thailand" county="Muang district" latitude="19.821" longLatPrecision="12" longitude="99.869835" municipality="Thasai" name="Doi Pui Reverse Signal Station" pageId="11" pageNumber="226" stateProvince="Chiang Rai">Doi Pui Reverse Signal Station</location>
, 
<location id="5B2C601233077456FE4DDE7DFDC6F881" LSID="urn:lsid:plazi:treatment:D65A87DF33077450FF10DB6BFDD3FDF5:5B2C601233077456FE4DDE7DFDC6F881" box="[469,563,1852,1877]" country="Thailand" county="Muang district" latitude="19.821" longLatPrecision="12" longitude="99.869835" municipality="Thasai" name="Doi Pui" pageId="11" pageNumber="226" stateProvince="Chiang Rai">Doi Pui</location>
, 
<geoCoordinate id="3BC7500E33077456FDDBDE7DFD4FF881" box="[579,698,1852,1877]" degrees="19" direction="north" minutes="49.26" orientation="latitude" pageId="11" pageNumber="226" precision="9" value="19.821">N19º49.26</geoCoordinate>
<geoCoordinate id="3BC7500E33077456FD5EDE7DFCCBF881" box="[710,830,1852,1877]" degrees="99" direction="east" minutes="52.19" orientation="longitude" pageId="11" pageNumber="226" precision="9" value="99.869835">E99º52.19</geoCoordinate>
, alt. 
<quantity id="990B9B2C33077456FCE5DE7DFC3EF881" box="[893,971,1852,1877]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="11" pageNumber="226" unit="m" value="655.0">
<elevation id="D5DED1FA33077456FCE5DE7DFC3EF881" box="[893,971,1852,1877]" metricMagnitude="2" metricUnit="m" metricValue="6.55" pageId="11" pageNumber="226" unit="m" value="655.0">655 m</elevation>
</quantity>
, 
<date id="2A4D100933077456FC43DE7DFBABF881" box="[987,1118,1852,1877]" pageId="11" pageNumber="226" value="2012-07-31">
<collectingDate id="3A09E9E133077456FC43DE7DFBABF881" box="[987,1118,1852,1877]" pageId="11" pageNumber="226" value="2012-07-31">31/07/2012</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FBF6DE7DFB32F881" box="[1134,1223,1852,1877]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW373</specimenCode>
(
<typeStatus id="8148886B33077456FB42DE7DFAB5F881" box="[1242,1344,1852,1877]" pageId="11" pageNumber="226" type="holotype">holotype</typeStatus>
, 
<collectionCode id="38E2AE0C33077456FAC8DE7DFA56F881" box="[1360,1443,1852,1877]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<typeStatus id="8148886B33077456FF10DE21FF29F8AD" box="[136,220,1888,1913]" pageId="11" pageNumber="226" type="isotype">isotype</typeStatus>
, 
<collectionCode id="38E2AE0C33077456FF7EDE21FECBF8AD" box="[230,318,1888,1913]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077456FEC6DE21FCA6F8AD" box="[350,851,1888,1913]" collectingDate="2011-07-25" collectionCode="GENT, MFLU" country="Thailand" county="Muang district" elevation="655" latitude="19.821" location="Doi Pui Reverse Signal Station" longLatPrecision="12" longitude="99.869835" municipality="Thasai" pageId="11" pageNumber="226" specimenCode="KW111" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933077456FE05DE21FDEBF8AD" box="[413,542,1888,1913]" pageId="11" pageNumber="226" value="2011-07-25">
<collectingDate id="3A09E9E133077456FE05DE21FDEBF8AD" box="[413,542,1888,1913]" pageId="11" pageNumber="226" value="2011-07-25">25/07/2011</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FDB1DE21FD8AF8AD" box="[553,639,1888,1913]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW111</specimenCode>
(
<collectionCode id="38E2AE0C33077456FD15DE21FD14F8AD" box="[653,737,1888,1913]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FD77DE21FCB2F8AD" box="[751,839,1888,1913]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077456FCFFDE21FAA8F8AD" box="[871,1373,1888,1913]" collectingDate="2011-07-25" collectionCode="GENT, MFLU" country="Thailand" county="Muang district" elevation="655" latitude="19.821" location="Doi Pui Reverse Signal Station" longLatPrecision="12" longitude="99.869835" municipality="Thasai" pageId="11" pageNumber="226" specimenCode="KW112" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933077456FC3FDE21FBDDF8AD" box="[935,1064,1888,1913]" pageId="11" pageNumber="226" value="2011-07-25">
