266 lines
33 KiB
XML
266 lines
33 KiB
XML
<document id="F299EBD7183C3E4A45651607E8C6D835" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFE16D52D6DDDD09BF3705AE" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba atramentaria Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="261" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="259" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5616B831D84C90DDBCAA74C8FDC06E34">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
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<mods:namePart id="E4836777F813E6A4C0A85F2C071B4384">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="67426AC831D74889896B2F0DE2BF6DDE">Zootaxa</mods:title>
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<mods:part id="F94C5819E249BE82F46E68144665558B">
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<mods:date id="CDD03741F9D46143729BAB655EB9E96C">2024</mods:date>
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<mods:number id="EAE31B4A8A70DFA6AA3F600AD613DEE1">2024-03-14</mods:number>
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<mods:number id="7DCE7048A21D062DF0B73D1ADC49C859">5424</mods:number>
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<mods:classification id="3D7A617B52BA411BC0B5DEF88A21D64B">journal article</mods:classification>
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<mods:identifier id="CCC88C39F581E57DF86C662FF908B5D7" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
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<mods:identifier id="22C08C788FE68E1F1EA70AD8BB40C8C7" type="ISSN">1175-5326</mods:identifier>
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<treatment id="A15C878DFFE16D52D6DDDD09BF3705AE" LSID="urn:lsid:plazi:treatment:A15C878DFFE16D52D6DDDD09BF3705AE" httpUri="http://treatment.plazi.org/id/A15C878DFFE16D52D6DDDD09BF3705AE" lastPageId="10" lastPageNumber="261" pageId="8" pageNumber="259">
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<subSubSection id="61EF6510FFE16D50D6DDDD09BC4C021A" box="[151,587,1086,1113]" pageId="8" pageNumber="259" type="nomenclature">
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<paragraph id="294A369BFFE16D50D6DDDD09BC4C021A" blockId="8.[151,587,1086,1149]" box="[151,587,1086,1113]" pageId="8" pageNumber="259">
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<heading id="720281F7FFE16D50D6DDDD09BC4C021A" bold="true" box="[151,587,1086,1113]" fontSize="11" level="1" pageId="8" pageNumber="259" reason="1">
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<emphasis id="1B81EA89FFE16D50D6DDDD09BC4C021A" bold="true" box="[151,587,1086,1113]" pageId="8" pageNumber="259">
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<taxonomicName id="EEF54D18FFE16D50D6DDDD09BC05021A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,514,1086,1113]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. nov.">
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<emphasis id="1B81EA89FFE16D50D6DDDD09BC05021A" bold="true" box="[151,514,1086,1113]" italics="true" pageId="8" pageNumber="259">Paraphloeostiba atramentaria</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="00B257F2FFE16D50D443DD08BC4C021A" box="[521,587,1087,1113]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFE16D50D6DDDD54BF44023E" box="[151,323,1123,1149]" pageId="8" pageNumber="259" type="description">
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<paragraph id="294A369BFFE16D50D6DDDD54BF44023E" blockId="8.[151,587,1086,1149]" box="[151,323,1123,1149]" pageId="8" pageNumber="259">
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(
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<figureCitation id="B1CE2A1EFFE16D50D6D5DD54BEE0023E" box="[159,231,1123,1149]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="8" pageNumber="259">Figs 3</figureCitation>
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,
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<figureCitation id="B1CE2A1EFFE16D50D6BEDD54BF3D023E" box="[244,314,1123,1149]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="8" pageNumber="259">16–19</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFE16D50D6DDDD9CBF58034E" pageId="8" pageNumber="259" type="materials_examined">
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<paragraph id="294A369BFFE16D50D6DDDD9CBF58034E" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">
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<emphasis id="1B81EA89FFE16D50D6DDDD9CBF430287" bold="true" box="[151,324,1194,1221]" pageId="8" pageNumber="259">Type material.</emphasis>
