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<document id="1FD7563DAE0345AC17A4D8CC54C2506A" ID-DOI="10.11646/zootaxa.5424.3.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="10821214" ID-ZooBank="DE26201A-252D-40D3-A5CF-FCCC6B3C2D01" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1710500677057" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2024" docId="A15C878DFFC16D71D6DDD9A0BF3700A2" docLanguage="en" docName="zootaxa.5424.3.1.pdf" docOrigin="Zootaxa 5424 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5424.3.1" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Paraphloeostiba riedeli Shavrin 2024, sp. n." docType="treatment" docVersion="2" lastPageNumber="292" masterDocId="5D65FFF5FFE96D58D64AD937BE070643" masterDocTitle="Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960" masterLastPageNumber="307" masterPageNumber="251" pageNumber="291" updateTime="1710501296679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="6B547A0217F52F66B230E053E3335DFC" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="EC679DC93A2F428A3643C8AE8F5180F9">Twenty-one new species of Omaliini from the Papuan Region (Coleoptera: Staphylinidae: Omaliinae), with diagnostic and faunistic notes on some species of the genus Paraphloeostiba Steel, 1960</mods:title>
</mods:titleInfo>
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<mods:namePart id="1B137CE4193C11DFBFFA8152826E3134">Shavrin, Alexey V.</mods:namePart>
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<mods:title id="212D658CFC9A0B7C4C6695A55103EF43">Zootaxa</mods:title>
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<mods:date id="14205D5CC85C830F742A0787FC2B12CB">2024</mods:date>
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<mods:number id="3AD77318E88E980F2B8136C12F42B508">2024-03-14</mods:number>
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<mods:number id="E426FA7802C25391F4FC092076660144">3</mods:number>
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<mods:url id="FA4E8097EF961DEBD780218F1134C259">http://dx.doi.org/10.11646/zootaxa.5424.3.1</mods:url>
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<mods:classification id="4A09BE8F2CCCBE0C0924196C3276C2A2">journal article</mods:classification>
<mods:identifier id="1FAB7AED8FA2F67B302FFB1C35EA7067" type="DOI">10.11646/zootaxa.5424.3.1</mods:identifier>
<mods:identifier id="3CE0203AC8AF29D4FA6669DD2B53A868" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="A28E1F392D1FA250D1F1D145D9441867" type="Zenodo-Dep">10821214</mods:identifier>
<mods:identifier id="BEE3427CBF3B4F8B900E51F2BC4A0CAD" type="ZooBank">DE26201A-252D-40D3-A5CF-FCCC6B3C2D01</mods:identifier>
</mods:mods>
<treatment id="A15C878DFFC16D71D6DDD9A0BF3700A2" ID-DOI="http://doi.org/10.5281/zenodo.10821305" ID-Zenodo-Dep="10821305" LSID="urn:lsid:plazi:treatment:A15C878DFFC16D71D6DDD9A0BF3700A2" httpUri="http://treatment.plazi.org/id/A15C878DFFC16D71D6DDD9A0BF3700A2" lastPageId="41" lastPageNumber="292" pageId="40" pageNumber="291">
<subSubSection id="61EF6510FFC16D70D6DDD9A0BFFD06F2" box="[151,506,151,178]" pageId="40" pageNumber="291" type="nomenclature">
<paragraph id="294A369BFFC16D70D6DDD9A0BFFD06F2" blockId="40.[151,506,151,214]" box="[151,506,151,178]" pageId="40" pageNumber="291">
<heading id="720281F7FFC16D70D6DDD9A0BFFD06F2" bold="true" box="[151,506,151,178]" fontSize="11" level="1" pageId="40" pageNumber="291" reason="1">
<emphasis id="1B81EA89FFC16D70D6DDD9A0BFFD06F2" bold="true" box="[151,506,151,178]" pageId="40" pageNumber="291">
