273 lines
34 KiB
XML
273 lines
34 KiB
XML
<document id="E850B0130C08266BE78DD87F861B6588" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFE76D48D6DDDD95BAEB0557" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba betlephila Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="267" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="265" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo id="31F6222B88CB409A9B6AFC12224CE830">
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<mods:title id="E6B5E3948E504492FFD835138BD3FAFA">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<mods:namePart id="ECD01D23AE1EFB1720DF384D1B50523A">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="3FFEE6231B6FD9ECFDFD0154406D7F6D">Zootaxa</mods:title>
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<mods:part id="50E68D34EC3ABA503E35AA9CF1BF210F">
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<mods:date id="4902B28CF2E9855A3F16FB541E870987">2024</mods:date>
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<mods:number id="DD748ED29189D764C974147B7E628568">2024-03-14</mods:number>
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<mods:number id="CD77C06DFB98B43637FB083C03A13886">5424</mods:number>
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<mods:url id="BF71BDC4C5B7EFCFAF5CEF342CF3632A">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
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<mods:classification id="E1ECC30A7E2F3639E7E93326F19DD12A">journal article</mods:classification>
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<mods:identifier id="1CA5AB0BF21C1E66923C62A0ACD610F0" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
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<mods:identifier id="001C9A2735BC991FBB78D0378FEB54EB" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A15C878DFFE76D48D6DDDD95BAEB0557" LSID="urn:lsid:plazi:treatment:A15C878DFFE76D48D6DDDD95BAEB0557" httpUri="http://treatment.plazi.org/id/A15C878DFFE76D48D6DDDD95BAEB0557" lastPageId="16" lastPageNumber="267" pageId="14" pageNumber="265">
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<subSubSection id="61EF6510FFE76D56D6DDDD95BC2302FE" box="[151,548,1186,1213]" pageId="14" pageNumber="265" type="nomenclature">
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<paragraph id="294A369BFFE76D56D6DDDD95BC2302FE" blockId="14.[151,548,1186,1249]" box="[151,548,1186,1213]" pageId="14" pageNumber="265">
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<heading id="720281F7FFE76D56D6DDDD95BC2302FE" bold="true" box="[151,548,1186,1213]" fontSize="11" level="1" pageId="14" pageNumber="265" reason="1">
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<emphasis id="1B81EA89FFE76D56D6DDDD95BC2302FE" bold="true" box="[151,548,1186,1213]" pageId="14" pageNumber="265">
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<taxonomicName id="EEF54D18FFE76D56D6DDDD95BFDC02FE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,475,1186,1213]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. nov.">
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<emphasis id="1B81EA89FFE76D56D6DDDD95BFDC02FE" bold="true" box="[151,475,1186,1213]" italics="true" pageId="14" pageNumber="265">Paraphloeostiba betlephila</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFE76D56D7A8DD94BC2302FE" box="[482,548,1187,1213]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFE76D56D6DDDDF0BF4402A2" box="[151,323,1222,1249]" pageId="14" pageNumber="265" type="description">
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<paragraph id="294A369BFFE76D56D6DDDDF0BF4402A2" blockId="14.[151,548,1186,1249]" box="[151,323,1222,1249]" pageId="14" pageNumber="265">
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(
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<figureCitation id="B1CE2A1EFFE76D56D6D5DDF0BEEF02A2" box="[159,232,1223,1249]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="14" pageNumber="265">Figs 6</figureCitation>
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,
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<figureCitation id="B1CE2A1EFFE76D56D6BEDDF1BF3C02A2" box="[244,315,1222,1249]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="14" pageNumber="265">32–35</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFE76D56D6DDDC38BFA60332" pageId="14" pageNumber="265" type="materials_examined">
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<paragraph id="294A369BFFE76D56D6DDDC38BFA60332" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">
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<emphasis id="1B81EA89FFE76D56D6DDDC38BF42036B" bold="true" box="[151,325,1294,1321]" pageId="14" pageNumber="265">Type material.</emphasis>
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<typeStatus id="F64E8839FFE76D56D706DC38BFB3036A" box="[332,436,1295,1321]" pageId="14" pageNumber="265" type="holotype">Holotype</typeStatus>