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<typeStatus id="F64E8839FFE16D50D700DD9CBFB50286" box="[330,434,1195,1221]" pageId="8" pageNumber="259" type="holotype">Holotype</typeStatus>
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<specimenCount id="3FF3FD12FFE16D50D7F3DD9CBFC90286" box="[441,462,1195,1221]" count="1" pageId="8" pageNumber="259" type="male">♂</specimenCount>
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(
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<figureCitation id="B1CE2A1EFFE16D50D797DD9CBC270286" box="[477,544,1195,1221]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="8" pageNumber="259">Fig. 3</figureCitation>
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; dissected): ‘
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<collectingRegion id="EB31F879FFE16D50D4E5DD9CBD480286" box="[687,847,1195,1221]" country="Indonesia" name="Papua Barat" pageId="8" pageNumber="259">IRIAN JAYA</collectingRegion>
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: Paniai Prov. | Sinak,
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<quantity id="EE0D9B7EFFE16D50D274DD9CBACA0286" box="[1086,1229,1195,1221]" metricMagnitude="3" metricUnit="m" metricValue="2.1" metricValueMax="2.2" metricValueMin="2.0" pageId="8" pageNumber="259" unit="m" value="2100.0" valueMax="2200.0" valueMin="2000.0">2000-2200m</quantity>
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<date id="5D4B105BFFE16D50D295DD9CBB9B0286" box="[1247,1436,1194,1221]" pageId="8" pageNumber="259" value="1995-12-14" valueMax="1995-12-17" valueMin="1995-12-14">14. -17. 12. 1995</date>
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| leg. A. Riedel’ <printed>, ‘
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<typeStatus id="F64E8839FFE16D50D79FDDF8BC6902AA" box="[469,622,1231,1257]" pageId="8" pageNumber="259" type="holotype">HOLOTYPE</typeStatus>
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<taxonomicName id="EEF54D18FFE16D50D4CFDDF8BD4702AA" authorityName="Steel" authorityYear="1960" box="[645,832,1231,1257]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="genus">
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<emphasis id="1B81EA89FFE16D50D4CFDDF8BD4702AA" box="[645,832,1231,1257]" italics="true" pageId="8" pageNumber="259">Paraphloeostiba</emphasis>
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</taxonomicName>
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<taxonomicName id="EEF54D18FFE16D50D51DDDF8BDEC02AB" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[855,1003,1231,1256]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="259" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. nov.">
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<emphasis id="1B81EA89FFE16D50D51DDDF8BDEC02AB" box="[855,1003,1231,1256]" italics="true" pageId="8" pageNumber="259">atramentaria</emphasis>
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</taxonomicName>
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<emphasis id="1B81EA89FFE16D50D5B9DDF8BA3202AA" bold="true" box="[1011,1077,1231,1257]" pageId="8" pageNumber="259">
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<taxonomicNameLabel id="00B257F2FFE16D50D5B9DDF8BA3202AA" box="[1011,1077,1231,1257]" pageId="8" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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| Shavrin A.
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<collectionCode id="4FE4AE5EFFE16D50D289DDF8BADE02AA" box="[1219,1241,1231,1257]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="8" pageNumber="259" type="Museum">V</collectionCode>
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. des. 2024’ <red, printed> (
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<collectionCode id="4FE4AE5EFFE16D50D74FDDC4BF54034E" box="[261,339,1267,1293]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="8" pageNumber="259">NMW</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="61EF6510FFE16D52D68DDC21BC6E0556" lastPageId="10" lastPageNumber="261" pageId="8" pageNumber="259" type="description">
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<paragraph id="294A369BFFE16D50D68DDC21BB9A0316" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">
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<emphasis id="1B81EA89FFE16D50D68DDC21BF5E0373" bold="true" box="[199,345,1302,1328]" pageId="8" pageNumber="259">Description.</emphasis>
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Measurements: HW: 0.47; HL: 0.32; OL: 0.17; AL: 0.63; PL: 0.42; PWmax: 0.62; PWmin: 0.51; ESL: 0.68; EW: 0.80; MTbL: 0.40; MTrL: 0.22 (MTrL 1–4: 0.07; MTrL 5: 0.15); AW: 0.82; AedL: 0.51; BL: 2.75.