<taxonomicName id="EEF54D18FFC16D70D6DDD9A0BFB606F1" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[151,433,151,178]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC16D70D6DDD9A0BFB606F1" bold="true" box="[151,433,151,178]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba riedeli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00B257F2FFC16D70D7F2D9A0BFFD06F2" box="[440,506,151,177]" pageId="40" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC16D70D6DDD98CBF6B0695" box="[151,364,187,214]" pageId="40" pageNumber="291" type="description">
<paragraph id="294A369BFFC16D70D6DDD98CBF6B0695" blockId="40.[151,506,151,214]" box="[151,364,187,214]" pageId="40" pageNumber="291">
(
<figureCitation id="B1CE2A1EFFC16D70D6D5D98CBEF10696" box="[159,246,187,213]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="40" pageNumber="291">Figs 78</figureCitation>
,
<figureCitation id="B1CE2A1EFFC16D70D748D98CBF630695" box="[258,356,187,214]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="40" pageNumber="291">119–125</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC16D70D6DDD834BEE707B6" pageId="40" pageNumber="291" type="materials_examined">
<paragraph id="294A369BFFC16D70D6DDD834BB480702" blockId="40.[151,1437,259,501]" pageId="40" pageNumber="291">
<emphasis id="1B81EA89FFC16D70D6DDD834BF4E075E" bold="true" box="[151,329,259,285]" pageId="40" pageNumber="291">Type material.</emphasis>
<typeStatus id="F64E8839FFC16D70D719D834BFBC075E" box="[339,443,259,285]" pageId="40" pageNumber="291" type="holotype">Holotype</typeStatus>
<specimenCount id="3FF3FD12FFC16D70D78CD833BFD9075D" box="[454,478,260,286]" count="1" pageId="40" pageNumber="291" type="male">♂</specimenCount>
: ‘
<collectingRegion id="EB31F879FFC16D70D7BBD834BC91075E" box="[497,662,259,285]" country="Indonesia" name="Papua Barat" pageId="40" pageNumber="291">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Bilogay,
<date id="5D4B105BFFC16D70D5FBD834BA28075D" box="[945,1071,259,286]" pageId="40" pageNumber="291" value="1995-12-29">29.12.1995</date>
|
<quantity id="EE0D9B7EFFC16D70D200D834BADE075E" box="[1098,1241,259,285]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="40" pageNumber="291" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
| leg. A. Riedel’ &lt;printed&gt;, ‘
<typeStatus id="F64E8839FFC16D70D752D810BFB60702" box="[280,433,295,321]" pageId="40" pageNumber="291" type="holotype">HOLOTYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC16D70D78FD810BC870702" authorityName="Steel" authorityYear="1960" box="[453,640,295,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC16D70D78FD810BC870702" box="[453,640,295,321]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC16D70D4D9D810BCDB0702" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[659,732,295,321]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC16D70D4D9D810BCDB0702" box="[659,732,295,321]" italics="true" pageId="40" pageNumber="291">riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC16D70D4A9D810BD240702" bold="true" box="[739,803,295,321]" pageId="40" pageNumber="291">
<taxonomicNameLabel id="00B257F2FFC16D70D4A9D810BD240702" box="[739,803,295,321]" pageId="40" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC16D70D5E6D810BDC50702" box="[940,962,295,321]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="40" pageNumber="291" type="Museum">V</collectionCode>
. des. 2024’ &lt;red, printed&gt; (
<collectionCode id="4FE4AE5EFFC16D70D2BFD810BB440702" box="[1269,1347,295,321]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="291">NMW</collectionCode>
).
</paragraph>
<paragraph id="294A369BFFC16D70D68DD87BBEE707B6" blockId="40.[151,1437,259,501]" pageId="40" pageNumber="291">
<typeStatus id="F64E8839FFC16D70D68DD87BBF3F0725" box="[199,312,332,358]" pageId="40" pageNumber="291" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="3FF3FD12FFC16D70D70FD87CBF930725" box="[325,404,331,358]" count="10" pageId="40" pageNumber="291" type="male">10 ♂♂</specimenCount>
(
<specimenCount id="3FF3FD12FFC16D70D7EFD87CBC480726" box="[421,591,331,357]" count="2" pageId="40" pageNumber="291" type="generic">two specimens</specimenCount>
dissected;
<specimenCount id="3FF3FD12FFC16D70D485D87BBD6C0726" box="[719,875,331,358]" count="1" pageId="40" pageNumber="291" type="generic">one specimen</specimenCount>
without apical part of the abdomen),
<specimenCount id="3FF3FD12FFC16D70D355D87CBB670725" box="[1311,1376,331,358]" count="5" pageId="40" pageNumber="291" type="female">5 ♀♀</specimenCount>
(
<specimenCount id="3FF3FD12FFC16D70D33BD87CBF0B07CA" count="2" pageId="40" pageNumber="291" type="generic">two specimens</specimenCount>
dissected): same data as the holotype (
<specimenCount id="3FF3FD12FFC16D70D486D858BD0807C9" box="[716,783,367,394]" count="2" pageId="40" pageNumber="291" type="male">2 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFC16D70D557D858BD4D07C9" box="[797,842,367,394]" count="1" pageId="40" pageNumber="291" type="female">1 ♀</specimenCount>
: cSh;
<specimenCount id="3FF3FD12FFC16D70D5D9D847BDD107C9" box="[915,982,368,394]" count="7" pageId="40" pageNumber="291" type="male">7 ♂♂</specimenCount>
,
<specimenCount id="3FF3FD12FFC16D70D5A9D858BA2107C9" box="[995,1062,367,394]" count="4" pageId="40" pageNumber="291" type="female">4 ♀♀</specimenCount>
:
<collectionCode id="4FE4AE5EFFC16D70D27ED858BA8607CA" box="[1076,1153,367,393]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="291">NMW</collectionCode>
);
<specimenCount id="3FF3FD12FFC16D70D2DFD858BAC607C9" box="[1173,1217,367,394]" count="1" pageId="40" pageNumber="291" type="male">1 ♂</specimenCount>
(left antennomeres 6–11 missing): ‘
<collectingRegion id="EB31F879FFC16D70D70DD8A4BFEF07EE" box="[327,488,403,429]" country="Indonesia" name="Papua Barat" pageId="40" pageNumber="291">IRIAN JAYA</collectingRegion>
: Paniai Prov. | Bilogay,
<quantity id="EE0D9B7EFFC16D70D4A4D8A4BD7A07EE" box="[750,893,403,429]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="40" pageNumber="291" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100-2200m</quantity>
|
<date id="5D4B105BFFC16D70D5C5D8A4BA0A07ED" box="[911,1037,403,430]" pageId="40" pageNumber="291" value="1995-12-29">29.12.1995</date>
| leg. A. Riedel’ &lt;printed&gt; (
<collectionCode id="4FE4AE5EFFC16D70D309D8A4BB9607EE" box="[1347,1425,403,429]" country="Austria" httpUri="http://grbio.org/cool/c7yt-hh77" name="Naturhistorisches Museum, Wien" pageId="40" pageNumber="291">NMW</collectionCode>
). All
<typeStatus id="F64E8839FFC16D70D689D88FBF290791" box="[195,302,440,466]" pageId="40" pageNumber="291" type="paratype">paratypes</typeStatus>
with additional red printed label: ‘
<typeStatus id="F64E8839FFC16D70D4FDD880BD400792" box="[695,839,439,465]" pageId="40" pageNumber="291" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="EEF54D18FFC16D70D514D880BA1E0792" authorityName="Steel" authorityYear="1960" box="[862,1049,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="genus">
<emphasis id="1B81EA89FFC16D70D514D880BA1E0792" box="[862,1049,439,465]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba</emphasis>
</taxonomicName>
|
<taxonomicName id="EEF54D18FFC16D70D265D880BA7F0792" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1071,1144,439,465]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC16D70D265D880BA7F0792" box="[1071,1144,439,465]" italics="true" pageId="40" pageNumber="291">riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC16D70D2CAD880BAC60792" bold="true" box="[1152,1217,439,465]" pageId="40" pageNumber="291">
<taxonomicNameLabel id="00B257F2FFC16D70D2CAD880BAC60792" box="[1152,1217,439,465]" pageId="40" pageNumber="291" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="4FE4AE5EFFC16D70D304D880BB630792" box="[1358,1380,439,465]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="40" pageNumber="291" type="Museum">V</collectionCode>
. des. 2024’.
</paragraph>
</subSubSection>
<caption id="7D8A6613FFC16D70D6DDDE63BDDE01F7" ID-DOI="http://doi.org/10.5281/zenodo.10821275" ID-Zenodo-Dep="10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="40" pageNumber="291" startId="40.[151,264,1876,1900]" targetBox="[159,1428,533,1844]" targetPageId="40" targetType="figure">
<paragraph id="294A369BFFC16D70D6DDDE63BDDE01F7" blockId="40.[151,1437,1876,1973]" pageId="40" pageNumber="291">
<emphasis id="1B81EA89FFC16D70D6DDDE63BF77012F" bold="true" box="[151,368,1876,1900]" pageId="40" pageNumber="291">FIGURES 119–125.</emphasis>
<taxonomicName id="EEF54D18FFC16D70D733DE63BC69012F" authorityName="Shavrin" authorityYear="2024" box="[377,622,1876,1900]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="40" pageNumber="291" phylum="Arthropoda" rank="species" species="riedeli">
<emphasis id="1B81EA89FFC16D70D733DE63BC69012F" box="[377,622,1876,1900]" italics="true" pageId="40" pageNumber="291">Paraphloeostiba riedeli</emphasis>
</taxonomicName>
(paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm.
</paragraph>
</caption>
<subSubSection id="61EF6510FFC06D71D68DD9A1BC60006E" pageId="41" pageNumber="292" type="description">
<paragraph id="294A369BFFC06D71D68DD9A1BEDE075E" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DD9A1BF5E06F3" bold="true" box="[199,345,150,176]" pageId="41" pageNumber="292">Description.</emphasis>
Measurements (n=17): HW: 0.42–0.56; HL: 0.25–0.26; OL: 0.14; TL: 0.02; AL (
<typeStatus id="F64E8839FFC06D71D2A2D9A0BB5606F2" box="[1256,1361,151,177]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
): 0.62; PL: 0.29–0.34; PWmax: 0.54–0.56; PWmin: 0.46–0.47; ESL: 0.55–0.60; EW: 0.70–0.76; MTbL (
<typeStatus id="F64E8839FFC06D71D2ADD98CBB570696" box="[1255,1360,187,213]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
): 0.30; MTrL (
<typeStatus id="F64E8839FFC06D71D6A1D9E8BF5306BA" box="[235,340,223,249]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
): 0.15 (MTrL 1–4: 0.05; MTrL 5: 0.10); AW: 0.61–0.79; AedL: 0.42; BL: 1.80–2.40 (
<typeStatus id="F64E8839FFC06D71D37BD9E8BB9E06BA" box="[1329,1433,223,249]" pageId="41" pageNumber="292" type="holotype">holotype</typeStatus>
: 1.87).
</paragraph>
<paragraph id="294A369BFFC06D71D68DD810BD720422" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
Habitus as in
<figureCitation id="B1CE2A1EFFC06D71D722D810BFBA0702" box="[360,445,295,321]" captionStart="FIGURES 75–80" captionStartId="28.[151,264,1598,1622]" captionTargetBox="[212,1375,188,1560]" captionTargetId="figure-22@28.[200,1387,181,1574]" captionTargetPageId="28" captionText="FIGURES 75–80. Habitus of Paraphloeostiba: 75–P. okapensis (paratype), 76–P. penelopeae (paratype), 77–P. pilosa (holotype), 78–P. riedeli (paratype), 79–P. rhopalocera, 80–P. rufula (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821253" httpUri="https://zenodo.org/record/10821253/files/figure.png" pageId="41" pageNumber="292">Fig. 78</figureCitation>
. Body yellowish brown to reddish-brown, with slightly paler elytra (several
<typeStatus id="F64E8839FFC06D71D37BD810BB9B0702" box="[1329,1436,295,321]" pageId="41" pageNumber="292" type="paratype">paratypes</typeStatus>
with paler lateral and basal portion of pronotum); antennomeres 6–11 brown; mouthparts, antennomeres 1–5 and legs yellow (some
<typeStatus id="F64E8839FFC06D71D7E9D858BC0907CA" box="[419,526,367,393]" pageId="41" pageNumber="292" type="paratype">paratypes</typeStatus>
with distinctly darker preapical maxillary palpomere). Pronotum with fine and sparse punctation, slightly larger in lateral and finer and sparser in middle portions (some
<typeStatus id="F64E8839FFC06D71D2C9D8A4BAE907EE" box="[1155,1262,403,429]" pageId="41" pageNumber="292" type="paratype">paratypes</typeStatus>