<collectingDate id="3A09E9E133077456FC3FDE21FBDDF8AD" box="[935,1064,1888,1913]" pageId="11" pageNumber="226" value="2011-07-25">25/07/2011</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FBAADE21FB7FF8AD" box="[1074,1162,1888,1913]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW112</specimenCode>
(
<collectionCode id="38E2AE0C33077456FB00DE21FB19F8AD" box="[1176,1260,1888,1913]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FB62DE21FAA7F8AD" box="[1274,1362,1888,1913]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077456FAEADE21FDB8F849" collectingDate="2011-09-01" collectionCode="GENT, MFLU" country="Thailand" county="Muang district" elevation="655" latitude="19.821" location="Doi Pui Reverse Signal Station" longLatPrecision="12" longitude="99.869835" municipality="Thasai" pageId="11" pageNumber="226" specimenCode="KW126" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933077456FF10DEC5FEFFF849" box="[136,266,1924,1949]" pageId="11" pageNumber="226" value="2011-09-01">
<collectingDate id="3A09E9E133077456FF10DEC5FEFFF849" box="[136,266,1924,1949]" pageId="11" pageNumber="226" value="2011-09-01">01/09/2011</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FE80DEC5FE85F849" box="[280,368,1924,1949]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW126</specimenCode>
(
<collectionCode id="38E2AE0C33077456FE1BDEC5FE22F849" box="[387,471,1924,1949]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FE72DEC5FDB7F849" box="[490,578,1924,1949]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077456FDFADEC5FB9FF849" box="[610,1130,1924,1949]" collectingDate="2011-09-01" collectionCode="GENT, MFLU" country="Thailand" county="Muang district" elevation="655" latitude="19.821" location="Doi Pui Reverse Signal Station" longLatPrecision="12" longitude="99.869835" municipality="Thasai" pageId="11" pageNumber="226" specimenCode="KW128" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933077456FD3DDEC5FCD2F849" box="[677,807,1924,1949]" pageId="11" pageNumber="226" value="2011-09-01">
<collectingDate id="3A09E9E133077456FD3DDEC5FCD2F849" box="[677,807,1924,1949]" pageId="11" pageNumber="226" value="2011-09-01">01/09/2011</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FCADDEC5FC7BF849" box="[821,910,1924,1949]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW128</specimenCode>
(
<collectionCode id="38E2AE0C33077456FC39DEC5FC00F849" box="[929,1013,1924,1949]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FB9FDEC5FBAAF849" box="[1031,1119,1924,1949]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077456FBE7DEC5FEA2F815" collectingDate="2011-09-01" collectionCode="GENT, MFLU" country="Thailand" county="Muang district" elevation="655" latitude="19.821" location="Doi Pui Reverse Signal Station" longLatPrecision="12" longitude="99.869835" municipality="Thasai" pageId="11" pageNumber="226" specimenCode="KW129" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933077456FB5BDEC5FAB1F849" box="[1219,1348,1924,1949]" pageId="11" pageNumber="226" value="2011-09-01">
<collectingDate id="3A09E9E133077456FB5BDEC5FAB1F849" box="[1219,1348,1924,1949]" pageId="11" pageNumber="226" value="2011-09-01">01/09/2011</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FACBDEC5FA5EF849" box="[1363,1451,1924,1949]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW129</specimenCode>
(
<collectionCode id="38E2AE0C33077456FF08DEE9FF11F815" box="[144,228,1960,1985]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FF6CDEE9FEB9F815" box="[244,332,1960,1985]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077456FEF4DEE9FC9CF815" box="[364,873,1960,1985]" collectingDate="2012-07-25" collectionCode="GENT, MFLU" country="Thailand" county="Muang district" elevation="655" latitude="19.821" location="Doi Pui Reverse Signal Station" longLatPrecision="12" longitude="99.869835" municipality="Thasai" pageId="11" pageNumber="226" specimenCode="KW370" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933077456FE35DEE9FDDAF815" box="[429,559,1960,1985]" pageId="11" pageNumber="226" value="2012-07-25">
<collectingDate id="3A09E9E133077456FE35DEE9FDDAF815" box="[429,559,1960,1985]" pageId="11" pageNumber="226" value="2012-07-25">25/07/2012</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FDA2DEE9FD66F815" box="[570,659,1960,1985]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW370</specimenCode>
(
<collectionCode id="38E2AE0C33077456FD3BDEE9FD02F815" box="[675,759,1960,1985]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FC9EDEE9FCABF815" box="[774,862,1960,1985]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
; 
<materialsCitation id="EE9B3C9433077456FCEDDEE9FF19F7DD" collectingDate="2012-05-30" collectionCode="GENT, MFLU" country="Thailand" county="Mae Fah Luang district" elevation="1269" latitude="20.281666" location="Doi Mae Salong" municipality="Doi Mae Salong Nok" pageId="11" pageNumber="226" specimenCode="KW221" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
<collectingRegion id="9C37F82B33077456FCEDDEE9FB94F815" box="[885,1121,1960,1985]" country="Thailand" name="Chiang Rai" pageId="11" pageNumber="226">Chiang Rai province</collectingRegion>
, 
<collectingCounty id="B72D4E4533077456FBF4DEE9FA83F815" box="[1132,1398,1960,1985]" pageId="11" pageNumber="226">Mae Fah Luang district</collectingCounty>
, 
<collectingMunicipality id="BE28ACB333077456FA19DEE9FEBEF831" pageId="11" pageNumber="226">Doi Mae Salong Nok</collectingMunicipality>
sub-district, 
<location id="5B2C601233077456FE7CDE8DFD50F831" LSID="urn:lsid:plazi:treatment:D65A87DF33077450FF10DB6BFDD3FDF5:5B2C601233077456FE7CDE8DFD50F831" box="[484,677,1996,2021]" country="Thailand" county="Mae Fah Luang district" latitude="20.281666" municipality="Doi Mae Salong Nok" name="Doi Mae Salong" pageId="11" pageNumber="226" stateProvince="Chiang Rai">Doi Mae Salong</location>
, alt. 
<quantity id="990B9B2C33077456FD47DE8DFCCCF831" box="[735,825,1996,2021]" metricMagnitude="3" metricUnit="m" metricValue="1.269" pageId="11" pageNumber="226" unit="m" value="1269.0">
<elevation id="D5DED1FA33077456FD47DE8DFCCCF831" box="[735,825,1996,2021]" metricMagnitude="3" metricUnit="m" metricValue="1.269" pageId="11" pageNumber="226" unit="m" value="1269.0">1269 m</elevation>
</quantity>
, 
<geoCoordinate id="3BC7500E33077456FCD0DE8DFC37F831" box="[840,962,1996,2021]" degrees="20" direction="north" minutes="16.90" orientation="latitude" pageId="11" pageNumber="226" precision="9" value="20.281666">N20°16.90</geoCoordinate>
E99°62.30, 
<date id="2A4D100933077456FBCCDE8DFB22F831" box="[1108,1239,1996,2021]" pageId="11" pageNumber="226" value="2012-05-30">
<collectingDate id="3A09E9E133077456FBCCDE8DFB22F831" box="[1108,1239,1996,2021]" pageId="11" pageNumber="226" value="2012-05-30">30/05/2012</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FB7DDE8DFAC8F831" box="[1253,1341,1996,2021]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW221</specimenCode>
(
<collectionCode id="38E2AE0C33077456FAD7DE8DFA56F831" box="[1359,1443,1996,2021]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FF10DEB1FF15F7DD" box="[136,224,2032,2057]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077456FE98DEB1FD01F7DD" box="[256,756,2032,2057]" collectingDate="2012-05-30" collectionCode="GENT, MFLU" country="Thailand" county="Mae Fah Luang district" elevation="1269" latitude="20.281666" location="Doi Mae Salong" municipality="Doi Mae Salong Nok" pageId="11" pageNumber="226" specimenCode="KW223" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933077456FEA6DEB1FE34F7DD" box="[318,449,2032,2057]" pageId="11" pageNumber="226" value="2012-05-30">
<collectingDate id="3A09E9E133077456FEA6DEB1FE34F7DD" box="[318,449,2032,2057]" pageId="11" pageNumber="226" value="2012-05-30">30/05/2012</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FE52DEB1FDD6F7DD" box="[458,547,2032,2057]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW223</specimenCode>
(
<collectionCode id="38E2AE0C33077456FDA8DEB1FD71F7DD" box="[560,644,2032,2057]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FD08DEB1FD1DF7DD" box="[656,744,2032,2057]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
; 
<materialsCitation id="EE9B3C9433077456FD66DEB1FAA8F7F9" collectingDate="2012-07-22" collectionCode="GENT, MFLU" country="Thailand" county="Mae Fah Luang district" elevation="1193" latitude="20.287167" location="Doi Mae Salong" municipality="Doi Mae Salong Nok" pageId="11" pageNumber="226" specimenCode="KW365" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
<collectingRegion id="9C37F82B33077456FD66DEB1FC11F7DD" box="[766,996,2032,2057]" country="Thailand" name="Chiang Rai" pageId="11" pageNumber="226">Chiang Rai province</collectingRegion>
, 
<collectingCounty id="B72D4E4533077456FC75DEB1FB05F7DD" box="[1005,1264,2032,2057]" pageId="11" pageNumber="226">Mae Fah Luang district</collectingCounty>
, 
<collectingMunicipality id="BE28ACB333077456FB60DEB1FF4DF7F9" pageId="11" pageNumber="226">Doi Mae Salong Nok</collectingMunicipality>
sub-district, 
<location id="5B2C601233077456FED4D155FDF2F7F9" LSID="urn:lsid:plazi:treatment:D65A87DF33077450FF10DB6BFDD3FDF5:5B2C601233077456FED4D155FDF2F7F9" box="[332,519,2068,2093]" country="Thailand" county="Mae Fah Luang district" latitude="20.287167" municipality="Doi Mae Salong Nok" name="Doi Mae Salong" pageId="11" pageNumber="226" stateProvince="Chiang Rai">Doi Mae Salong</location>
, 
<geoCoordinate id="3BC7500E33077456FD8AD155FD7FF7F9" box="[530,650,2068,2093]" degrees="20" direction="north" minutes="17.23" orientation="latitude" pageId="11" pageNumber="226" precision="9" value="20.287167">N20º17.23</geoCoordinate>
E99º61.69, alt. 
<quantity id="990B9B2C33077456FCA6D155FC61F7F9" box="[830,916,2068,2093]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="11" pageNumber="226" unit="m" value="1193.0">
<elevation id="D5DED1FA33077456FCA6D155FC61F7F9" box="[830,916,2068,2093]" metricMagnitude="3" metricUnit="m" metricValue="1.193" pageId="11" pageNumber="226" unit="m" value="1193.0">1193 m</elevation>
</quantity>
, 
<date id="2A4D100933077456FC38D155FBD6F7F9" box="[928,1059,2068,2093]" pageId="11" pageNumber="226" value="2012-07-22">
<collectingDate id="3A09E9E133077456FC38D155FBD6F7F9" box="[928,1059,2068,2093]" pageId="11" pageNumber="226" value="2012-07-22">22/07/2012</collectingDate>
</date>
, 
<specimenCode id="0E559EB233077456FBB6D155FB72F7F9" box="[1070,1159,2068,2093]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="11" pageNumber="226" type="Herbarium">KW365</specimenCode>
(
<collectionCode id="38E2AE0C33077456FB0ED155FB1FF7F9" box="[1174,1258,2068,2093]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="11" pageNumber="226" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33077456FB62D155FAA7F7F9" box="[1274,1362,2068,2093]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="11" pageNumber="226">MFLU</collectionCode>
!)
</materialsCitation>
;− 
<materialsCitation id="EE9B3C9433077451FAEAD155FDB4FF6D" collectingDate="2013-08-28" collectionCode="GENT, MFLU" country="Thailand" county="Mae Fah Luang district" elevation="1193" lastPageId="12" lastPageNumber="227" latitude="20.287167" location="Doi Mae Salong" municipality="Doi Mae Salong Nok" pageId="11" pageNumber="226" specimenCode="KW466" specimenCount="1" stateProvince="Chiang Rai" typeStatus="holotype">
ibid., 
<date id="2A4D100933007451FF10D9E1FEFFFF6D" box="[136,266,160,185]" pageId="12" pageNumber="227" value="2013-08-28">
<collectingDate id="3A09E9E133007451FF10D9E1FEFFFF6D" box="[136,266,160,185]" pageId="12" pageNumber="227" value="2013-08-28">28/08/2013</collectingDate>
</date>
, 
<specimenCode id="0E559EB233007451FE8DD9E1FE98FF6D" box="[277,365,160,185]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="12" pageNumber="227" type="Herbarium">KW466</specimenCode>
(
<collectionCode id="38E2AE0C33007451FEE3D9E1FE3AFF6D" box="[379,463,160,185]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="12" pageNumber="227" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33007451FE45D9E1FDC0FF6D" box="[477,565,160,185]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="12" pageNumber="227">MFLU</collectionCode>
!)
</materialsCitation>
; 
<materialsCitation id="EE9B3C9433007451FDD3D9E1FC14FF09" collectingDate="2012-06-02" collectionCode="M, GENT, MFLU" country="Thailand" elevation="1040" latitude="18.884666" location="Bahn Pok" longLatPrecision="13" longitude="99.3575" municipality="Huaikaew" pageId="12" pageNumber="227" specimenCode="KW236" specimenCount="1" stateProvince="Chiang Mai" typeStatus="holotype">
<collectingRegion id="9C37F82B33007451FDD3D9E1FCCFFF6D" box="[587,826,160,185]" country="Thailand" name="Chiang Mai" pageId="12" pageNumber="227">Chiang Mai province</collectingRegion>
, Mae-On district, 
<collectingMunicipality id="BE28ACB333007451FB9AD9E1FB81FF6D" box="[1026,1140,160,185]" pageId="12" pageNumber="227">Huaikaew</collectingMunicipality>
sub-district, 
<location id="5B2C601233007451FA9CD9E1FA84FF6D" LSID="urn:lsid:plazi:treatment:D65A87DF33077450FF10DB6BFDD3FDF5:5B2C601233007451FA9CD9E1FA84FF6D" box="[1284,1393,160,185]" country="Thailand" latitude="18.884666" longLatPrecision="13" longitude="99.3575" municipality="Huaikaew" name="Bahn Pok" pageId="12" pageNumber="227" stateProvince="Chiang Mai">Bahn Pok</location>
<collectionCode id="38E2AE0C33007451FAEED9E1FA62FF6D" box="[1398,1431,160,185]" country="Germany" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="12" pageNumber="227" type="Herbarium">M</collectionCode>
.1, 
<geoCoordinate id="3BC7500E33007451FF10D985FEF5FF09" box="[136,256,196,221]" degrees="18" direction="north" minutes="53.08" orientation="latitude" pageId="12" pageNumber="227" precision="9" value="18.884666">N18º53.08</geoCoordinate>
<geoCoordinate id="3BC7500E33007451FE91D985FE75FF09" box="[265,384,196,221]" degrees="99" direction="east" minutes="21.45" orientation="longitude" pageId="12" pageNumber="227" precision="9" value="99.3575">E99º21.45</geoCoordinate>
, alt. 
<quantity id="990B9B2C33007451FE21D985FDE7FF09" box="[441,530,196,221]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="12" pageNumber="227" unit="m" value="1040.0">
<elevation id="D5DED1FA33007451FE21D985FDE7FF09" box="[441,530,196,221]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="12" pageNumber="227" unit="m" value="1040.0">1040 m</elevation>
</quantity>
, 
<date id="2A4D100933007451FDB8D985FD57FF09" box="[544,674,196,221]" pageId="12" pageNumber="227" value="2012-06-02">
<collectingDate id="3A09E9E133007451FDB8D985FD57FF09" box="[544,674,196,221]" pageId="12" pageNumber="227" value="2012-06-02">02/06/2012</collectingDate>
</date>
, 
<specimenCode id="0E559EB233007451FD37D985FCFDFF09" box="[687,776,196,221]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="12" pageNumber="227" type="Herbarium">KW236</specimenCode>
(
<collectionCode id="38E2AE0C33007451FC81D985FC98FF09" box="[793,877,196,221]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="12" pageNumber="227" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33007451FCE6D985FC23FF09" box="[894,982,196,221]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="12" pageNumber="227">MFLU</collectionCode>
!)
</materialsCitation>
; 
<materialsCitation id="EE9B3C9433007451FC77D985FF19FEF1" collectingDate="2012-06-03" collectionCode="GENT, MFLU" country="Thailand" elevation="1450" latitude="18.857166" location="Bahn Mae Kampong" longLatPrecision="13" longitude="99.368164" municipality="Huaikaew" pageId="12" pageNumber="227" specimenCode="KW249" specimenCount="1" stateProvince="Chiang Mai" typeStatus="holotype">
<collectingRegion id="9C37F82B33007451FC77D985FB11FF09" box="[1007,1252,196,221]" country="Thailand" name="Chiang Mai" pageId="12" pageNumber="227">Chiang Mai province</collectingRegion>
, Mae-On district, 
<collectingMunicipality id="BE28ACB333007451FF10D9A9FF0FFED5" box="[136,250,232,257]" pageId="12" pageNumber="227">Huaikaew</collectingMunicipality>
sub-district, 
<location id="5B2C601233007451FE00D9A9FD65FED5" LSID="urn:lsid:plazi:treatment:D65A87DF33077450FF10DB6BFDD3FDF5:5B2C601233007451FE00D9A9FD65FED5" box="[408,656,232,257]" country="Thailand" latitude="18.857166" longLatPrecision="13" longitude="99.368164" municipality="Huaikaew" name="Bahn Mae Kampong" pageId="12" pageNumber="227" stateProvince="Chiang Mai">Bahn Mae Kampong</location>
, 
<geoCoordinate id="3BC7500E33007451FD3AD9A9FCEFFED5" box="[674,794,232,257]" degrees="18" direction="north" minutes="51.43" orientation="latitude" pageId="12" pageNumber="227" precision="9" value="18.857166">N18º51.43</geoCoordinate>
<geoCoordinate id="3BC7500E33007451FCBFD9A9FC6BFED5" box="[807,926,232,257]" degrees="99" direction="east" minutes="22.09" orientation="longitude" pageId="12" pageNumber="227" precision="9" value="99.368164">E99º22.09</geoCoordinate>
, alt. 
<quantity id="990B9B2C33007451FC47D9A9FBC9FED5" box="[991,1084,232,257]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="227" unit="m" value="1450.0">
<elevation id="D5DED1FA33007451FC47D9A9FBC9FED5" box="[991,1084,232,257]" metricMagnitude="3" metricUnit="m" metricValue="1.45" pageId="12" pageNumber="227" unit="m" value="1450.0">1450 m</elevation>
</quantity>
, 
<date id="2A4D100933007451FBD6D9A9FB24FED5" box="[1102,1233,232,257]" pageId="12" pageNumber="227" value="2012-06-03">
<collectingDate id="3A09E9E133007451FBD6D9A9FB24FED5" box="[1102,1233,232,257]" pageId="12" pageNumber="227" value="2012-06-03">03/06/2012</collectingDate>
</date>
, 
<specimenCode id="0E559EB233007451FB79D9A9FACFFED5" box="[1249,1338,232,257]" collectionCode="KW" country="Ukraine" lsid="urn:lsid:biocol.org:col:15713" name="National Academy of Sciences of Ukraine" pageId="12" pageNumber="227" type="Herbarium">KW249</specimenCode>
(
<collectionCode id="38E2AE0C33007451FAD7D9A9FA56FED5" box="[1359,1443,232,257]" country="Belgium" httpUri="http://grbio.org/cool/nz5a-xvmm" name="Ghent University, Biology Department" pageId="12" pageNumber="227" type="Herbarium">GENT</collectionCode>
!, 
<collectionCode id="38E2AE0C33007451FF10D84DFF15FEF1" box="[136,224,268,293]" country="Thailand" httpUri="http://grbio.org/cool/672j-mk16" name="Mae Fah Laung University Herbarium" pageId="12" pageNumber="227">MFLU</collectionCode>
!)
</materialsCitation>
.
</paragraph>
<caption id="0A8C664133007451FF10DEAFFC71F7F3" ID-DOI="http://doi.org/10.5281/zenodo.13637275" ID-Zenodo-Dep="13637275" httpUri="https://zenodo.org/record/13637275/files/figure.png" pageId="12" pageNumber="227" startId="12.[136,229,2030,2051]" targetBox="[230,1367,344,1978]" targetPageId="12" targetType="figure">
<paragraph id="5E4C36C933007451FF10DEAFFC71F7F3" blockId="12.[136,1452,2030,2087]" pageId="12" pageNumber="227">
<emphasis id="6C87EADB33007451FF10DEAFFF08F7D7" bold="true" box="[136,253,2030,2051]" pageId="12" pageNumber="227">FIGURE 5.</emphasis>
<taxonomicName id="99F34D4A33007451FE9ADEAFFE45F7D7" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[258,432,2030,2051]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="12" pageNumber="227" phylum="Basidiomycota" rank="species" species="fuscomaculatus">
<emphasis id="6C87EADB33007451FE9ADEAFFE45F7D7" box="[258,432,2030,2051]" italics="true" pageId="12" pageNumber="227">L. fuscomaculatus</emphasis>
</taxonomicName>
: a. basidiospore, b. cheilocystidia, c. marginal cell, d. basidia, e. pseudocystidia, 
<taxonomicName id="99F34D4A33007451FB2FDEAFFA64F7D7" box="[1207,1425,2030,2051]" form="pleuromacrocystidia" pageId="12" pageNumber="227" rank="form">f. pleuromacrocystidia</taxonomicName>
, g. pileipellis (a and c− f: KW365, b and g: KW373, holotype) (scale bar = 10 μm).
</paragraph>
</caption>
<paragraph id="5E4C36C933017450FF25D9E1FDD3FDF5" blockId="13.[136,1452,160,545]" pageId="13" pageNumber="228">
<emphasis id="6C87EADB33017450FF25D9E1FD9DFF6D" box="[189,616,160,185]" italics="true" pageId="13" pageNumber="228">
Comments: 
<taxonomicName id="99F34D4A33017450FEDED9E1FD9DFF6D" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[326,616,160,185]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="fuscomaculatus">Lactarius fuscomaculatus</taxonomicName>
</emphasis>
is widely distributed in tropical forests with 
<taxonomicName id="99F34D4A33017450FBF4D9E1FB00FF6D" box="[1132,1269,160,185]" class="Magnoliopsida" family="Fagaceae" genus="Castanopsis" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="228" phylum="Tracheophyta" rank="genus">
<emphasis id="6C87EADB33017450FBF4D9E1FB00FF6D" box="[1132,1269,160,185]" italics="true" pageId="13" pageNumber="228">Castanopsis</emphasis>
</taxonomicName>
and 
<taxonomicName id="99F34D4A33017450FAB7D9E1FA78FF6D" box="[1327,1421,160,185]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="13" pageNumber="228" phylum="Tracheophyta" rank="genus">
<emphasis id="6C87EADB33017450FAB7D9E1FA78FF6D" box="[1327,1421,160,185]" italics="true" pageId="13" pageNumber="228">Quercus</emphasis>
</taxonomicName>
in Northern 
<collectingCountry id="26E4765933017450FF6BD985FEAFFF09" box="[243,346,196,221]" name="Thailand" pageId="13" pageNumber="228">Thailand</collectingCountry>
. The species can be confused with a closely related species, 
<taxonomicName id="99F34D4A33017450FC64D985FB35FF09" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[1020,1216,196,221]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="rubrobrunneus">
<emphasis id="6C87EADB33017450FC64D985FB35FF09" box="[1020,1216,196,221]" italics="true" pageId="13" pageNumber="228">L. rubrobrunneus</emphasis>
</taxonomicName>
due to similarities in basidiocarps size,color,latex features and habitat.Both species are very similar in the immature stage, however,according to our experience, brown spots on the pileus are often found in fully mature fruiting bodies of 
<taxonomicName id="99F34D4A33017450FB0DD84DFA95FEF1" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[1173,1376,268,293]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="fuscomaculatus">
<emphasis id="6C87EADB33017450FB0DD84DFA95FEF1" box="[1173,1376,268,293]" italics="true" pageId="13" pageNumber="228">L. fuscomaculatus</emphasis>
</taxonomicName>
, while 
<taxonomicName id="99F34D4A33017450FF10D871FEA5FE9D" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[136,336,304,329]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="rubrobrunneus">
<emphasis id="6C87EADB33017450FF10D871FEA5FE9D" box="[136,336,304,329]" italics="true" pageId="13" pageNumber="228">L. rubrobrunneus</emphasis>
</taxonomicName>
is more unicolored. Under the microscope, both species have basidiospores with an incomplete reticulum and not many cystidia. A slight difference was observed in their pileipellis structures; 
<taxonomicName id="99F34D4A33017450FB7CD815FA5EFEB9" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[1252,1451,340,365]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="rubrobrunneus">
<emphasis id="6C87EADB33017450FB7CD815FA5EFEB9" box="[1252,1451,340,365]" italics="true" pageId="13" pageNumber="228">L. rubrobrunneus</emphasis>
</taxonomicName>
has in part a very thin layer of repent hyphae which is like a transition between an epithelium and a hyphoepithelium, whereas 
<taxonomicName id="99F34D4A33017450FF72D8DDFE43FE61" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[234,438,412,437]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="fuscomaculatus">
<emphasis id="6C87EADB33017450FF72D8DDFE43FE61" box="[234,438,412,437]" italics="true" pageId="13" pageNumber="228">L. fuscomaculatus</emphasis>
</taxonomicName>
has a more complete layer of repent hyphae covering the pileus. 
<taxonomicName id="99F34D4A33017450FB13D8DDFA5EFE61" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[1163,1451,412,437]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="fuscomaculatus">
<emphasis id="6C87EADB33017450FB13D8DDFA5EFE61" box="[1163,1451,412,437]" italics="true" pageId="13" pageNumber="228">Lactarius fuscomaculatus</emphasis>
</taxonomicName>
may also be confused with 
<taxonomicName id="99F34D4A33017450FE24D881FDA7FE0D" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[444,594,448,473]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="tangerinus">
<emphasis id="6C87EADB33017450FE24D881FDA7FE0D" box="[444,594,448,473]" italics="true" pageId="13" pageNumber="228">L. tangerinus</emphasis>
</taxonomicName>
in the field. 
<taxonomicName id="99F34D4A33017450FD7AD881FC39FE0D" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[738,972,448,473]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="tangerinus">
<emphasis id="6C87EADB33017450FD7AD881FC39FE0D" box="[738,972,448,473]" italics="true" pageId="13" pageNumber="228">Lactarius tangerinus</emphasis>
</taxonomicName>
has smaller basidiomata, is typically paler in color without dark brown spots and possesses transparent latex. For more details on the difference between these two species, see under 
<taxonomicName id="99F34D4A33017450FE11DB49FDEAFDF5" authorityName="Wisitrassameewong &amp; Nuytinck &amp; Le &amp; Crop &amp; Hampe &amp; Verbeken" authorityYear="2015" box="[393,543,520,545]" class="Agaricomycetes" family="Russulaceae" genus="Lactarius" kingdom="Fungi" order="Russulales" pageId="13" pageNumber="228" phylum="Basidiomycota" rank="species" species="tangerinus">
<emphasis id="6C87EADB33017450FE11DB49FDEAFDF5" box="[393,543,520,545]" italics="true" pageId="13" pageNumber="228">L. tangerinus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>