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</paragraph>
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<paragraph id="294A369BFFE16D50D68DDC68BFE300FE" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">
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Habitus as in
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<figureCitation id="B1CE2A1EFFE16D50D712DC68BF9F033A" box="[344,408,1375,1401]" captionStart="FIGURES 1–6" captionStartId="3.[151,264,1858,1882]" captionTargetBox="[164,1423,192,1823]" captionTargetId="figure-16@3.[151,1436,179,1832]" captionTargetPageId="3" captionText="FIGURES 1–6. Habitus of Omaliini: 1–Nipponophloeostiba hornabrooki (holotype), 2–Paraphloeostiba assimile (holotype), 3–P. atramentaria (holotype), 4–P. attenuata (paratype), 5–P. barclayi (paratype), 6–P. betlephila (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821216" httpUri="https://zenodo.org/record/10821216/files/figure.png" pageId="8" pageNumber="259">Fig.3</figureCitation>
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. Body and antennomeres 6–11 dark brown; mouthparts, antennomeres 1–5 and legs yellowish. Head with sparse and fine punctation, more indistinct in middle and slightly denser in infraorbital portions; neck without visible punctation; pronotum with fine and dense punctation, slightly larger than that in head, sparser in middle; scutellum without punctures; elytra with dense and fine punctation, slightly larger than that in pronotum, finer and sparser in middle, with narrow impunctate area along suture in basal half; abdominal tergites with very indistinct, fine and sparse punctation. Head with very dense and coarse microsculpture, transverse in frontal portion and distinctly isodiametric in middle; pronotum with very dense, coarse, isodiametric microsculpture; scutellum with very fine transverse meshes; elytra with dense isodiametric microreticulation, slightly larger than that in pronotum; abdominal tergites with dense microsculpture, transverse in abdominal tergites III–IV and isodiametric in abdominal tergites V–VIII.
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</paragraph>
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<paragraph id="294A369BFFE16D50D68DDFF0BD5A019E" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="259">Head slightly elevated in middle, with elongate anteriomedian depressions, 1.4 times as broad as long, with short and deep anteocellar foveae, slightly subdiagonally stretching anteriad and reaching basal third of length of eyes; postocular carina indistinct, acute. Ocelli small, located slightly below level of posterior margins of eyes; distance between ocelli about 1.6 times as long as distince between ocellus and posterior margin of eyes. Apical antennomere about 1.5 times as long as preapical segment. Antennomere 3 about as long as and distinctly narrower than 2, 4 about as wide as and about twice shorter than 3, 5 distinctly longer and broader than 4, 6–9 slightly longer and distinctly broader than 5, 10 indistinctly longer than 9, apical antennomere about 1.5 times as long as 10, from about middle gradually narrowed apicad toward subacute apex.</paragraph>
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<caption id="7D8A6613FFE06D51D6DDDF8FBD9900B0" ID-DOI="http://doi.org/10.5281/zenodo.10821225" ID-Zenodo-Dep="10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="9" pageNumber="260" startId="9.[151,264,1720,1744]" targetBox="[172,1413,189,1688]" targetPageId="9" targetType="figure">
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<paragraph id="294A369BFFE06D51D6DDDF8FBD9900B0" blockId="9.[151,1436,1720,1780]" pageId="9" pageNumber="260">
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<emphasis id="1B81EA89FFE06D51D6DDDF8FBF540093" bold="true" box="[151,339,1720,1744]" pageId="9" pageNumber="260">FIGURES 16–19.</emphasis>
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<taxonomicName id="EEF54D18FFE06D51D71DDF8FBC8C0093" authorityName="Shavrin" authorityYear="2024" box="[343,651,1720,1744]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="260" phylum="Arthropoda" rank="species" species="atramentaria">
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<emphasis id="1B81EA89FFE06D51D71DDF8FBC8C0093" box="[343,651,1720,1744]" italics="true" pageId="9" pageNumber="260">Paraphloeostiba atramentaria</emphasis>
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</taxonomicName>
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(holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm.