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<specimenCount id="3FF3FD12FFE76D56D7F6DC38BFD6036A" box="[444,465,1295,1321]" count="1" pageId="14" pageNumber="265" type="male">♂</specimenCount>
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(
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<figureCitation id="B1CE2A1EFFE76D56D7AADC38BC22036A" box="[480,549,1295,1321]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="14" pageNumber="265">Fig. 6</figureCitation>
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; dissected): ‘
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<collectingRegion id="EB31F879FFE76D56D4FFDC38BD15036A" box="[693,786,1295,1321]" country="Indonesia" name="Papua Barat" pageId="14" pageNumber="265">PAPUA</collectingRegion>
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<taxonomicNameLabel id="00B257F2FFE76D56D553DC38BD54036A" box="[793,851,1295,1321]" pageId="14" pageNumber="265" rank="genus">N.G.</taxonomicNameLabel>
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:
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<collectingRegion id="EB31F879FFE76D56D515DC38BDFE036A" box="[863,1017,1295,1321]" country="Papua New Guinea" name="Morobe" pageId="14" pageNumber="265">Morobe Prov.</collectingRegion>
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| Aseki,
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<quantity id="EE0D9B7EFFE76D56D211DC38BAB7036A" box="[1115,1200,1295,1321]" metricMagnitude="3" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="265" unit="m" value="1700.0">1700 m</quantity>
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<date id="5D4B105BFFE76D56D28EDC38BB30036A" box="[1220,1335,1295,1321]" pageId="14" pageNumber="265" value="1998-02-22">22.2.1998</date>
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, betelnut - | flower, leg. A. Riedel’ <printed>, ‘
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<typeStatus id="F64E8839FFE76D56D474DC04BCD0030E" box="[574,727,1331,1357]" pageId="14" pageNumber="265" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="EEF54D18FFE76D56D4A4DC04BDAE030E" authorityName="Steel" authorityYear="1960" box="[750,937,1331,1357]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="genus">
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<emphasis id="1B81EA89FFE76D56D4A4DC04BDAE030E" box="[750,937,1331,1357]" italics="true" pageId="14" pageNumber="265">Paraphloeostiba</emphasis>
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</taxonomicName>
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<taxonomicName id="EEF54D18FFE76D56D5F5DC04BA28030E" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[959,1071,1331,1357]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="265" phylum="Arthropoda" rank="species" species="betlephila" status="sp. nov.">
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<emphasis id="1B81EA89FFE76D56D5F5DC04BA28030E" box="[959,1071,1331,1357]" italics="true" pageId="14" pageNumber="265">betlephila</emphasis>
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</taxonomicName>
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<emphasis id="1B81EA89FFE76D56D27DDC04BA7F030E" bold="true" box="[1079,1144,1331,1357]" pageId="14" pageNumber="265">
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<taxonomicNameLabel id="00B257F2FFE76D56D27DDC04BA7F030E" box="[1079,1144,1331,1357]" pageId="14" pageNumber="265" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.
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<collectionCode id="4FE4AE5EFFE76D56D34CDC04BB1B030E" box="[1286,1308,1331,1357]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="14" pageNumber="265" type="Museum">V</collectionCode>
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. des. 2024’ <red, printed> (
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<collectionCode id="4FE4AE5EFFE76D56D70DDC60BF920332" box="[327,405,1367,1393]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="14" pageNumber="265">NMW</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFE76D48D68DDC4DBCCD0423" lastPageId="16" lastPageNumber="267" pageId="14" pageNumber="265" type="description">
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<paragraph id="294A369BFFE76D56D68DDC4DBF45039E" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">
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<emphasis id="1B81EA89FFE76D56D68DDC4DBF5E03D7" bold="true" box="[199,345,1402,1428]" pageId="14" pageNumber="265">Description.</emphasis>
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Measurements: HW: 0.30; HL: 0.23; OL: 0.10; TL: 0.02; AL: 0.57; PL: 0.25; PWmax: 0.45; PWmin: 0.38; ESL: 0.45; EW: 0.56; MTbL: 0.30; MTrL: 0.14 (MTrL 1–4: 0.04; MTrL 5: 0.10); AW: 0.57; AedL: 0.30; BL: 1.42.