without visible punctation in middle); elytra with moderately dense punctation, distinctly larger and deeper than that on pronotum, finer and sparser in middle; abdominal tergites with distinct, moderately fine and dense punctation. Middle portion of head with dense and coarse isodiametric microsculpture; pronotum with dense microreticulation somewhat similar to that in middle portion of head, finer in middle; microsculpture of elytra about as that in pronotum, slightly finer in middle; abdominal tergites with dense transverse meshes.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDB5CBFB0057A" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">Head 1.6 times to twice as broad as long, with narrow and moderately deep linear anteocellar foveae; postocular carinae acute. Ocelli large, located at level of postocular carinae; distance between ocelli 1.6–1.8 times as long as distance between ocellus and posterior margin of eyes. Apical maxillary palpomere about 1.6 times as long as preapical segment. Antennomere 4 indistinctly longer than broad, twice shorter than 3, 5 slightly broader than 4, 6 slightly longer and about twice broader than 5, 7 slightly longer and broader than 5, 8–9 slightly shorter than 7, 10 indisitnctly longer than 9.</paragraph>
<paragraph id="294A369BFFC06D71D68DDA74BC7405E6" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">Pronotum 1.6–1.8 times as broad as long, 1.1–1.2 times as broad as head, widest in middle, slightly more narrowed posteriad than anteriad; middle portion with indistinct wide longitudinal impressions; laterobasal impressions moderately wide and shallow.</paragraph>
<paragraph id="294A369BFFC06D71D68DDA98BDCB058A" blockId="41.[151,1437,150,1761]" box="[199,972,943,969]" pageId="41" pageNumber="292">Elytra 1.2 times as broad as long, 1.7–1.8 times as long as pronotum.</paragraph>
<paragraph id="294A369BFFC06D71D68DDAE4BCFA05AE" blockId="41.[151,1437,150,1761]" box="[199,765,979,1005]" pageId="41" pageNumber="292">Abdomen slightly narrower or broader than elytra.</paragraph>
<paragraph id="294A369BFFC06D71D68DDAC0BD630286" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
Male. Posterior margin of abdominal tergite VIII truncate (
<figureCitation id="B1CE2A1EFFC06D71D506DAC0BDAB0252" box="[844,940,1015,1041]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 121</figureCitation>
). Posterior margin of abdominal sternite VIII concave (
<figureCitation id="B1CE2A1EFFC06D71D749DD2CBF610276" box="[259,358,1051,1077]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 122</figureCitation>
). Aedeagus with wide basal portion, gradually narrowed toward small rounded apex; parameres slightly not reaching apex of median lobe, slightly broadened in preapical and narrowed in apical portions, each with three apical and three preapical moderately short setae, inner margins of each paramere in middle with several moderately short setae; internal sac long and moderately narrow, with two elongate sclerotized structures in basal portion (
<figureCitation id="B1CE2A1EFFC06D71D6BDDD9CBF5E0286" box="[247,345,1195,1221]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 119</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="B1CE2A1EFFC06D71D54ADD9CBD670286" box="[768,864,1195,1221]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 120</figureCitation>
.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDDF8BD3B0372" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
Female. Posterior margin of abdominal tergite VIII rounded (
<figureCitation id="B1CE2A1EFFC06D71D53CDDF8BDDF02AA" box="[886,984,1231,1257]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 123</figureCitation>
). Posterior margin of abdominal sternite VIII truncate (
<figureCitation id="B1CE2A1EFFC06D71D770DDC4BF9A034E" box="[314,413,1267,1293]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 124</figureCitation>
). Accessory sclerite triangular, with wide basal portion, gradually narrowed apicad toward small rounded apex (
<figureCitation id="B1CE2A1EFFC06D71D7C8DC20BFE30372" box="[386,484,1302,1329]" captionStart="FIGURES 119–125" captionStartId="40.[151,264,1876,1900]" captionTargetBox="[159,1428,533,1844]" captionTargetId="figure-136@40.[151,1436,525,1852]" captionTargetPageId="40" captionText="FIGURES 119–125. Paraphloeostiba riedeli (paratypes): 119–aedeagus, parameral view (holotype), 120–aedeagus, lateral view (holotype), 121–male abdominal tergite VIII, 122–male abdominal sternite VIII, 123–female abdominal tergite VIII, 124– female abdominal sternite VIII, 125–female accessory sclerite. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.10821275" httpUri="https://zenodo.org/record/10821275/files/figure.png" pageId="41" pageNumber="292">Fig. 125</figureCitation>
). Spermatheca not recognized.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDC0DBC60006E" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDC0DBFA80316" bold="true" box="[199,431,1338,1365]" pageId="41" pageNumber="292">Comparative notes.</emphasis>
Based on the general structure of the aedeagus,
<taxonomicName id="EEF54D18FFC06D71D5F4DC0CBA260316" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[958,1057,1339,1365]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="riedeli" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D5F4DC0CBA260316" box="[958,1057,1339,1365]" italics="true" pageId="41" pageNumber="292">P. riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D26CDC0CBA630316" bold="true" box="[1062,1124,1339,1365]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D26CDC0CBA630316" box="[1062,1124,1339,1365]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEF54D18FFC06D71D2A5DC0CBB700317" authorityName="Shavrin" authorityYear="2023" box="[1263,1399,1339,1364]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFC06D71D2A5DC0CBB700317" box="[1263,1399,1339,1364]" italics="true" pageId="41" pageNumber="292">P. conjuncta</emphasis>
</taxonomicName>
,
<taxonomicName id="EEF54D18FFC06D71D3CEDC0CBF5C033A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D3CEDC0CBB920317" box="[1412,1429,1339,1364]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D6DDDC68BF5C033A" box="[151,347,1375,1401]" italics="true" pageId="41" pageNumber="292">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D728DC68BFA5033A" bold="true" box="[354,418,1375,1401]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D728DC68BFA5033A" box="[354,418,1375,1401]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, and
<taxonomicName id="EEF54D18FFC06D71D795DC68BC45033B" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[479,578,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D795DC68BFF7033B" box="[479,496,1375,1400]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D7B4DC68BC45033B" box="[510,578,1375,1400]" italics="true" pageId="41" pageNumber="292">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D403DC68BC8E033A" bold="true" box="[585,649,1375,1401]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D403DC68BC8E033A" box="[585,649,1375,1401]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
It can be distinguished from
<taxonomicName id="EEF54D18FFC06D71D59ADC68BA5D033B" authorityName="Shavrin" authorityYear="2023" box="[976,1114,1375,1400]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFC06D71D59ADC68BA5D033B" box="[976,1114,1375,1400]" italics="true" pageId="41" pageNumber="292">P. conjuncta</emphasis>
</taxonomicName>
and
<taxonomicName id="EEF54D18FFC06D71D2DBDC68BB73033A" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1169,1396,1375,1401]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="guadalcanalensis" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D2DBDC68BAA5033B" box="[1169,1186,1375,1400]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D2FADC68BB73033A" box="[1200,1396,1375,1401]" italics="true" pageId="41" pageNumber="292">guadalcanalensis</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D331DC68BEA903DE" bold="true" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D331DC68BEA903DE" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
by the less distinct longitudinal impressions in the middle of the pronotum. From
<taxonomicName id="EEF54D18FFC06D71D209DCB4BAA203DF" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[1091,1189,1411,1437]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="iriana" status="sp. n.">
<emphasis id="1B81EA89FFC06D71D209DCB4BA5303DF" box="[1091,1108,1411,1436]" italics="true" pageId="41" pageNumber="292">P</emphasis>
.
<emphasis id="1B81EA89FFC06D71D22BDCB4BAA203DF" box="[1121,1189,1411,1436]" italics="true" pageId="41" pageNumber="292">iriana</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D2E7DCB4BAEB03DE" bold="true" box="[1197,1260,1411,1437]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D2E7DCB4BAEB03DE" box="[1197,1260,1411,1437]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
it differs by the slightly shorter parameres, the absence of the punctation of the head, and the less distinct longitudinal impressions in the middle of the pronotum. Additionally, it differs from
<taxonomicName id="EEF54D18FFC06D71D545DCFCBD9F03A7" authorityName="Shavrin" authorityYear="2023" box="[783,920,1483,1508]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="conjuncta">
<emphasis id="1B81EA89FFC06D71D545DCFCBD9F03A7" box="[783,920,1483,1508]" italics="true" pageId="41" pageNumber="292">P. conjuncta</emphasis>
</taxonomicName>
by the pronotum with not concave laterobasal margins. From all these species it differs by the narrower median lobe, the different structure of the internal sac and the shape of the female accessory sclerite
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC06D71D68DDF01BB930012" box="[199,1428,1590,1617]" pageId="41" pageNumber="292" type="distribution">
<paragraph id="294A369BFFC06D71D68DDF01BB930012" blockId="41.[151,1437,150,1761]" box="[199,1428,1590,1617]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDF01BF660013" bold="true" box="[199,353,1590,1616]" pageId="41" pageNumber="292">Distribution.</emphasis>
<taxonomicName id="EEF54D18FFC06D71D72DDF00BC750012" authority="Shavrin, 2024" authorityName="Shavrin" authorityYear="2024" box="[359,626,1591,1617]" class="Insecta" family="Staphylinidae" genus="Paraphloeostiba" kingdom="Animalia" order="Coleoptera" pageId="41" pageNumber="292" phylum="Arthropoda" rank="species" species="riedeli" status="sp. nov.">
<emphasis id="1B81EA89FFC06D71D72DDF00BC750012" box="[359,626,1591,1617]" italics="true" pageId="41" pageNumber="292">Paraphloeostiba riedeli</emphasis>
</taxonomicName>
<emphasis id="1B81EA89FFC06D71D433DF00BCBE0012" bold="true" box="[633,697,1591,1617]" pageId="41" pageNumber="292">
<taxonomicNameLabel id="00B257F2FFC06D71D433DF00BCBE0012" box="[633,697,1591,1617]" pageId="41" pageNumber="292" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is known from two localities in Western New
<collectingCountry id="51E2760BFFC06D71D28BDF00BB120012" box="[1217,1301,1591,1617]" name="Guinea-Bissau" pageId="41" pageNumber="292">Guinea</collectingCountry>
,
<collectingCountry id="51E2760BFFC06D71D36ADF00BB960012" box="[1312,1425,1591,1617]" name="Indonesia" pageId="41" pageNumber="292">Indonesia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="61EF6510FFC06D71D68DDF6CBF3700A2" pageId="41" pageNumber="292" type="etymology">
<paragraph id="294A369BFFC06D71D68DDF6CBF5C00DA" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDF6CBF490036" bold="true" box="[199,334,1627,1653]" pageId="41" pageNumber="292">Etymology.</emphasis>
Patronymic, the species is named after coleopterologist Alexander Riedel (Karlsruhe), collector of the
<typeStatus id="F64E8839FFC06D71D68ADF48BEF700DA" box="[192,240,1663,1689]" pageId="41" pageNumber="292">type</typeStatus>
material.
</paragraph>
<paragraph id="294A369BFFC06D71D68DDF95BF3700A2" blockId="41.[151,1437,150,1761]" pageId="41" pageNumber="292">
<emphasis id="1B81EA89FFC06D71D68DDF95BF4C00FF" bold="true" box="[199,331,1698,1724]" pageId="41" pageNumber="292">Bionomics.</emphasis>
Specimens were collected at elevations from
<quantity id="EE0D9B7EFFC06D71D512DF94BA0A00FE" box="[856,1037,1699,1725]" metricMagnitude="3" metricUnit="m" metricValue="2.15" metricValueMax="2.2" metricValueMin="2.1" pageId="41" pageNumber="292" unit="m" value="2150.0" valueMax="2200.0" valueMin="2100.0">2100 to 2200 m</quantity>
a.s.l. The detailed bionomical data are unknown.
</paragraph>
</subSubSection>
</treatment>
</document>
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