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</paragraph>
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</caption>
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<paragraph id="294A369BFFE06D51D68DDE70BD7D01C7" blockId="9.[151,1437,1862,2033]" pageId="9" pageNumber="260">Pronotum 1.4 times as broad as long, slightly wider than head, widest in middle, distinctly more narrowed posteriad than anteriad; posteriolateral impressions wide, shallow.</paragraph>
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<paragraph id="294A369BFFE06D51D68DDEB9BC6D01EA" blockId="9.[151,1437,1862,2033]" box="[199,618,1934,1961]" pageId="9" pageNumber="260">Elytra 1.6 times as long as pronotum.</paragraph>
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<paragraph id="294A369BFFE06D51D68DDE85BD23018F" blockId="9.[151,1437,1862,2033]" box="[199,804,1970,1997]" pageId="9" pageNumber="260">Metatarsi slightly less than twice as long as metatibia.</paragraph>
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<paragraph id="294A369BFFE06D52D68DDEE1BEEA0702" blockId="9.[151,1437,1862,2033]" lastBlockId="10.[151,1437,151,1005]" lastPageId="10" lastPageNumber="261" pageId="9" pageNumber="260">
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Male. Posterior margin of abdominal tergite VIII truncate (
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<figureCitation id="B1CE2A1EFFE06D51D51EDEE1BDA101B3" box="[852,934,2006,2032]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="9" pageNumber="260">Fig. 18</figureCitation>
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). Posterior margin of abdominal sternite VIII widely and deeply concave (
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<figureCitation id="B1CE2A1EFFE36D52D793D9A0BC2A06F2" box="[473,557,151,177]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="10" pageNumber="261">Fig. 19</figureCitation>
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). Aedeagus with wide basal portion, strongly narrowed toward narrow middle portion of elongate median lobe with small rounded apex; parameres moderately narrow, reaching apex of median lobe, each with three moderately short apical and six setae on inner margin; internal sac narrow and moderately short, with several long curved sclerotized structures in basal portion (
|
||
<figureCitation id="B1CE2A1EFFE36D52D5E3D834BDFB075E" box="[937,1020,259,285]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="10" pageNumber="261">Fig. 16</figureCitation>
|
||
). Lateral aspect of the aedeagus as in
|
||
<figureCitation id="B1CE2A1EFFE36D52D6DDD810BEED0702" box="[151,234,295,321]" captionStart="FIGURES 16–19" captionStartId="9.[151,264,1720,1744]" captionTargetBox="[172,1413,189,1688]" captionTargetId="figure-16@9.[164,1423,181,1696]" captionTargetPageId="9" captionText="FIGURES 16–19. Paraphloeostiba atramentaria (holotype): 16–aedeagus, parameral view, 17–aedeagus, lateral view, 18–male abdominal tergite VIII, 19–male abdominal sternite VIII. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821225" httpUri="https://zenodo.org/record/10821225/files/figure.png" pageId="10" pageNumber="261">Fig. 17</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DD87CBF960726" blockId="10.[151,1437,151,1005]" box="[199,401,331,357]" pageId="10" pageNumber="261">Female unknown.</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DD859BCBC0792" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
<emphasis id="1B81EA89FFE36D52D68DD859BFA807CA" bold="true" box="[199,431,366,393]" pageId="10" pageNumber="261">Comparative notes.</emphasis>
|
||
Based on the very narrow median lobe and the parameres,
|
||
<taxonomicName id="EEF54D18FFE36D52D271D858BAEA07CB" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1083,1261,367,393]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D271D858BA4B07CB" box="[1083,1100,367,392]" italics="true" pageId="10" pageNumber="261">P</emphasis>
|
||
.
|
||
<emphasis id="1B81EA89FFE36D52D213D858BAEA07CB" box="[1113,1261,367,392]" italics="true" pageId="10" pageNumber="261">atramentaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D2B8D858BB3607CA" bold="true" box="[1266,1329,367,393]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D2B8D858BB3607CA" box="[1266,1329,367,393]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="EEF54D18FFE36D52D6F9D8A4BF2B07EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[179,300,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D6F9D8A4BEC307EF" box="[179,196,403,428]" italics="true" pageId="10" pageNumber="261">P</emphasis>
|
||
.
|
||
<emphasis id="1B81EA89FFE36D52D69AD8A4BF2B07EE" box="[208,300,403,429]" italics="true" pageId="10" pageNumber="261">barclayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D778D8A4BF7707EE" bold="true" box="[306,368,403,429]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D778D8A4BF7707EE" box="[306,368,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEF54D18FFE36D52D737D8A4BC0007EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[381,519,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D737D8A4BC0007EE" box="[381,519,403,429]" italics="true" pageId="10" pageNumber="261">P. betlephila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D446D8A4BC4D07EE" bold="true" box="[524,586,403,429]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D446D8A4BC4D07EE" box="[524,586,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="EEF54D18FFE36D52D41DD8A4BCEC07EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[599,747,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D41DD8A4BCEC07EE" box="[599,747,403,429]" italics="true" pageId="10" pageNumber="261">P. coprophila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D4BAD8A4BD2907EE" bold="true" box="[752,814,403,429]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D4BAD8A4BD2907EE" box="[752,814,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, and
|
||
<taxonomicName id="EEF54D18FFE36D52D523D8A4BDB907EE" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[873,958,403,429]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D523D8A4BDB907EE" box="[873,958,403,429]" italics="true" pageId="10" pageNumber="261">P. steeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D589D8A4BA0507EE" bold="true" box="[963,1026,403,429]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D589D8A4BA0507EE" box="[963,1026,403,429]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
It can be distinguished from all these species by the following morphological features:
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DD8ECBB1207B6" blockId="10.[151,1437,151,1005]" box="[199,1301,475,501]" pageId="10" pageNumber="261">
|
||
from
|
||
<taxonomicName id="EEF54D18FFE36D52D74FD8ECBF7B07B6" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[261,380,475,501]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="barclayi" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D74FD8ECBF7B07B6" box="[261,380,475,501]" italics="true" pageId="10" pageNumber="261">P. barclayi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D7C9D8ECBFC407B6" bold="true" box="[387,451,475,501]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D7C9D8ECBFC407B6" box="[387,451,475,501]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the darker elytra, slightly narrower pronotum and the shorter parameres;
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DD8C8BD9A047E" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
from
|
||
<taxonomicName id="EEF54D18FFE36D52D74ED8C8BF97045A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[260,400,511,537]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="betlephila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D74ED8C8BF97045A" box="[260,400,511,537]" italics="true" pageId="10" pageNumber="261">P. betlephila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D7DDD8C8BFD1045A" bold="true" box="[407,470,511,537]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D7DDD8C8BFD1045A" box="[407,470,511,537]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the presence of the punctation on the head, coarser punctation of the pronotum, the narrower pronotum, the shorter elytra and slightly shorter parameres;
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DDB70BD2E04C6" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
from
|
||
<taxonomicName id="EEF54D18FFE36D52D74CDB70BF9A0422" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[262,413,583,609]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="coprophila" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D74CDB70BF9A0422" box="[262,413,583,609]" italics="true" pageId="10" pageNumber="261">P. coprophila</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D7ECDB70BFE00422" bold="true" box="[422,487,583,609]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D7ECDB70BFE00422" box="[422,487,583,609]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the presence of the punctation on the head, coarser punctation of the pronotum, shorter elytra, and slightly narrower and longer parameres;
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DDBB8BD4C048E" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
from
|
||
<taxonomicName id="EEF54D18FFE36D52D74DDBB8BF5804EA" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[263,351,655,681]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="steeli" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D74DDBB8BF5804EA" box="[263,351,655,681]" italics="true" pageId="10" pageNumber="261">P. steeli</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D722DBB8BFAD04EA" bold="true" box="[360,426,655,681]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D722DBB8BFAD04EA" box="[360,426,655,681]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by the darker coloration, the absence of elevations between punctures on the elytra, the shorter parameres with significantly narrower apical portions.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DDBE0BC6E0556" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
From all these species,
|
||
<taxonomicName id="EEF54D18FFE36D52D780DBE0BC7B04B3" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[458,636,727,753]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. n.">
|
||
<emphasis id="1B81EA89FFE36D52D780DBE0BFDC04B3" box="[458,475,727,752]" italics="true" pageId="10" pageNumber="261">P</emphasis>
|
||
.
|
||
<emphasis id="1B81EA89FFE36D52D7A2DBE0BC7B04B3" box="[488,636,727,752]" italics="true" pageId="10" pageNumber="261">atramentaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D4C8DBE0BCC704B2" bold="true" box="[642,704,727,753]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D4C8DBE0BCC704B2" box="[642,704,727,753]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be distinguished by the larger body and details of the external and internal morphology of the aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61EF6510FFE36D52D68DDA29BF6C051E" pageId="10" pageNumber="261" type="distribution">
|
||
<paragraph id="294A369BFFE36D52D68DDA29BF6C051E" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
<emphasis id="1B81EA89FFE36D52D68DDA29BF66057B" bold="true" box="[199,353,798,824]" pageId="10" pageNumber="261">Distribution.</emphasis>
|
||
<taxonomicName id="EEF54D18FFE36D52D726DA28BCC1057B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[364,710,799,825]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="261" phylum="Arthropoda" rank="species" species="atramentaria" status="sp. nov.">
|
||
<emphasis id="1B81EA89FFE36D52D726DA28BCC1057B" box="[364,710,799,825]" italics="true" pageId="10" pageNumber="261">Paraphloeostiba atramentaria</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="1B81EA89FFE36D52D498DA28BD11057A" bold="true" box="[722,790,799,825]" pageId="10" pageNumber="261">
|
||
<taxonomicNameLabel id="00B257F2FFE36D52D498DA28BD11057A" box="[722,790,799,825]" pageId="10" pageNumber="261" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known only from the
|
||
<typeStatus id="F64E8839FFE36D52D20EDA28BA73057A" box="[1092,1140,799,825]" pageId="10" pageNumber="261">type</typeStatus>
|
||
locality in Western New
|
||
<collectingCountry id="51E2760BFFE36D52D6DDDA74BEEC051E" box="[151,235,835,861]" name="Guinea-Bissau" pageId="10" pageNumber="261">Guinea</collectingCountry>
|
||
,
|
||
<collectingCountry id="51E2760BFFE36D52D6BCDA74BF60051E" box="[246,359,835,861]" name="Indonesia" pageId="10" pageNumber="261">Indonesia</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="61EF6510FFE36D52D68DDA50BF3705AE" pageId="10" pageNumber="261" type="etymology">
|
||
<paragraph id="294A369BFFE36D52D68DDA50BEF905E6" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
<emphasis id="1B81EA89FFE36D52D68DDA50BF4905C2" bold="true" box="[199,334,871,897]" pageId="10" pageNumber="261">Etymology.</emphasis>
|
||
The specific epithet is the Latin
|
||
<emphasis id="1B81EA89FFE36D52D482DA50BD6005C3" box="[712,871,871,896]" italics="true" pageId="10" pageNumber="261">atramentarius</emphasis>
|
||
, -
|
||
<emphasis id="1B81EA89FFE36D52D535DA50BDCF05C3" box="[895,968,871,896]" italics="true" pageId="10" pageNumber="261">a, -um</emphasis>
|
||
(blackish). It alludes to the coloration of the body.
|
||
</paragraph>
|
||
<paragraph id="294A369BFFE36D52D68DDA99BF3705AE" blockId="10.[151,1437,151,1005]" pageId="10" pageNumber="261">
|
||
<emphasis id="1B81EA89FFE36D52D68DDA99BF4C058B" bold="true" box="[199,331,942,968]" pageId="10" pageNumber="261">Bionomics.</emphasis>
|
||
The
|
||
<typeStatus id="F64E8839FFE36D52D7CCDA98BFEF058A" box="[390,488,943,969]" pageId="10" pageNumber="261" type="holotype">holotype</typeStatus>
|
||
was collected at elevations from
|
||
<quantity id="EE0D9B7EFFE36D52D52DDA98BA1C058A" box="[871,1051,943,969]" metricMagnitude="3" metricUnit="m" metricValue="2.2" metricValueMax="2.4" metricValueMin="2.0" pageId="10" pageNumber="261" unit="m" value="2200.0" valueMax="2400.0" valueMin="2000.0">2000 to 2400 m</quantity>
|
||
a.s.l. The detailed bionomial data are unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |