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<document id="23E0C3756FBD01820C9DF8CE6713C9C2" ID-DOI="10.11646/zootaxa.4028.2.2" ID-GBIF-Dataset="07168623-b3d8-461a-8a1d-7a4751fbbc8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="233672" ID-ZooBank="2ED815F3-73F8-41EF-905E-0BB98444E7FA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461285306969" checkinUser="plazi" docAuthor="Câmara, Josenir T., Takiya, Daniela M., Plant, Adrian R. &amp; Rafael, José A." docDate="2015" docId="A1138055FFB2CB02A7C329C3FDCAFEC8" docLanguage="en" docName="zt04028p214.pdf" docOrigin="Zootaxa 4028 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hemerodromia mourai Câmara, Takiya, Plant &amp; Rafael, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="210" masterDocId="5D2AF82DFFBFCB0DA7542954FFD8FFD4" masterDocTitle="Neotropical Hemerodromia Meigen (Diptera: Empididae), a world of discovery II: New species from Atlantic forest, Brazil" masterLastPageNumber="214" masterPageNumber="197" pageNumber="210" updateTime="1698615035940" updateUser="plazi">
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<mods:title id="6BAC64CCE95881129342CC789724E6D5">Neotropical Hemerodromia Meigen (Diptera: Empididae), a world of discovery II: New species from Atlantic forest, Brazil</mods:title>
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<mods:namePart id="7D4ADB56ABB9A2CFC4153A29D59596C9">Câmara, Josenir T.</mods:namePart>
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<mods:namePart id="54EE67DCBA24EC8DE736393D9CCE2D0B">Takiya, Daniela M.</mods:namePart>
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<mods:namePart id="3EAC233EB2663B454A9F49AC18A1DE34">Plant, Adrian R.</mods:namePart>
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<mods:namePart id="0F4057C722766580FEF2556E7A85EC8D">Rafael, José A.</mods:namePart>
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<treatment id="A1138055FFB2CB02A7C329C3FDCAFEC8" ID-DOI="http://doi.org/10.5281/zenodo.6102304" ID-GBIF-Taxon="119641298" ID-Zenodo-Dep="6102304" LSID="urn:lsid:plazi:treatment:A1138055FFB2CB02A7C329C3FDCAFEC8" httpUri="http://treatment.plazi.org/id/A1138055FFB2CB02A7C329C3FDCAFEC8" lastPageId="15" lastPageNumber="210" pageId="13" pageNumber="210">
<subSubSection id="61A062C8FFB2CB00A7C329C3FEFFFF07" pageId="13" pageNumber="210" type="nomenclature">
<paragraph id="29053143FFB2CB00A7C329C3FDCAFF66" blockId="13.[151,530,151,211]" box="[151,530,151,178]" pageId="13" pageNumber="210">
<heading id="724D862FFFB2CB00A7C329C3FDCAFF66" bold="true" box="[151,530,151,178]" fontSize="11" level="1" pageId="13" pageNumber="210" reason="1">
<emphasis id="1BCEED51FFB2CB00A7C329C3FDCAFF66" bold="true" box="[151,530,151,178]" pageId="13" pageNumber="210">
<taxonomicName id="EEBA4AC0FFB2CB00A7C329C3FE76FF65" ID-CoL="9HXX9" box="[151,430,151,177]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="210" phylum="Arthropoda" rank="species" species="mourai" status="sp. nov.">
<emphasis id="1BCEED51FFB2CB00A7C329C3FE76FF65" bold="true" box="[151,430,151,177]" italics="true" pageId="13" pageNumber="210">Hemerodromia mourai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00FD502AFFB2CB00A6E129CCFDCAFF66" box="[437,530,152,178]" pageId="13" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="29053143FFB2CB00A7C329EEFEFFFF07" blockId="13.[151,530,151,211]" box="[151,295,186,211]" pageId="13" pageNumber="210">