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</paragraph>
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<paragraph id="294A369BFFE76D56D68DDCD0BEE4009A" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">
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Habitus as in
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<figureCitation id="B1CE2A1EFFE76D56D726DCD0BFB30042" box="[364,436,1511,1537]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="14" pageNumber="265">Fig. 6</figureCitation>
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. Body and antennomeres 6–11 brown; mouthparts, antennomere 1–5 and legs yellow. Head and neck without punctation; pronotum with indistinct, fine and moderately dense punctation, invisible in mediobasal portion; elytra with distinct, very dense, moderately large and deep punctation, finer and sparser around scutellum and along suture. Head with dense coarse isodiametric microsculpture; pronotum with very dense isodiametric microsculpture, slightly finer than that in middle portion of head; scutellum with coarse isodiametric microreticulation; microsculpture of elytra very dense, transverse, finer around scutellum and longitudinal along suture.
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</paragraph>
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<paragraph id="294A369BFFE76D56D68DDFD4BF7401F2" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">Head 1.3 times as broad as long, with short and deep narrow anteocellar foveae almost reaching level of middle length of eyes. Ocelli very small, located about at level of posterior margins of eyes; distance between ocelli slightly larger than distance between ocellus and posterior margin of eyes. Antennae with slightly transverse antennomeres 6–8 and distinctly transverse antennomeres 9–10; antennomere 5 slightly broader than 4, 6 slightly longer and broader than 5, 7–8 distinctly longer and broader than 6, 9–10 slightly longer than 8, apical antennomere about 1.8 times as long as 10.</paragraph>
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<paragraph id="294A369BFFE76D56D69ADE8CBDD801BA" blockId="14.[151,1437,1294,2041]" pageId="14" pageNumber="265">Pronotum 1.8 times as broad as long, 1.5 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; laterobasal portions slightly and widely impressed.</paragraph>
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<caption id="7D8A6613FFE66D57D6DDDF27BA2D0033" ID-DOI="http://doi.org/10.5281/zenodo.10821231" ID-Zenodo-Dep="10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266" startId="15.[151,264,1552,1576]" targetBox="[176,1428,191,1520]" targetPageId="15" targetType="figure">
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<paragraph id="294A369BFFE66D57D6DDDF27BA2D0033" blockId="15.[151,1436,1552,1649]" pageId="15" pageNumber="266">
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<emphasis id="1B81EA89FFE66D57D6DDDF27BF53006B" bold="true" box="[151,340,1552,1576]" pageId="15" pageNumber="266">FIGURES 32–38.</emphasis>
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<taxonomicName id="EEF54D18FFE66D57D713DF27BC6A006B" authorityName="Shavrin" authorityYear="2024" box="[345,621,1552,1576]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="betlephila">
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<emphasis id="1B81EA89FFE66D57D713DF27BC6A006B" box="[345,621,1552,1576]" italics="true" pageId="15" pageNumber="266">Paraphloeostiba betlephila</emphasis>
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</taxonomicName>
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(Figs. 32
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<emphasis id="1B81EA89FFE66D57D487DF27BCDD006B" bold="true" box="[717,730,1552,1576]" pageId="15" pageNumber="266">–</emphasis>
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35: holotype) and
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<taxonomicName id="EEF54D18FFE66D57D5DBDF26BA0A006B" authorityName="Shavrin" authorityYear="2023" box="[913,1037,1553,1576]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="266" phylum="Arthropoda" rank="species" species="conjuncta">
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<emphasis id="1B81EA89FFE66D57D5DBDF26BA0A006B" box="[913,1037,1553,1576]" italics="true" pageId="15" pageNumber="266">P. conjuncta</emphasis>
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</taxonomicName>
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(Figs. 36
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<emphasis id="1B81EA89FFE66D57D227DF27BA7D006B" bold="true" box="[1133,1146,1552,1576]" pageId="15" pageNumber="266">–</emphasis>
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38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm.
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</paragraph>
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</caption>
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<paragraph id="294A369BFFE66D57D68DDF89BDCF009A" blockId="15.[151,1437,1726,2040]" box="[199,968,1726,1753]" pageId="15" pageNumber="266">Elytra 1.2 times as broad as long and slightly shorter than pronotum.</paragraph>
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<paragraph id="294A369BFFE66D57D68DDFD5BD2300BF" blockId="15.[151,1437,1726,2040]" box="[199,804,1762,1789]" pageId="15" pageNumber="266">Metatarsi slightly less than twice as long as metatibia.</paragraph>
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<paragraph id="294A369BFFE66D57D68DDE31BEE90197" blockId="15.[151,1437,1726,2040]" pageId="15" pageNumber="266">
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Male. Posterior margin of abdominal tergite VIII truncate (
|
||
<figureCitation id="B1CE2A1EFFE66D57D51EDE31BDA00163" box="[852,935,1798,1824]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 34</figureCitation>
|
||
). Posterior margin of abdominal sternite VIII concave (
|
||
<figureCitation id="B1CE2A1EFFE66D57D749DE1DBF500106" box="[259,343,1834,1861]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 35</figureCitation>
|
||
). Aedeagus with wide basal portion, gradually narrowed toward very narrow apical portion, with small rounded apex; parameres slightly not reaching apex of median lobe, moderately wide, with rounded apices, each with three long apical and one preapical setae, apical half of inner margin with four to five short setae; internal sac moderately short and wide, without large sclerotized structures (
|
||
<figureCitation id="B1CE2A1EFFE66D57D5D7DEA1BDF401F3" box="[925,1011,1942,1968]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 32</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="B1CE2A1EFFE66D57D6DDDE8DBEED0197" box="[151,234,1978,2004]" captionStart="FIGURES 32–38" captionStartId="15.[151,264,1552,1576]" captionTargetBox="[176,1428,191,1520]" captionTargetId="figure-16@15.[151,1436,181,1528]" captionTargetPageId="15" captionText="FIGURES 32–38. Paraphloeostiba betlephila (Figs. 32–35: holotype) and P. conjuncta (Figs. 36–38): 32–aedeagus, parameral view, 33–aedeagus, lateral view, 34–male abdominal tergite VIII, 35–male abdominal sternite VIII, 36–female abdominal tergite VIII, 37–female abdominal sternite VIII, 38–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821231" httpUri="https://zenodo.org/record/10821231/files/figure.png" pageId="15" pageNumber="266">Fig. 33</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE66D57D68DDEE9BF9601BB" blockId="15.[151,1437,1726,2040]" box="[199,401,2014,2040]" pageId="15" pageNumber="266">Female unknown.</paragraph>
|
||
<paragraph id="294A369BFFF96D48D68DD9A1BCFC06BB" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
|
||
<emphasis id="1B81EA89FFF96D48D68DD9A1BFB506F3" bold="true" box="[199,434,150,176]" pageId="16" pageNumber="267">Comparative notes.</emphasis>
|
||
Based on the very narrow median lobe and the parameres,
|
||
<taxonomicName id="EEF54D18FFF96D48D21CD9A0BAE406F3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1110,1251,150,176]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D21CD9A0BAE406F3" box="[1110,1251,150,176]" italics="true" pageId="16" pageNumber="267">P. betlephila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D2A1D9A1BB2B06F3" bold="true" box="[1259,1324,150,176]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D2A1D9A1BB2B06F3" box="[1259,1324,150,176]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="EEF54D18FFF96D48D6FED98CBF600697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[180,359,187,213]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D6FED98CBEC20697" box="[180,197,187,212]" italics="true" pageId="16" pageNumber="267">P</emphasis>
|
||
.
|
||
<emphasis id="1B81EA89FFF96D48D699D98CBF600697" box="[211,359,187,212]" italics="true" pageId="16" pageNumber="267">atramentaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D727D98DBFAA0697" bold="true" box="[365,429,186,212]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D727D98DBFAA0697" box="[365,429,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEF54D18FFF96D48D7F0D98CBC360697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[442,561,186,212]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D7F0D98CBC360697" box="[442,561,186,212]" italics="true" pageId="16" pageNumber="267">P. barclayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D472D98DBC7F0697" bold="true" box="[568,632,186,212]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D472D98DBC7F0697" box="[568,632,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEF54D18FFF96D48D4CFD98CBD1D0697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[645,794,186,212]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D4CFD98CBD1D0697" box="[645,794,186,212]" italics="true" pageId="16" pageNumber="267">P. coprophila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D56BD98DBD670697" bold="true" box="[801,864,186,212]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D56BD98DBD670697" box="[801,864,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="EEF54D18FFF96D48D5D7D98CBDF40697" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[925,1011,186,212]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D5D7D98CBDF40697" box="[925,1011,186,212]" italics="true" pageId="16" pageNumber="267">P. steeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D5B0D98DBA3D0697" bold="true" box="[1018,1082,186,212]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D5B0D98DBA3D0697" box="[1018,1082,186,212]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
It can be distinguished from all these species by the following morphological features:
|
||
</paragraph>
|
||
<paragraph id="294A369BFFF96D48D68DD835BD760702" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
|
||
from
|
||
<taxonomicName id="EEF54D18FFF96D48D742D834BFB9075F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[264,446,259,285]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D742D834BF1E075F" box="[264,281,259,284]" italics="true" pageId="16" pageNumber="267">P</emphasis>
|
||
.
|
||
<emphasis id="1B81EA89FFF96D48D760D834BFB9075F" box="[298,446,259,284]" italics="true" pageId="16" pageNumber="267">atramentaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D782D835BC0D075F" bold="true" box="[456,522,258,284]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D782D835BC0D075F" box="[456,522,258,284]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the absence of the punctation of the head, the finer punctation of the more transverse pronotum, the longer elytra and the shorter parameres;
|
||
</paragraph>
|
||
<paragraph id="294A369BFFF96D48D68DD87DBCFE07CA" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
|
||
from
|
||
<taxonomicName id="EEF54D18FFF96D48D74ED87CBF7C0727" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,379,330,356]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D74ED87CBF7C0727" box="[260,379,330,356]" italics="true" pageId="16" pageNumber="267">P. barclayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D7C8D87DBFC60727" bold="true" box="[386,449,330,356]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D7C8D87DBFC60727" box="[386,449,330,356]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the darker body, the absence of the punctation of the head, the finer punctation on the pornotum, the longer elytra and the shorter parameres;
|
||
</paragraph>
|
||
<paragraph id="294A369BFFF96D48D68DD8A5BAB207EE" blockId="16.[151,1437,150,789]" box="[199,1205,402,429]" pageId="16" pageNumber="267">
|
||
from
|
||
<taxonomicName id="EEF54D18FFF96D48D74FD8A4BF9D07EF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,410,402,428]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D74FD8A4BF9D07EF" box="[261,410,402,428]" italics="true" pageId="16" pageNumber="267">P. coprophila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D7EBD8A5BFE707EF" bold="true" box="[417,480,402,428]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D7EBD8A5BFE707EF" box="[417,480,402,428]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the slightly shorter elytra and moderately broader parameres;
|
||
</paragraph>
|
||
<paragraph id="294A369BFFF96D48D68DD881BF42045B" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
|
||
from
|
||
<taxonomicName id="EEF54D18FFF96D48D74FD880BF5C0793" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,347,438,464]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D74FD880BF5C0793" box="[261,347,438,464]" italics="true" pageId="16" pageNumber="267">P. steeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D729D881BFA50793" bold="true" box="[355,418,438,464]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D729D881BFA50793" box="[355,418,438,464]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the paler body, the absence of the punctation of the head and the presence of elevations between punctures on the elytra, the finer punctation of the pronotum, and slightly shorter parameres with narrower apical portions.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFF96D48D68DDB14BCCD0423" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
|
||
From all these species,
|
||
<taxonomicName id="EEF54D18FFF96D48D793DB14BC6C047F" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[473,619,546,573]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="267" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFF96D48D793DB14BFED047F" box="[473,490,547,572]" italics="true" pageId="16" pageNumber="267">P</emphasis>
|
||
.