(
<figureCitation id="B1812DC6FFB2CB00A7CB29EEFEC7FF07" box="[159,287,186,211]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Figs 4450</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="61A062C8FFB2CB00A7C32856FE2BFEB7" pageId="13" pageNumber="210" type="diagnosis">
<paragraph id="29053143FFB2CB00A7C32856FE2BFEB7" blockId="13.[151,1437,258,2022]" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A7C32856FED2FECF" bold="true" box="[151,266,258,283]" pageId="13" pageNumber="210">Diagnosis</emphasis>
. Yellow species. Fore femur with 2 basiventral denticles; cercus greatly enlarged, suboval in lateral view (
<figureCitation id="B1812DC6FFB2CB00A7CB2872FF2DFEEB" box="[159,245,294,319]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 46</figureCitation>
) and with row of spine-like setae ventrally (
<figureCitation id="B1812DC6FFB2CB00A4522872FC85FEEB" box="[774,861,294,319]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 48</figureCitation>
); subepandrial sclerite with two long, sclerotized posterior process (
<figureCitation id="B1812DC6FFB2CB00A630281EFE3FFEB7" box="[356,487,330,355]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Figs 46, 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="61A062C8FFB2CB00A793283AFD03FA3A" pageId="13" pageNumber="210" type="description">
<paragraph id="29053143FFB2CB00A793283AFDB7FA1E" blockId="13.[151,1437,258,2022]" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A793283AFE89FE53" bold="true" box="[199,337,366,391]" pageId="13" pageNumber="210">Description</emphasis>
.
<emphasis id="1BCEED51FFB2CB00A630283AFE79FE53" bold="true" box="[356,417,366,391]" pageId="13" pageNumber="210">Male</emphasis>
(
<figureCitation id="B1812DC6FFB2CB00A6E1283AFDD7FE53" box="[437,527,366,391]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 44</figureCitation>
).
<emphasis id="1BCEED51FFB2CB00A573283AFDBFFE53" bold="true" box="[551,615,366,391]" pageId="13" pageNumber="210">Head</emphasis>
. Dark brown to black, antenna and mouthparts yellow with all setae whitish. Ocellar triangle with 1 pair of proclinate setae; anterior ocellus larger. Eyes iridescent black, very narrowly separated on face bearing fine reclinate setulae. Frons with 12 frontal setulae; 1 pair of distinct vertical setae contiguous with uniseriate row of smaller postocular setae. Occiput bearing scattered fine hairs. Clypeus with rather dense, short downwardly directed pile. Proboscis slightly curved with yellow setae. Antenna with scape and pedicel bearing distinct, short dorsal setulae; postpedicel about 1.5X as long as wide, stylus ~ 1.2X as long as postpedicel.
<emphasis id="1BCEED51FFB2CB00A67C2B12FE59FD8B" bold="true" box="[296,385,582,607]" pageId="13" pageNumber="210">Thorax</emphasis>
(
<figureCitation id="B1812DC6FFB2CB00A6C72B12FE33FD8B" box="[403,491,582,607]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 44</figureCitation>
). Elongate, slightly arched ventrally; thoracic dorsum yellow except brownish on prothoracic collar, narrowly behind postpronotal lobe, on scutum posterolaterally above wing base, anatergite, suture between katepisternum and meron, katepisternum ventrally behind C1, laterally on scutellum and mediotergite; antepronotum with anterior margin almost straight and scutum with yellow setae, very small and fine except 1 notopleural and 2 pairs of scutellars.
<emphasis id="1BCEED51FFB2CB00A5E12B82FD35FD3B" bold="true" box="[693,749,726,751]" pageId="13" pageNumber="210">Legs</emphasis>
(
<figureCitation id="B1812DC6FFB2CB00A4542B82FC80FD3B" box="[768,856,726,751]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 44</figureCitation>
). Whitish yellow. C1 ~ 1.3X longer than distance between C1 and C2,
<date id="5D041783FFB2CB00A6D72BAEFE7DFCC7" box="[387,421,762,787]" pageId="13" pageNumber="210">6X</date>
as long as wide with some pale dorsoapical setae. F1 about 1.1X as long as C1, 3.03.5X as long as wide. Femoral formula 78/1617/2+2021/7; denticles confined to distal 0.9, becoming closer together distally, black, 2 rows linear, without distal discontinuity, diverging abruptly at extreme apex. T1 ~ 0.7X as long as F1, evenly curved, ventral face shallowly concave; about 2023 sharply pointed spinose setae ventrally; dorsal ciliation of decumbent short, pale setulae, denser distally; without apicoventral extension on T1 but with strong apical spine. Mid- and hind legs slender with fine setae; T3 with comb of short setae posteroapically.
<emphasis id="1BCEED51FFB2CB00A24A2AFAFA87FC13" bold="true" box="[1310,1375,942,967]" pageId="13" pageNumber="210">Wing</emphasis>
(
<figureCitation id="B1812DC6FFB2CB00A23A2AFAFF6FFC3F" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 45</figureCitation>
). Membrane slightly yellow, veins yellowish; R2+3 long and straight, not strongly curved towards C apically, joining C ~ 0.70.8 distance between end of R1 and R4+5; R4+5 fork acute (~ 60°), distal to position of M1+2 fork by about same length as R4; R5 and M1 convergent distally, diverging slightly at extreme apex; R5 ~
<date id="5D041783FFB2CB00A38D2D74FB23FBEC" box="[1241,1275,1056,1080]" pageId="13" pageNumber="210">2X</date>
as long as R4; cell bm+dm short, ending at level of R1, ~
<date id="5D041783FFB2CB00A53D2D13FD55FB8B" box="[617,653,1095,1119]" pageId="13" pageNumber="210">2X</date>
as long as cell br. Halter whitish yellow.
<emphasis id="1BCEED51FFB2CB00A30C2D12FB15FB8B" bold="true" box="[1112,1229,1094,1119]" pageId="13" pageNumber="210">Abdomen</emphasis>
(
<figureCitation id="B1812DC6FFB2CB00A3882D12FAE8FB8B" box="[1244,1328,1094,1119]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 44</figureCitation>
). Tergites 16 brown; tergites 78 and sternites yellow; tergite 8 shorter than sternite 8.
<emphasis id="1BCEED51FFB2CB00A3702D39FB7FFB52" bold="true" box="[1060,1191,1133,1158]" pageId="13" pageNumber="210">Terminalia</emphasis>
. Brownish (
<figureCitation id="B1812DC6FFB2CB00A26C2D39FA48FB52" box="[1336,1424,1133,1158]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 46</figureCitation>
). Cercus greatly enlarged, suboval with small median projection apically (
<figureCitation id="B1812DC6FFB2CB00A4A52DC5FB92FB7E" box="[1009,1098,1169,1194]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 46</figureCitation>
), left and right cerci closely approximated anterodorsally, distinctly setose on outer face and spine-like setae ventrally (
<figureCitation id="B1812DC6FFB2CB00A3E62DE1FAD1FB1A" box="[1202,1289,1205,1230]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 47</figureCitation>
). Epandrium narrow, rather inflated basally, narrower apically, with distinct, strong row of setae on outer face (
<figureCitation id="B1812DC6FFB2CB00A2612D8DFA57FB26" box="[1333,1423,1241,1266]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 46</figureCitation>
); articulated to the hypandrium by weak point anteriorly. Surstylus absent. Hypandrium subrectangular (
<figureCitation id="B1812DC6FFB2CB00A2452DA9FABCFAC2" box="[1297,1380,1277,1302]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 50</figureCitation>
), not markedly inflated, with strong setae; gonocoxal apodeme projecting as narrow process just beyond anterior margin of hypandrium (
<figureCitation id="B1812DC6FFB2CB00A61A2C11FE7CFA8A" box="[334,420,1349,1374]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 46</figureCitation>
). Subepandrial sclerite with 2 lateral sclerotized, very long finger-like posterior processes with setae apically (
<figureCitation id="B1812DC6FFB2CB00A6292C3DFE0BFA56" box="[381,467,1385,1410]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 48</figureCitation>
). Phallus strongly sclerotized basally, extending well beyond hypandrium apex, with apical membranous area covered with spicules (
<figureCitation id="B1812DC6FFB2CB00A5E02CD9FCD1FA72" box="[692,777,1421,1446]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 49</figureCitation>
). Ejaculatory apodeme absent.
<emphasis id="1BCEED51FFB2CB00A33D2CD9FAF7FA72" bold="true" box="[1129,1327,1421,1446]" pageId="13" pageNumber="210">Specimen length</emphasis>
:
<quantity id="EE429CA6FFB2CB00A26A2CDAFA40FA72" box="[1342,1432,1422,1446]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="13" pageNumber="210" unit="mm" value="2.9">2.9 mm</quantity>
;
<emphasis id="1BCEED51FFB2CB00A7C32CE5FEFAFA1E" bold="true" box="[151,290,1457,1482]" pageId="13" pageNumber="210">wing length</emphasis>
:
<quantity id="EE429CA6FFB2CB00A6652CE5FE52FA1D" box="[305,394,1457,1482]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="13" pageNumber="210" unit="mm" value="2.5">2.5 mm</quantity>
.
<emphasis id="1BCEED51FFB2CB00A6C02CE5FE33FA1E" bold="true" box="[404,491,1457,1482]" pageId="13" pageNumber="210">Female</emphasis>
. Unknown.
</paragraph>
<paragraph id="29053143FFB2CB00A7932C81FD03FA3A" blockId="13.[151,1437,258,2022]" box="[199,731,1493,1518]" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A7932C81FE3FFA3A" bold="true" box="[199,487,1493,1518]" pageId="13" pageNumber="210">Geographic distribution</emphasis>
.
<collectingCountry id="51AD71D3FFB2CB00A6A12C81FDE1FA3A" box="[501,569,1493,1518]" name="Brazil" pageId="13" pageNumber="210">Brazil</collectingCountry>
(Ceará State).
</paragraph>
</subSubSection>
<subSubSection id="61A062C8FFB2CB00A7932CADFD7CF976" pageId="13" pageNumber="210" type="materials_examined">
<paragraph id="29053143FFB2CB00A7932CADFCDCF9AA" blockId="13.[151,1437,258,2022]" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A7932CADFEA8F9C6" bold="true" box="[199,368,1529,1554]" pageId="13" pageNumber="210">
<typeStatus id="F6018FE1FFB2CB00A7932CADFEDAF9C6" box="[199,258,1529,1554]" pageId="13" pageNumber="210">Type</typeStatus>
material
</emphasis>
.
<emphasis id="1BCEED51FFB2CB00A6D52CADFDF8F9C6" bold="true" box="[385,544,1529,1554]" pageId="13" pageNumber="210">
<typeStatus id="F6018FE1FFB2CB00A6D52CADFDF8F9C6" box="[385,544,1529,1554]" pageId="13" pageNumber="210" type="holotype">HOLOTYPE</typeStatus>
</emphasis>
♂, “
<collectingCountry id="51AD71D3FFB2CB00A50F2CADFD7CF9C6" box="[603,676,1529,1554]" name="Brazil" pageId="13" pageNumber="210">Brazil</collectingCountry>
, CE[Ceará], Ubajara, P.[arque] N.[acional] Ubajara, Cachoeira do Cafundó.
<geoCoordinate id="4C8E5784FFB2CB00A6442F49FE5EF9E2" box="[272,390,1565,1590]" direction="south" orientation="latitude" pageId="13" pageNumber="210" precision="15" value="-3.8369443">03º50'13''S</geoCoordinate>
<geoCoordinate id="4C8E5784FFB2CB00A6C22F49FDC7F9E2" box="[406,543,1565,1590]" direction="west" orientation="longitude" pageId="13" pageNumber="210" precision="15" value="-40.909725">40º54'35''W</geoCoordinate>
, Malaise,
<date id="5D041783FFB2CB00A5F22F49FC8CF9E2" box="[678,852,1565,1590]" pageId="13" pageNumber="210" value="2012-07-20" valueMax="2012-07-22" valueMin="2012-07-20">2022.vii.2012</date>
. J.A.Rafael &amp; F.Limeira-de-Oliveira.” (
<collectionCode id="4FABA986FFB2CB00A26D2F4AFA48F9E2" box="[1337,1424,1566,1590]" pageId="13" pageNumber="210">CZMA</collectionCode>
).
<typeStatus id="F6018FE1FFB2CB00A7C32F15FED7F98E" box="[151,271,1601,1626]" pageId="13" pageNumber="210" type="paratype">
<emphasis id="1BCEED51FFB2CB00A7C32F15FED7F98E" bold="true" box="[151,271,1601,1626]" pageId="13" pageNumber="210">Paratypes</emphasis>
</typeStatus>
: Same data as
<typeStatus id="F6018FE1FFB2CB00A6E12F15FDCFF98E" box="[437,535,1601,1626]" pageId="13" pageNumber="210" type="holotype">holotype</typeStatus>
(
<specimenCount id="3FBCFACAFFB2CB00A5722F15FD8AF98D" box="[550,594,1601,1626]" pageId="13" pageNumber="210" type="generic">5 ♂</specimenCount>
,
<collectionCode id="4FABA986FFB2CB00A5092F16FD69F98E" box="[605,689,1602,1626]" pageId="13" pageNumber="210">CZMA</collectionCode>
,
<specimenCount id="3FBCFACAFFB2CB00A5E82F16FD3FF98D" box="[700,743,1602,1626]" pageId="13" pageNumber="210" type="generic">2 ♂</specimenCount>
,
<collectionCode id="4FABA986FFB2CB00A5A72F16FCEFF98E" box="[755,823,1602,1626]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="13" pageNumber="210">INPA</collectionCode>
).
<emphasis id="1BCEED51FFB2CB00A41C2F16FC58F98E" box="[840,896,1602,1626]" italics="true" pageId="13" pageNumber="210">Idem</emphasis>
,
<date id="5D041783FFB2CB00A4D92F15FBEBF98E" box="[909,1075,1601,1626]" pageId="13" pageNumber="210" value="2012-11-13" valueMax="2012-11-17" valueMin="2012-11-13">1317.xi.2012</date>
. F. Limeira-de-Oliveira, D.W.A. Marques, cols. (
<specimenCount id="3FBCFACAFFB2CB00A61F2F32FEAFF9AA" box="[331,375,1638,1662]" pageId="13" pageNumber="210" type="generic">3 ♂</specimenCount>
,
<collectionCode id="4FABA986FFB2CB00A6D72F32FE38F9AA" box="[387,480,1638,1662]" pageId="13" pageNumber="210">MNWC</collectionCode>
,
<specimenCount id="3FBCFACAFFB2CB00A6BF2F32FDCFF9AA" box="[491,535,1638,1662]" pageId="13" pageNumber="210" type="generic">8 ♂</specimenCount>
,
<collectionCode id="4FABA986FFB2CB00A5772F32FDBDF9AA" box="[547,613,1638,1662]" httpUri="http://grbio.org/cool/rm88-499z" name="Instituto Nacional de Pesquisas da Amazonia" pageId="13" pageNumber="210">INPA</collectionCode>
,
<specimenCount id="3FBCFACAFFB2CB00A5242F31FD44F9AA" box="[624,668,1637,1662]" pageId="13" pageNumber="210" type="generic">5 ♂</specimenCount>
,
<collectionCode id="4FABA986FFB2CB00A5FC2F32FD20F9AA" box="[680,760,1638,1662]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="13" pageNumber="210">MNRJ</collectionCode>
).
</paragraph>
<paragraph id="29053143FFB2CB00A7932FDDFD7CF976" blockId="13.[151,1437,258,2022]" box="[199,676,1673,1698]" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A7932FDDFE73F976" bold="true" box="[199,427,1673,1698]" pageId="13" pageNumber="210">
<typeStatus id="F6018FE1FFB2CB00A7932FDDFEECF976" box="[199,308,1673,1698]" pageId="13" pageNumber="210" type="holotype">Holotype</typeStatus>
condition
</emphasis>
. Good. Not dissected.
</paragraph>
</subSubSection>
<subSubSection id="61A062C8FFB2CB00A7932FF9FB34F8DA" pageId="13" pageNumber="210" type="etymology">
<paragraph id="29053143FFB2CB00A7932FF9FDEFF93E" blockId="13.[151,1437,258,2022]" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A7932FF9FE9EF912" bold="true" box="[199,326,1709,1734]" pageId="13" pageNumber="210">Etymology</emphasis>
. The specific epithet is in tribute to Antonio Moura, the senior author's husband, partner and friend, for his contribution during this work.
</paragraph>
<paragraph id="29053143FFB2CB00A7932FA1FB34F8DA" blockId="13.[151,1437,258,2022]" box="[199,1260,1781,1806]" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A7932FA1FE98F8DA" bold="true" box="[199,320,1781,1806]" pageId="13" pageNumber="210">Variations</emphasis>
. Body length varying from
<quantity id="EE429CA6FFB2CB00A52F2FA1FCDAF8DA" box="[635,770,1781,1806]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="3.6" metricValueMin="2.4" pageId="13" pageNumber="210" unit="mm" value="3.0" valueMax="3.6" valueMin="2.4">2.43.6 mm</quantity>
and wing length varying from
<quantity id="EE429CA6FFB2CB00A30A2FA2FB30F8DA" box="[1118,1256,1782,1806]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" metricValueMax="2.9" metricValueMin="1.3" pageId="13" pageNumber="210" unit="mm" value="2.1" valueMax="2.9" valueMin="1.3">1.32.9 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="61A062C8FFB2CB02A7932E4DFDCAFEC8" lastPageId="15" lastPageNumber="212" pageId="13" pageNumber="210" type="discussion">
<paragraph id="29053143FFB2CB02A7932E4DFDCAFEC8" blockId="13.[151,1437,258,2022]" lastBlockId="15.[151,1436,151,285]" lastPageId="15" lastPageNumber="212" pageId="13" pageNumber="210">
<emphasis id="1BCEED51FFB2CB00A7932E4DFEECF8E6" bold="true" box="[199,308,1817,1842]" pageId="13" pageNumber="210">Remarks</emphasis>
.
<taxonomicName id="EEBA4AC0FFB2CB00A6172E4EFD9DF8E5" box="[323,581,1818,1842]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="210" phylum="Arthropoda" rank="species" species="mourai" status="sp. nov.">
<emphasis id="1BCEED51FFB2CB00A6172E4EFD9DF8E5" box="[323,581,1818,1842]" italics="true" pageId="13" pageNumber="210">Hemerodromia mourai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00FD502AFFB2CB00A5182E4DFD7FF8E6" box="[588,679,1817,1842]" pageId="13" pageNumber="210" rank="species">
<emphasis id="1BCEED51FFB2CB00A5182E4DFD7FF8E6" bold="true" box="[588,679,1817,1842]" pageId="13" pageNumber="210">sp. nov.</emphasis>
</taxonomicNameLabel>
appears to belong to the informal
<emphasis id="1BCEED51FFB2CB00A37A2E4EFB71F8E6" box="[1070,1193,1818,1842]" italics="true" pageId="13" pageNumber="210">melangyna</emphasis>
species
<emphasis id="1BCEED51FFB2CB00A2562E4EFAD3F8E5" box="[1282,1291,1818,1841]" italics="true" pageId="13" pageNumber="210">-</emphasis>
group (
<emphasis id="1BCEED51FFB2CB00A20A2E4EFA44F8E5" box="[1374,1436,1818,1841]" italics="true" pageId="13" pageNumber="210">sensu</emphasis>
<bibRefCitation id="4D2B4CB2FFB2CB00A7C32E69FE1AF882" author="Vaillant" box="[151,450,1853,1878]" pageId="13" pageNumber="210" refString="Vaillant, F. &amp; Gagneur, J. (1998) The Diptera Empididae Hemerodromiinae from Western Algeria and Middle Atlas of Morocco. Annales de la Societe Entomologique de France, 34, 365 - 84." type="journal article" year="1998">Vaillant &amp; Gagneur, 1998</bibRefCitation>
: 376) in having the cercus greatly enlarged, fore tibia without an apical extension but with a long, slender and pointed apical spine.
<taxonomicName id="EEBA4AC0FFB2CB00A5EB2E36FC1EF8AD" box="[703,966,1890,1914]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="210" phylum="Arthropoda" rank="species" species="mourai" status="sp. nov.">
<emphasis id="1BCEED51FFB2CB00A5EB2E36FC1EF8AD" box="[703,966,1890,1914]" italics="true" pageId="13" pageNumber="210">Hemerodromia mourai</emphasis>
</taxonomicName>
<emphasis id="1BCEED51FFB2CB00A4852E35FBE8F8AE" bold="true" box="[977,1072,1889,1914]" pageId="13" pageNumber="210">
<taxonomicNameLabel id="00FD502AFFB2CB00A4852E35FBE8F8AE" box="[977,1072,1889,1914]" pageId="13" pageNumber="210" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is similar to
<taxonomicName id="EEBA4AC0FFB2CB00A3812E36FE31F84A" authority="Camara, Plant &amp; Rafael, 2014" authorityName="Camara, Plant &amp; Rafael" authorityYear="2014" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="210" phylum="Arthropoda" rank="species" species="cercusdilatata">
<emphasis id="1BCEED51FFB2CB00A3812E36FA44F8AE" box="[1237,1436,1890,1914]" italics="true" pageId="13" pageNumber="210">H. cercusdilatata</emphasis>
<bibRefCitation id="4D2B4CB2FFB2CB00A7C32ED2FE31F84A" author="Camara" box="[151,489,1925,1950]" pageId="13" pageNumber="210" refString="Camara, J. T., Plant, A. R. &amp; Rafael, J. A. (2014) Neotropical Hemerodromia Meigen (Diptera: Empididae), a world of discovery I: New generic record and new species from Brazilian Amazon basin. Zootaxa, 3893 (2), 209 - 231. http: // dx. doi. org / 10.11646 / zootaxa. 3893.2.3" type="journal article" year="2014">
Câmara, Plant &amp; Rafael,
<quantity id="EE429CA6FFB2CB00A6E52ED2FE31F84A" box="[433,489,1926,1950]" metricMagnitude="1" metricUnit="m" metricValue="5.11556" pageId="13" pageNumber="210" unit="in" value="2014.0">2014</quantity>
</bibRefCitation>
</taxonomicName>
in having tergites 78 and sternites yellow, fore tibia with 2 rows of pointed spinose setae ventrally, subepandrial sclerite with 2 finger-like posterior processes and similar shaped epandrium. In
<taxonomicName id="EEBA4AC0FFB2CB00A2D52EFEFF30F832" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="210" phylum="Arthropoda" rank="species" species="mourai" status="sp. nov.">
<emphasis id="1BCEED51FFB2CB00A2D52EFEFF30F832" italics="true" pageId="13" pageNumber="210">H. mourai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="00FD502AFFB2CB00A7BA2E99FE9FF832" box="[238,327,1997,2022]" pageId="13" pageNumber="210" rank="species">
<emphasis id="1BCEED51FFB2CB00A7BA2E99FE9FF832" bold="true" box="[238,327,1997,2022]" pageId="13" pageNumber="210">sp. nov.</emphasis>
</taxonomicNameLabel>
fore femur has 2 basal denticles; cercus suboval (
<figureCitation id="B1812DC6FFB2CB00A43B2E99FC19F832" box="[879,961,1997,2022]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="13" pageNumber="210">Fig. 46</figureCitation>
) and with row of spine-like setae ventrally (
<figureCitation id="B1812DC6FFB0CB02A7CB29C3FF2CFF64" box="[159,244,151,176]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="15" pageNumber="212">Fig. 48</figureCitation>
); and epandrium with a long, stronger seta at apex (
<figureCitation id="B1812DC6FFB0CB02A46329C3FC53FF64" box="[823,907,151,176]" captionStart="FIGURES 44 50" captionStartId="14.[151,264,1928,1951]" captionTargetBox="[210,1359,193,1899]" captionTargetId="figure@14.[210,1376,193,1908]" captionTargetPageId="14" captionText="FIGURES 44 50. Hemerodromia mourai sp. nov., 44 45 holotype ♂. 46 50, paratype ♂. 44, habitus, lateral view; 45, wing; 46, terminalia, lateral view; 47, cercus, ventral view; 48, epandrium and subepandrial sclerite, ventral view; 49, phallus and hypandrium, lateral view; 50, terminalia, ventral view." httpUri="https://zenodo.org/record/233679/files/figure.png" pageId="15" pageNumber="212">Fig. 46</figureCitation>
). In
<taxonomicName id="EEBA4AC0FFB0CB02A4E929CDFBA6FF64" box="[957,1150,152,176]" class="Insecta" family="Empididae" genus="Hemerodromia" kingdom="Animalia" order="Diptera" pageId="15" pageNumber="212" phylum="Arthropoda" rank="species" species="cercusdilatata">
<emphasis id="1BCEED51FFB0CB02A4E929CDFBA6FF64" box="[957,1150,152,176]" italics="true" pageId="15" pageNumber="212">H. cercusdilatata</emphasis>
</taxonomicName>
the fore femur is without basal denticles; the cercus has with a bluntly pointed apical process incurved with serrate posteroapical margin (
<bibRefCitation id="4D2B4CB2FFB0CB02A7CA29B4FEAFFF2C" author="Camara" box="[158,375,224,248]" pageId="15" pageNumber="212" refString="Camara, J. T., Plant, A. R. &amp; Rafael, J. A. (2014) Neotropical Hemerodromia Meigen (Diptera: Empididae), a world of discovery I: New generic record and new species from Brazilian Amazon basin. Zootaxa, 3893 (2), 209 - 231. http: // dx. doi. org / 10.11646 / zootaxa. 3893.2.3" type="journal article" year="2014">
Câmara
<emphasis id="1BCEED51FFB0CB02A7AA29B5FEEEFF2C" box="[254,310,224,248]" italics="true" pageId="15" pageNumber="212">et al.</emphasis>
2014
</bibRefCitation>
, fig. 20), and without spine-like setae ventrally; and epandrium with a spine-like setae on inner face (
<bibRefCitation id="4D2B4CB2FFB0CB02A7802850FE76FEC8" author="Camara" box="[212,430,260,284]" pageId="15" pageNumber="212" refString="Camara, J. T., Plant, A. R. &amp; Rafael, J. A. (2014) Neotropical Hemerodromia Meigen (Diptera: Empididae), a world of discovery I: New generic record and new species from Brazilian Amazon basin. Zootaxa, 3893 (2), 209 - 231. http: // dx. doi. org / 10.11646 / zootaxa. 3893.2.3" type="journal article" year="2014">
Câmara
<emphasis id="1BCEED51FFB0CB02A6602851FEB4FEC8" box="[308,364,260,284]" italics="true" pageId="15" pageNumber="212">et al.</emphasis>
2014
</bibRefCitation>
, fig. 22).
</paragraph>
</subSubSection>
</treatment>
</document>