|
||
<emphasis id="1B81EA89FFF96D48D7B1DB15BC6C047F" box="[507,619,546,572]" italics="true" pageId="16" pageNumber="267">betlephila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFF96D48D43FDB15BCBF047F" bold="true" box="[629,696,546,572]" pageId="16" pageNumber="267">
|
||
<taxonomicNameLabel id="00B257F2FFF96D48D43FDB15BCBF047F" box="[629,696,546,572]" pageId="16" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by the smaller body and the details of the external and internal morphology of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61EF6510FFF96D48D68DDB5DBF2B04EB" pageId="16" pageNumber="267" type="distribution">
|
||
<paragraph id="294A369BFFF96D48D68DDB5DBF2B04EB" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
|
||
<emphasis id="1B81EA89FFF96D48D68DDB5DBF6604C7" bold="true" box="[199,353,618,644]" pageId="16" pageNumber="267">Distribution.</emphasis>
|
||
The new species is known only from the
|
||
<typeStatus id="F64E8839FFF96D48D563DB5CBD5E04C6" box="[809,857,619,645]" pageId="16" pageNumber="267">type</typeStatus>
|
||
locality in Morobi Province of eastern part of
|
||
<collectingCountry id="51E2760BFFF96D48D312DB5CBF2F04EB" name="Papua New Guinea" pageId="16" pageNumber="267">Papua New Guinea</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61EF6510FFF96D48D68DDB85BAEB0557" pageId="16" pageNumber="267" type="etymology">
|
||
<paragraph id="294A369BFFF96D48D68DDB85BB0B04B3" blockId="16.[151,1437,150,789]" pageId="16" pageNumber="267">
|
||
<emphasis id="1B81EA89FFF96D48D68DDB85BF49048F" bold="true" box="[199,334,690,716]" pageId="16" pageNumber="267">Etymology.</emphasis>
|
||
The name of the new species is derived from the fact that the
|
||
<typeStatus id="F64E8839FFF96D48D252DB85BA7D048F" box="[1048,1146,690,716]" pageId="16" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
was collected in betelnut flower (
|
||
<taxonomicName id="EEF54D18FFF96D48D6BBDBE0BF6B04B3" box="[241,364,726,752]" class="Magnoliopsida" family="Piperaceae" genus="Piper" kingdom="Plantae" order="Piperales" pageId="16" pageNumber="267" phylum="Tracheophyta" rank="species" species="betle">
|
||
<emphasis id="1B81EA89FFF96D48D6BBDBE0BF6B04B3" box="[241,364,726,752]" italics="true" pageId="16" pageNumber="267">Piper betle</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="EEF54D18FFF96D48D731DBE1BFFE04B3" box="[379,505,726,752]" class="Magnoliopsida" family="Piperaceae" kingdom="Plantae" order="Piperales" pageId="16" pageNumber="267" phylum="Tracheophyta" rank="family">Piperaceae</taxonomicName>
|
||
), the most commonly used as flavoring in chewing in Asian countries.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFF96D48D68DDBCDBAEB0557" blockId="16.[151,1437,150,789]" box="[199,1260,762,789]" pageId="16" pageNumber="267">
|
||
<emphasis id="1B81EA89FFF96D48D68DDBCDBF4C0557" bold="true" box="[199,331,762,788]" pageId="16" pageNumber="267">Bionomics.</emphasis>
|
||
The
|
||
<typeStatus id="F64E8839FFF96D48D7CEDBCDBFE10557" box="[388,486,762,788]" pageId="16" pageNumber="267" type="holotype">holotype</typeStatus>
|
||
was collected in the flower of betelnut at an elevation of
|
||
<quantity id="EE0D9B7EFFF96D48D220DBCDBAB60556" box="[1130,1201,762,789]" metricMagnitude="2" metricUnit="m" metricValue="1.7" pageId="16" pageNumber="267" unit="m" value="170.0">170 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |