484 lines
57 KiB
XML
484 lines
57 KiB
XML
<document id="5E5AC83488155209A72B8909B1DD01EB" ID-DOI="10.11646/zootaxa.5020.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222931" ID-ZooBank="882BBB9D-6E5B-4CE5-99DF-E91AC7971EB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629386356657" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2021" docId="785C87DD333F002AC8F7FEF7CC5F460A" docLanguage="en" docName="zootaxa.5020.1.2.pdf" docOrigin="Zootaxa 5020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus fuscoaeneus Dejean 1829" docType="treatment" docVersion="3" lastPageNumber="45" masterDocId="8465FFA533330024C860FF99CD16402C" masterDocTitle="On some Afrotropical Harpalus, with description of two new species, and remarks on Hypharpax australis (Coleoptera: Carabidae): misidentification, mislabeling and introduction to the Australian region" masterLastPageNumber="56" masterPageNumber="31" pageNumber="43" updateTime="1699070990175" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="7B5B486FBD3AF0C1F4C826F0764D7F41">On some Afrotropical Harpalus, with description of two new species, and remarks on Hypharpax australis (Coleoptera: Carabidae): misidentification, mislabeling and introduction to the Australian region</mods:title>
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<mods:namePart id="1AC0D605EC6CEB2B2800EC64D0AC9D30">Kataev, Boris M.</mods:namePart>
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<mods:title id="288A1A804D2E8E963B3CE5E6D0347705">Zootaxa</mods:title>
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<mods:part id="A89FC4A6A2C0017F1F58DE8F85EE04FB">
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<mods:date id="FF044734C8D64069967D36FBCB1437BB">2021</mods:date>
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<mods:number id="9F38C577BEA59A990CB203ECF69938FF">2021-08-11</mods:number>
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<mods:number id="3B4416C5803AD6DECAF52CBECF57DFAF">5020</mods:number>
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<mods:start id="76A48E792F25CE71668B18F3773BB559">31</mods:start>
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<mods:identifier id="96133E98C1DC6066C2C2E58B6D9310B8" type="DOI">10.11646/zootaxa.5020.1.2</mods:identifier>
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<mods:identifier id="854570C12B90314BF8D9C921ABF2405E" type="Zenodo-Dep">5222931</mods:identifier>
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<treatment id="785C87DD333F002AC8F7FEF7CC5F460A" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A" httpUri="http://treatment.plazi.org/id/785C87DD333F002AC8F7FEF7CC5F460A" lastPageId="14" lastPageNumber="45" pageId="12" pageNumber="43">
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<subSubSection id="B8EF6540333F0028C8F7FEF7CF4641A4" box="[151,592,366,393]" pageId="12" pageNumber="43" type="nomenclature">
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<paragraph id="F04A36CB333F0028C8F7FEF7CF4641A4" blockId="12.[151,592,366,430]" box="[151,592,366,393]" pageId="12" pageNumber="43">
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<heading id="AB0281A7333F0028C8F7FEF7CF4641A4" bold="true" box="[151,592,366,393]" fontSize="11" level="1" pageId="12" pageNumber="43" reason="1">
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<taxonomicName id="37F54D48333F0028C8F7FEF7CF4641A4" ID-CoL="3JSL4" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" box="[151,592,366,393]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">
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<emphasis id="C281EAD9333F0028C8F7FEF7CF4641A4" bold="true" box="[151,592,366,393]" pageId="12" pageNumber="43">
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<emphasis id="C281EAD9333F0028C8F7FEF7CCB141A4" bold="true" box="[151,423,366,393]" italics="true" pageId="12" pageNumber="43">Harpalus fuscoaeneus</emphasis>
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<bibRefCitation id="94644B3A333F0028C9CEFEF7CF4641A4" author="Dejean, P. F. M. A." box="[430,592,366,392]" pageId="12" pageNumber="43" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="B8EF6540333F0028C8F7FE0ACC6D4182" box="[151,379,403,430]" pageId="12" pageNumber="43" type="description">
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<paragraph id="F04A36CB333F0028C8F7FE0ACC6D4182" blockId="12.[151,592,366,430]" box="[151,379,403,430]" pageId="12" pageNumber="43">
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(
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<figureCitation id="68CE2A4E333F0028C8FFFE0ACC364181" box="[159,288,403,429]" captionStart="FIGURES 29–32" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[180,1402,189,1896]" captionTargetId="figure-16@13.[172,1415,181,1901]" captionTargetPageId="13" captionText="FIGURES 29–32. Harpalus spp., habitus. 29–31, H. fuscoaeneus Dejean, 1929 (29, S Africa; 30, lectotype; 31, holotype of H. nanus Dejean, 1829); 32, H. australasiae Dejean, 1829 (holotype). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222947" httpUri="https://zenodo.org/record/5222947/files/figure.png" pageId="12" pageNumber="43">Figs 29–31</figureCitation>
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,
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<figureCitation id="68CE2A4E333F0028C94CFE0ACC624182" box="[300,372,403,430]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1045,1069]" captionTargetBox="[158,1426,188,1013]" captionTargetId="figure-23@14.[151,1436,181,1021]" captionTargetPageId="14" captionText="FIGURES 33–36. Harpalus fuscoaeneus Dejean, 1929 (Cape of Good Hope). 33, gonocoxite; 34, laterotergite, gonosubcoxite, and gonocoxite; 35, 36, median lobe of aedeagus. Lateral (33, 35), ventral (34) and dorsal (36) views. Scale bar = 0. 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222949" httpUri="https://zenodo.org/record/5222949/files/figure.png" pageId="12" pageNumber="43">33–36</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B8EF6540333F0028C8F7FE4ECF554201" pageId="12" pageNumber="43" type="reference_group">
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<paragraph id="F04A36CB333F0028C8F7FE4ECF5741C3" blockId="12.[151,579,471,557]" box="[151,577,471,495]" pageId="12" pageNumber="43">
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<treatmentCitationGroup id="D0E511E5333F0028C8F7FE4ECF5741C3" box="[151,577,471,495]" pageId="12" pageNumber="43">
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<treatmentCitation id="715410DA333F0028C8F7FE4ECF2B41C3" author="Dejean, P. F. M. A." box="[151,573,471,495]" page="392" pageId="12" pageNumber="43" year="1829">
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<taxonomicName id="37F54D48333F0028C8F7FE4ECF2B41C3" ID-CoL="3JSL4" authority="Dejean, 1829: 392" authorityName="Dejean" authorityPageNumber="392" authorityYear="1829" box="[151,573,471,495]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">
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<emphasis id="C281EAD9333F0028C8F7FE4ECC6F41C3" box="[151,377,471,495]" italics="true" pageId="12" pageNumber="43">Harpalus fuscoaeneus</emphasis>
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<bibRefCitation id="94644B3A333F0028C91FFE4ECF2B41C3" author="Dejean, P. F. M. A." box="[383,573,471,495]" pageId="12" pageNumber="43" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829: 392</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="F04A36CB333F0028C8F7FE6FCF144222" blockId="12.[151,579,471,557]" box="[151,514,502,526]" pageId="12" pageNumber="43">
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<treatmentCitationGroup id="D0E511E5333F0028C8F7FE6FCF144222" box="[151,514,502,526]" pageId="12" pageNumber="43">
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<treatmentCitation id="715410DA333F0028C8F7FE6FCCE84222" author="Dejean, P. F. M. A." box="[151,510,502,526]" page="395" pageId="12" pageNumber="43" year="1829">
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<taxonomicName id="37F54D48333F0028C8F7FE6FCCE84222" ID-CoL="3JTMD" authority="Dejean, 1829: 395" authorityName="Dejean" authorityPageNumber="395" authorityYear="1829" box="[151,510,502,526]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="nanus">
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<emphasis id="C281EAD9333F0028C8F7FE6FCC2C4222" box="[151,314,502,526]" italics="true" pageId="12" pageNumber="43">Harpalus nanus</emphasis>
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<bibRefCitation id="94644B3A333F0028C920FE6FCCE84222" author="Dejean, P. F. M. A." box="[320,510,502,526]" pageId="12" pageNumber="43" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829: 395</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
<paragraph id="F04A36CB333F0028C8F7FD8DCF554201" blockId="12.[151,579,471,557]" box="[151,579,532,557]" pageId="12" pageNumber="43">
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<treatmentCitationGroup id="D0E511E5333F0028C8F7FD8DCF554201" box="[151,579,532,557]" pageId="12" pageNumber="43">
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<treatmentCitation id="715410DA333F0028C8F7FD8DCF294201" author="Peringuey, L." box="[151,575,532,557]" page="443" pageId="12" pageNumber="43" year="1896">
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||
<taxonomicName id="37F54D48333F0028C8F7FD8DCF294201" ID-CoL="3JTT2" authority="Peringuey, 1896: 443" authorityName="Peringuey" authorityPageNumber="443" authorityYear="1896" box="[151,575,532,557]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="precarius">
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<emphasis id="C281EAD9333F0028C8F7FD8DCC4B4200" box="[151,349,532,556]" italics="true" pageId="12" pageNumber="43">Harpalus precarius</emphasis>
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||
<bibRefCitation id="94644B3A333F0028C904FD8DCF294201" author="Peringuey, L." box="[356,575,532,557]" pageId="12" pageNumber="43" pagination="99 - 623" refId="ref16860" refString="Peringuey, L. (1896) Descriptive catalogue of the Coleoptera of South Africa. Part II. Cicindelidae supplement. Carabidae. The Transactions of the South African Philosophical Society, 7, 99 - 623 + i - xiv." type="journal article" year="1896">Péringuey, 1896: 443</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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||
.
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</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B8EF6540333F0028C8F7FDC2CC9E4441" pageId="12" pageNumber="43" type="materials_examined">
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||
<paragraph id="F04A36CB333F0028C8F7FDC2C94642B5" blockId="12.[151,1437,602,1997]" pageId="12" pageNumber="43">
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||
<emphasis id="C281EAD9333F0028C8F7FDC2CCD64258" bold="true" box="[151,448,602,629]" pageId="12" pageNumber="43">Type material examined.</emphasis>
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||
<materialsCitation id="409D3C96333F0028C9A7FDC2C94642B5" collectionCode="MNHN" pageId="12" pageNumber="43" specimenCount="2" specimenCount-male="2" typeStatus="lectotype">
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||
<typeStatus id="2F4E8869333F0028C9A7FDC2CF214259" box="[455,567,603,629]" pageId="12" pageNumber="43" type="lectotype">Lectotype</typeStatus>
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(present designation) of
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<taxonomicName id="37F54D48333F0028CB30FDC2CEEA4259" authorityName="Dejean" authorityYear="1829" box="[848,1020,603,629]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">
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<emphasis id="C281EAD9333F0028CB30FDC2CEEA4259" box="[848,1020,603,629]" italics="true" pageId="12" pageNumber="43">H. fuscoaeneus</emphasis>
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</taxonomicName>
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||
:
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<specimenCount id="E6F3FD42333F0028CC6BFDC2C9344259" box="[1035,1058,603,629]" pageId="12" pageNumber="43" type="male">♂</specimenCount>
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||
, labeled “
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<specimenCount id="E6F3FD42333F0028CCF2FDC2C9BF4259" box="[1170,1193,603,629]" pageId="12" pageNumber="43" type="male">♂</specimenCount>
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”, “
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<taxonomicName id="37F54D48333F0028CCACFDC2C88B4259" authority="Esch." authorityName="Esch." box="[1228,1437,603,629]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">fuscoaeneus Esch.</taxonomicName>
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ad Cap. Bon. Sp., D.” [Dejean’s handwriting], and “Ex Musaeo Chaudoir” (
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<collectionCode id="96E4AE0E333F0028CB88FDE6C95242B5" box="[1000,1092,639,665]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="43">MNHN</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
<paragraph id="F04A36CB333F0028C8A7FD3ACC9542CD" blockId="12.[151,1437,602,1997]" pageId="12" pageNumber="43">
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||
<materialsCitation id="409D3C96333F0028C8A7FD3ACC9542CD" collectionCode="MNHN" pageId="12" pageNumber="43" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
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||
<typeStatus id="2F4E8869333F0028C8A7FD3ACC394291" box="[199,303,675,701]" pageId="12" pageNumber="43" type="holotype">Holotype</typeStatus>
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||
of
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||
<taxonomicName id="37F54D48333F0028C932FD3ACCA04290" authorityName="Dejean" authorityYear="1829" box="[338,438,675,700]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="nanus">
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||
<emphasis id="C281EAD9333F0028C932FD3ACCA04290" box="[338,438,675,700]" italics="true" pageId="12" pageNumber="43">H. nanus</emphasis>
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||
</taxonomicName>
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||
:
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||
<specimenCount id="E6F3FD42333F0028C9A4FD3ACCCF4291" box="[452,473,675,701]" pageId="12" pageNumber="43" type="female">♀</specimenCount>
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||
, labeled “
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||
<specimenCount id="E6F3FD42333F0028CA2AFD3ACF4B4291" box="[586,605,675,701]" pageId="12" pageNumber="43" type="female">♀</specimenCount>
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||
”, “nanus mi..., ad Cap. Bon. Sp. [Dejean’s handwriting]”, and “Ex Musaeo Chaudoir” (
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||
<collectionCode id="96E4AE0E333F0028C97BFD5ECC6142CD" box="[283,375,711,737]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="12" pageNumber="43">MNHN</collectionCode>
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||
).
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||
</materialsCitation>
|
||
</paragraph>
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||
<paragraph id="F04A36CB333F0028C8A7FD73CC9E4441" blockId="12.[151,1437,602,1997]" pageId="12" pageNumber="43">
|
||
<emphasis id="C281EAD9333F0028C8A7FD73CF244328" bold="true" box="[199,562,746,772]" pageId="12" pageNumber="43">Additional material examined.</emphasis>
|
||
<materialsCitation id="409D3C96333F0028CA59FD73CE0E4305" collectionCode="ZIN" collectorName="Coll. Mus. Congo Col. P." country="South Africa" location="Spei." municipality="Cap." pageId="12" pageNumber="43" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<emphasis id="C281EAD9333F0028CA59FD73CFCF4328" bold="true" box="[569,729,746,772]" pageId="12" pageNumber="43">
|
||
<collectingCountry id="88E2765B333F0028CA59FD73CFC24328" box="[569,724,746,772]" name="South Africa" pageId="12" pageNumber="43">South Africa</collectingCountry>
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||
:
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||
</emphasis>
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||
<specimenCount id="E6F3FD42333F0028CABFFD72CE1C4329" box="[735,778,747,773]" pageId="12" pageNumber="43" type="male">1 ♂</specimenCount>
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||
,
|
||
<specimenCount id="E6F3FD42333F0028CB75FD72CE284329" box="[789,830,747,773]" pageId="12" pageNumber="43" type="female">1 ♀</specimenCount>
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||
, “
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||
<collectingMunicipality id="102EACB1333F0028CB36FD72CE9B4329" box="[854,909,747,773]" pageId="12" pageNumber="43">Cap.</collectingMunicipality>
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||
B.
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||
<location id="F52A6010333F0028CBD2FD72CEF94329" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028CBD2FD72CEF94329" box="[946,1007,747,773]" country="South Africa" municipality="Cap." name="Spei." pageId="12" pageNumber="43">Spei.</location>
|
||
”, “
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||
<collectorName id="5D00531D333F0028CC6FFD72C83E4329" box="[1039,1320,747,773]" pageId="12" pageNumber="43">Coll. Mus. Congo Col. P.</collectorName>
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||
Basilews- ky”, “
|
||
<taxonomicName id="37F54D48333F0028C8B9FC96CF494305" authority="Dej. P. Basilewsky" box="[217,607,783,809]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">H. fuscoaeneus Dej. P. Basilewsky</taxonomicName>
|
||
det., 19 ” (
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<collectionCode id="96E4AE0E333F0028CABCFC96CE064305" box="[732,784,783,809]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
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||
)
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||
</materialsCitation>
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||
;
|
||
<materialsCitation id="409D3C96333F0028CB44FC96C8244305" box="[804,1330,783,809]" collectionCode="ZIN" collectorName="Coll. Tschitsherine" country="South Africa" location="Cap." pageId="12" pageNumber="43" specimenCount="1" specimenCount-female="1">
|
||
<specimenCount id="E6F3FD42333F0028CB44FC96CE584305" box="[804,846,783,809]" pageId="12" pageNumber="43" type="female">1 ♀</specimenCount>
|
||
, “
|
||
<location id="F52A6010333F0028CB07FC96CE8B4305" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028CB07FC96CE8B4305" box="[871,925,783,809]" country="South Africa" name="Cap." pageId="12" pageNumber="43">Cap.</location>
|
||
b. sp.”, “
|
||
<collectorName id="5D00531D333F0028CC65FC96C9CB4305" box="[1029,1245,783,809]" pageId="12" pageNumber="43">Coll. Tschitshérine</collectorName>
|
||
” (
|
||
<collectionCode id="96E4AE0E333F0028CC96FC96C83C4305" box="[1270,1322,783,809]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028CD5EFC96CF7D4361" collectionCode="ZIN" country="South Africa" pageId="12" pageNumber="43" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="E6F3FD42333F0028CD5EFC96C87C4305" box="[1342,1386,783,809]" pageId="12" pageNumber="43" type="male">1 ♂</specimenCount>
|
||
, “C. b. Sp.”, “
|
||
<taxonomicName id="37F54D48333F0028C89CFCAACF034361" authority="Eschs. Cap." authorityName="Eschs. Cap." box="[252,533,819,845]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">fuscoaeneus Eschs. Cap.</taxonomicName>
|
||
” (
|
||
<collectionCode id="96E4AE0E333F0028CA4EFCAACF744361" box="[558,610,819,845]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028CA16FCAAC88E435D" collectionCode="ZIN" collectorName="S. Afr." country="South Africa" location="South Africa Expedition" pageId="12" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Western Cape">
|
||
<specimenCount id="E6F3FD42333F0028CA16FCAACFB44361" box="[630,674,819,845]" pageId="12" pageNumber="43" type="male">1 ♂</specimenCount>
|
||
, “
|
||
<collectorName id="5D00531D333F0028CADBFCAACE124361" box="[699,772,819,845]" pageId="12" pageNumber="43">S. Afr.</collectorName>
|
||
<collectingRegion id="3231F829333F0028CB6BFCAACE904361" box="[779,902,819,845]" country="South Africa" name="Western Cape" pageId="12" pageNumber="43">Cape Prov.</collectingRegion>
|
||
<locationDeviation id="E1F82FE9333F0028CBEDFCABC9D04361" box="[909,1222,818,845]" location="Mount Frere." pageId="12" pageNumber="43">
|
||
<quantity id="370D9B2E333F0028CBEDFCABCEFB4361" box="[909,1005,818,845]" metricMagnitude="4" metricUnit="m" metricValue="2.414016" pageId="12" pageNumber="43" unit="mi" value="15.0">15 miles</quantity>
|
||
ENE Mount Frère.
|
||
</locationDeviation>
|
||
6.III.51. No. 209”, “Swedish
|
||
<location id="F52A6010333F0028C96AFCCECF0C435D" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028C96AFCCECF0C435D" box="[266,538,855,881]" country="South Africa" name="South Africa Expedition" pageId="12" pageNumber="43" stateProvince="Western Cape">South Africa Expedition</location>
|
||
1950–1951 Brinck—Rudebeck”, “
|
||
<taxonomicName id="37F54D48333F0028CBC4FCCEC819435D" authority="Dej., Basilevsky" authorityName="Dej., Basilevsky" box="[932,1295,855,881]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">H. fuscoaeneus Dej., Basilevsky</taxonomicName>
|
||
det.” (
|
||
<collectionCode id="96E4AE0E333F0028CD3BFCCEC899435D" box="[1371,1423,855,881]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028C8F7FCE2C82043B9" box="[151,1334,891,917]" collectingDate="1993-11-17" collectionCode="MFNB, ZIN" collectorName="M. Uhlig" country="South Africa" latitude="-34.45" location="De Hoop Nat. Res." longLatPrecision="1199" longitude="20.4" pageId="12" pageNumber="43" specimenCount="11" specimenCount-female="4" specimenCount-male="7" stateProvince="Western Cape">
|
||
<specimenCount id="E6F3FD42333F0028C8F7FCE2CDCE43B9" box="[151,216,891,917]" pageId="12" pageNumber="43" type="male">7 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="E6F3FD42333F0028C884FCE2CC3243B9" box="[228,292,891,917]" pageId="12" pageNumber="43" type="female">4 ♀♀</specimenCount>
|
||
,
|
||
<collectingRegion id="3231F829333F0028C951FCE2CCB843B9" box="[305,430,891,917]" country="South Africa" name="Western Cape" pageId="12" pageNumber="43">Cape Prov.</collectingRegion>
|
||
,
|
||
<location id="F52A6010333F0028C9D9FCE2CF8643B9" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028C9D9FCE2CF8643B9" box="[441,656,891,917]" country="South Africa" latitude="-34.45" longLatPrecision="1199" longitude="20.4" name="De Hoop Nat. Res." pageId="12" pageNumber="43" stateProvince="Western Cape">De Hoop Nat. Res.</location>
|
||
,
|
||
<geoCoordinate id="95C1500C333F0028CAFBFCE2CFE143B9" box="[667,759,891,917]" degrees="34" direction="south" minutes="27" orientation="latitude" pageId="12" pageNumber="43" precision="925" value="-34.45">34°27’S</geoCoordinate>
|
||
<geoCoordinate id="95C1500C333F0028CA9EFCE2CE4843B9" box="[766,862,891,917]" degrees="20" direction="east" minutes="24" orientation="longitude" pageId="12" pageNumber="43" precision="925" value="20.4">20°24’E</geoCoordinate>
|
||
,
|
||
<date id="844B100B333F0028CB09FCE2CEFA43B9" box="[873,1004,891,917]" pageId="12" pageNumber="43" value="1993-11-17">
|
||
<collectingDate id="940FE9E3333F0028CB09FCE2CEFA43B9" box="[873,1004,891,917]" pageId="12" pageNumber="43" value="1993-11-17">17.XI.1993</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5D00531D333F0028CB97FCE2C94843B9" box="[1015,1118,891,917]" pageId="12" pageNumber="43">M. Uhlig</collectorName>
|
||
leg. (
|
||
<collectionCode id="96E4AE0E333F0028CCFCFCE2C9E643B9" box="[1180,1264,891,917]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="43">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="96E4AE0E333F0028CC9BFCE2C83843B9" box="[1275,1326,891,917]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028CD21FCE2C97F4395" collectingDate="1992-03-08" collectingDateMax="1992-03-09" collectingDateMin="1992-03-08" collectionCode="MFNB, ZIN" collectorName="H. J. Bremer" country="South Africa" elevation="1800" location="Barkly-East" pageId="12" pageNumber="43" specimenCount="7" specimenCount-female="3" specimenCount-male="4" stateProvince="Western Cape">
|
||
<specimenCount id="E6F3FD42333F0028CD21FCE2C89443B9" box="[1345,1410,891,917]" pageId="12" pageNumber="43" type="male">4 ♂♂</specimenCount>
|
||
,
|
||
<specimenCount id="E6F3FD42333F0028CDEEFCE2CDD44395" pageId="12" pageNumber="43" type="female">3 ♀♀</specimenCount>
|
||
, NE
|
||
<collectingRegion id="3231F829333F0028C899FC06CC604395" box="[249,374,927,953]" country="South Africa" name="Western Cape" pageId="12" pageNumber="43">Cape Prov.</collectingRegion>
|
||
,
|
||
<location id="F52A6010333F0028C9E0FC06CF1F4395" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028C9E0FC06CF1F4395" box="[384,521,927,953]" country="South Africa" name="Barkly-East" pageId="12" pageNumber="43" stateProvince="Western Cape">Barkly-East</location>
|
||
,
|
||
<quantity id="370D9B2E333F0028CA72FC06CF7E4395" box="[530,616,927,953]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="43" unit="m" value="1800.0">
|
||
<elevation id="7BD8D1F8333F0028CA72FC06CF7E4395" box="[530,616,927,953]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="12" pageNumber="43" unit="m" value="1800.0">1800 m</elevation>
|
||
</quantity>
|
||
,
|
||
<date id="844B100B333F0028CA13FC06CE174395" box="[627,769,927,953]" pageId="12" pageNumber="43" value="1992-03-08" valueMax="1992-03-09" valueMin="1992-03-08">
|
||
<collectingDate id="940FE9E3333F0028CA13FC06CE174395" box="[627,769,927,953]" pageId="12" pageNumber="43" value="1992-03-08" valueMax="1992-03-09" valueMin="1992-03-08">8–9.III.1992</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5D00531D333F0028CB6CFC06CE854395" box="[780,915,927,953]" pageId="12" pageNumber="43">H.J. Bremer</collectorName>
|
||
leg. (
|
||
<collectionCode id="96E4AE0E333F0028CBB0FC06C9324395" box="[976,1060,927,953]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="43">MFNB</collectionCode>
|
||
,
|
||
<collectionCode id="96E4AE0E333F0028CC4DFC06C9764395" box="[1069,1120,927,953]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028CC13FC06CEB443F1" collectingDate="1992-03-11" collectionCode="MFNB" collectorName="H. J. Bremer" country="South Africa" location="Craddock" municipality="Mountain Zebra Nat. Park" pageId="12" pageNumber="43" specimenCount="1" specimenCount-female="1" stateProvince="Western Cape">
|
||
<specimenCount id="E6F3FD42333F0028CC13FC06C98A4395" box="[1139,1180,927,953]" pageId="12" pageNumber="43" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="3231F829333F0028CCC9FC06C8304395" box="[1193,1318,927,953]" country="South Africa" name="Western Cape" pageId="12" pageNumber="43">Cape Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="102EACB1333F0028CD4FFC06CC5B43F1" pageId="12" pageNumber="43">Mountain Zebra Nat. Park</collectingMunicipality>
|
||
, p.
|
||
<location id="F52A6010333F0028C915FC5ACCF343F1" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028C915FC5ACCF343F1" box="[373,485,963,989]" country="South Africa" municipality="Mountain Zebra Nat. Park" name="Craddock" pageId="12" pageNumber="43" stateProvince="Western Cape">Craddock</location>
|
||
,
|
||
<date id="844B100B333F0028C991FC5ACF6643F1" box="[497,624,963,989]" pageId="12" pageNumber="43" value="1992-03-11">
|
||
<collectingDate id="940FE9E3333F0028C991FC5ACF6643F1" box="[497,624,963,989]" pageId="12" pageNumber="43" value="1992-03-11">11.III.1992</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5D00531D333F0028CA1CFC5ACE1343F1" box="[636,773,963,989]" pageId="12" pageNumber="43">H.J. Bremer</collectorName>
|
||
leg. (
|
||
<collectionCode id="96E4AE0E333F0028CB25FC5ACE8D43F1" box="[837,923,963,989]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="12" pageNumber="43">MFNB</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028CBCEFC5ACF1F442D" country="South Africa" location="Cape Prov." pageId="12" pageNumber="43" specimenCount="2" specimenCount-female="2" stateProvince="Western Cape">
|
||
<specimenCount id="E6F3FD42333F0028CBCEFC5ACEF943F1" box="[942,1007,963,989]" pageId="12" pageNumber="43" type="female">2 ♀♀</specimenCount>
|
||
, “S. A.
|
||
<collectingRegion id="3231F829333F0028CC28FC5AC9D143F1" box="[1096,1223,963,989]" country="South Africa" name="Western Cape" pageId="12" pageNumber="43">Cape Prov.</collectingRegion>
|
||
, Addo Elephant N. P., 25.03.93, fc. Arndt in” (cWR)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028CA73FC7ECDF14409" collectingDate="1998-04-09" collectorName="C. Saure" country="South Africa" location="Sterkfontein Dam Nat. Park" municipality="Harrismith" pageId="12" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Free State">
|
||
<specimenCount id="E6F3FD42333F0028CA73FC7ECF29442D" box="[531,575,999,1025]" pageId="12" pageNumber="43" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingRegion id="3231F829333F0028CA2AFC7ECFAB442D" box="[586,701,999,1025]" country="South Africa" name="Free State" pageId="12" pageNumber="43">Free State</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="102EACB1333F0028CAA7FC7ECE55442D" box="[711,835,999,1025]" pageId="12" pageNumber="43">Harrismith</collectingMunicipality>
|
||
,
|
||
<location id="F52A6010333F0028CB2EFC7EC991442D" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028CB2EFC7EC991442D" box="[846,1159,999,1025]" country="South Africa" municipality="Harrismith" name="Sterkfontein Dam Nat. Park" pageId="12" pageNumber="43" stateProvince="Free State">Sterkfontein Dam Nat. Park</location>
|
||
,
|
||
<date id="844B100B333F0028CCF2FC7EC814442D" box="[1170,1282,999,1025]" pageId="12" pageNumber="43" value="1998-04-09">
|
||
<collectingDate id="940FE9E3333F0028CCF2FC7EC814442D" box="[1170,1282,999,1025]" pageId="12" pageNumber="43" value="1998-04-09">9.IV.1998</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5D00531D333F0028CD6DFC7EC87B442D" box="[1293,1389,999,1025]" pageId="12" pageNumber="43">C. Saure</collectorName>
|
||
leg. (cWR)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028C891FB92C88E4409" box="[241,1432,1034,1061]" collectingDate="2001-12" collectionCode="ZIN" collectorName="A. Frolov" country="South Africa" latitude="-26.9" location="Farm Abel" longLatPrecision="1239" longitude="27.583334" municipality="Parys Distr." pageId="12" pageNumber="43" specimenCount="1" specimenCount-male="1" stateProvince="Free State">
|
||
<specimenCount id="E6F3FD42333F0028C891FB92CC0B4409" box="[241,285,1035,1061]" pageId="12" pageNumber="43" type="male">1 ♂</specimenCount>
|
||
,
|
||
<collectingRegion id="3231F829333F0028C948FB92CCCD4409" box="[296,475,1035,1061]" country="South Africa" name="Free State" pageId="12" pageNumber="43">Free State Prov.</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="102EACB1333F0028C985FB92CF7C4409" box="[485,618,1035,1061]" pageId="12" pageNumber="43">Parys Distr.</collectingMunicipality>
|
||
,
|
||
<location id="F52A6010333F0028CA14FB92CFFB4409" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028CA14FB92CFFB4409" box="[628,749,1035,1061]" country="South Africa" latitude="-26.9" longLatPrecision="1239" longitude="27.583334" municipality="Parys Distr." name="Farm Abel" pageId="12" pageNumber="43" stateProvince="Free State">Farm Abel</location>
|
||
,
|
||
<geoCoordinate id="95C1500C333F0028CA98FB93CE424409" box="[760,852,1034,1061]" degrees="26" direction="south" minutes="54" orientation="latitude" pageId="12" pageNumber="43" precision="925" value="-26.9">26°54’S</geoCoordinate>
|
||
<geoCoordinate id="95C1500C333F0028CB3AFB93CEAC4409" box="[858,954,1034,1061]" degrees="27" direction="east" minutes="35" orientation="longitude" pageId="12" pageNumber="43" precision="925" value="27.583334">27°35’E</geoCoordinate>
|
||
, PFT dung,
|
||
<date id="844B100B333F0028CC20FB92C9BF4409" box="[1088,1193,1035,1061]" pageId="12" pageNumber="43" value="2001-12">
|
||
<collectingDate id="940FE9E3333F0028CC20FB92C9BF4409" box="[1088,1193,1035,1061]" pageId="12" pageNumber="43" value="2001-12">XII.2001</collectingDate>
|
||
</date>
|
||
,
|
||
<collectorName id="5D00531D333F0028CCD1FB92C8084409" box="[1201,1310,1035,1061]" pageId="12" pageNumber="43">A. Frolov</collectorName>
|
||
leg. (
|
||
<collectionCode id="96E4AE0E333F0028CD3BFB92C8994409" box="[1371,1423,1035,1061]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028C8F7FBB6CED64465" box="[151,960,1071,1097]" collectionCode="ZIN" collectorName="Coll. Tschitsherine" country="South Africa" location="Caffar." pageId="12" pageNumber="43" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
|
||
<specimenCount id="E6F3FD42333F0028C8F7FBB6CDD34465" box="[151,197,1071,1097]" pageId="12" pageNumber="43" type="male">1 ♂</specimenCount>
|
||
,
|
||
<specimenCount id="E6F3FD42333F0028C8B3FBB6CC164465" box="[211,256,1071,1097]" pageId="12" pageNumber="43" type="female">1 ♀</specimenCount>
|
||
, “
|
||
<location id="F52A6010333F0028C97AFBB6CC784465" LSID="urn:lsid:plazi:treatment:785C87DD333F002AC8F7FEF7CC5F460A:F52A6010333F0028C97AFBB6CC784465" box="[282,366,1071,1097]" country="South Africa" name="Caffar." pageId="12" pageNumber="43">Caffar.</location>
|
||
”, “
|
||
<taxonomicName id="37F54D48333F0028C9F1FBB6CF7A4465" authority="Boh." authorityName="Boh." box="[401,620,1071,1097]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="subaeneus">H. subaeneus Boh.</taxonomicName>
|
||
”, “
|
||
<collectorName id="5D00531D333F0028CAEFFBB6CE704465" box="[655,870,1071,1097]" pageId="12" pageNumber="43">Coll. Tschitshérine</collectorName>
|
||
” (
|
||
<collectionCode id="96E4AE0E333F0028CBE3FBB6CEA14465" box="[899,951,1071,1097]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation id="409D3C96333F0028CBADFBB6CC924441" collectionCode="ZIN" collectorName="Coll. Tschitsherine" country="South Africa" pageId="12" pageNumber="43" specimenCount="1" specimenCount-male="1">
|
||
<specimenCount id="E6F3FD42333F0028CBADFBB6CEED4465" box="[973,1019,1071,1097]" pageId="12" pageNumber="43" type="male">1 ♂</specimenCount>
|
||
, “
|
||
<taxonomicName id="37F54D48333F0028CC75FBB6C81C4465" authority="Esch." authorityName="Esch." box="[1045,1290,1071,1097]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">H. fuscoaeneus Esch.</taxonomicName>
|
||
leg.”, “
|
||
<collectorName id="5D00531D333F0028CD03FBB6CC394441" pageId="12" pageNumber="43">Coll. Tschitshérine</collectorName>
|
||
” (
|
||
<collectionCode id="96E4AE0E333F0028C928FBCACC6A4441" box="[328,380,1107,1133]" country="Russia" httpUri="http://grbio.org/cool/z2y8-2jfr" name="Russian Academy of Sciences, Zoological Institute, Zoological Museum" pageId="12" pageNumber="43" type="Museum">ZIN</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8EF6540333F0028C8A7FBEFCDFD4735" pageId="12" pageNumber="43" type="diagnosis">
|
||
<paragraph id="F04A36CB333F0028C8A7FBEFCE7E46A5" blockId="12.[151,1437,602,1997]" pageId="12" pageNumber="43">
|
||
<emphasis id="C281EAD9333F0028C8A7FBEFCC5444BC" bold="true" box="[199,322,1142,1168]" pageId="12" pageNumber="43">Diagnosis.</emphasis>
|
||
Among
|
||
<taxonomicName id="37F54D48333F0028C9C6FBEECF7F44BD" box="[422,617,1143,1169]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="C281EAD9333F0028C9C6FBEECF0644BD" box="[422,528,1143,1169]" italics="true" pageId="12" pageNumber="43">Harpalus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
from Southern Africa,
|
||
<taxonomicName id="37F54D48333F0028CB13FBEEC93644BD" box="[883,1056,1143,1169]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333F0028CB13FBEEC93644BD" box="[883,1056,1143,1169]" italics="true" pageId="12" pageNumber="43">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
can be recognized by the combination of the following distinctive features: small (length
|
||
<quantity id="370D9B2E333F0028CB4EFB02CEAD4499" box="[814,955,1179,1205]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" metricValueMax="6.1" metricValueMin="4.9" pageId="12" pageNumber="43" unit="mm" value="5.5" valueMax="6.1" valueMin="4.9">4.9–6.1 mm</quantity>
|
||
); body (
|
||
<figureCitation id="68CE2A4E333F0028CC7AFB02C98B4499" box="[1050,1181,1179,1205]" captionStart="FIGURES 29–32" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[180,1402,189,1896]" captionTargetId="figure-16@13.[172,1415,181,1901]" captionTargetPageId="13" captionText="FIGURES 29–32. Harpalus spp., habitus. 29–31, H. fuscoaeneus Dejean, 1929 (29, S Africa; 30, lectotype; 31, holotype of H. nanus Dejean, 1829); 32, H. australasiae Dejean, 1829 (holotype). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222947" httpUri="https://zenodo.org/record/5222947/files/figure.png" pageId="12" pageNumber="43">Figs 29–31</figureCitation>
|
||
) dark brown to black, usually with more or less distinct green or copper luster on dorsum; palpi, antennomeres 3–11, in many specimens also femora, tibial apices and tarsi more or less markedly infuscate; pronotum widest before or in the middle, almost equally narrowed apically and basally, with rounded (occasionally almost straight basally) sides and rounded (occasionally obtusangular) basal angles; pronotal basal edge glabrous; pronotal disc impunctate, at most with a few very fine punctures within small and shallow basal foveae; elytra with very shallow, almost indistinct preapical sinuation, with (occasionally without) discal pore on interval 3, and with parascutellar striole in many specimens strongly reduced; microsculpture on head, pronotum and elytra consisting of more or less isodiametric meshes; metepisternum notably longer than wide, strongly narrowed posteriorly; metacoxa without additional median and posteromedial pores; median lobe of aedeagus (
|
||
<figureCitation id="68CE2A4E333F0028CAD3FA46CE2045D6" box="[691,822,1502,1530]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1045,1069]" captionTargetBox="[158,1426,188,1013]" captionTargetId="figure-23@14.[151,1436,181,1021]" captionTargetPageId="14" captionText="FIGURES 33–36. Harpalus fuscoaeneus Dejean, 1929 (Cape of Good Hope). 33, gonocoxite; 34, laterotergite, gonosubcoxite, and gonocoxite; 35, 36, median lobe of aedeagus. Lateral (33, 35), ventral (34) and dorsal (36) views. Scale bar = 0. 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222949" httpUri="https://zenodo.org/record/5222949/files/figure.png" pageId="12" pageNumber="43">Figs 35, 36</figureCitation>
|
||
) S-formed, strongly bent ventrally at the middle, with apical half markedly curved to the right; its ventral margin in lateral view not or only very slightly convex; terminal lamella in dorsal view notably longer than wide, almost parallel-sided, rounded apically, in lateral view markedly protruding ventrally just at apex; internal sac with two short spines: distal spine at the middle, and proximal one in basal portion close to basal bulb. Female genitalia—
|
||
<figureCitation id="68CE2A4E333F0028CA84F9F6CE7546A5" box="[740,867,1647,1673]" captionStart="FIGURES 33–36" captionStartId="14.[151,264,1045,1069]" captionTargetBox="[158,1426,188,1013]" captionTargetId="figure-23@14.[151,1436,181,1021]" captionTargetPageId="14" captionText="FIGURES 33–36. Harpalus fuscoaeneus Dejean, 1929 (Cape of Good Hope). 33, gonocoxite; 34, laterotergite, gonosubcoxite, and gonocoxite; 35, 36, median lobe of aedeagus. Lateral (33, 35), ventral (34) and dorsal (36) views. Scale bar = 0. 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5222949" httpUri="https://zenodo.org/record/5222949/files/figure.png" pageId="12" pageNumber="43">Figs 33, 34</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="F04A36CB333F0028C8A7F90ACDFD4735" blockId="12.[151,1437,602,1997]" pageId="12" pageNumber="43">
|
||
Proportions (
|
||
<specimenCount id="E6F3FD42333F0028C93CF90BCC8B4681" box="[348,413,1682,1709]" pageId="12" pageNumber="43" type="male">5 ♂♂</specimenCount>
|
||
and
|
||
<specimenCount id="E6F3FD42333F0028C9B7F90BCF0E4681" box="[471,536,1682,1709]" pageId="12" pageNumber="43" type="female">5 ♀♀</specimenCount>
|
||
measured): HWmax/PWmax 0.64–0.71; HWmin/PWmax 0.50–0.57; HWmax/ HWmin 1.18–1.32; PWmax/PL 1.36–1.48; PWmax/PWmin 1.13–1.24; EL/EW
|
||
<quantity id="370D9B2E333F0028CC44F92FC9A546FD" box="[1060,1203,1718,1745]" metricMagnitude="-2" metricUnit="m" metricValue="3.8862" metricValueMax="3.9624" metricValueMin="3.81" pageId="12" pageNumber="43" unit="in" value="1.53" valueMax="1.56" valueMin="1.5">1.50–1.56 in</quantity>
|
||
male and
|
||
<quantity id="370D9B2E333F0028CD4CF92ECDBB46D9" metricMagnitude="-2" metricUnit="m" metricValue="3.7084" metricValueMax="3.7592" metricValueMin="3.6576" pageId="12" pageNumber="43" unit="in" value="1.46" valueMax="1.48" valueMin="1.44">1.44–1.48 in</quantity>
|
||
female; EL/PL
|
||
<quantity id="370D9B2E333F0028C908F943CCE146D9" box="[360,503,1754,1781]" metricMagnitude="-2" metricUnit="m" metricValue="6.4516" metricValueMax="6.731" metricValueMin="6.1722" pageId="12" pageNumber="43" unit="in" value="2.54" valueMax="2.65" valueMin="2.43">2.43–2.65 in</quantity>
|
||
male and
|
||
<quantity id="370D9B2E333F0028CA13F943CE1546D9" box="[627,771,1754,1781]" metricMagnitude="-2" metricUnit="m" metricValue="6.4643" metricValueMax="6.7056" metricValueMin="6.223" pageId="12" pageNumber="43" unit="in" value="2.545" valueMax="2.64" valueMin="2.45">2.45–2.64 in</quantity>
|
||
female; EW/PWmax
|
||
<quantity id="370D9B2E333F0028CC61F942C98746D9" box="[1025,1169,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="2.9972" metricValueMax="3.0225999999999997" metricValueMin="2.9718" pageId="12" pageNumber="43" unit="in" value="1.18" valueMax="1.19" valueMin="1.17">1.17–1.19 in</quantity>
|
||
male and
|
||
<quantity id="370D9B2E333F0028CD6DF942C88A46D9" box="[1293,1436,1755,1781]" metricMagnitude="-2" metricUnit="m" metricValue="3.1369" metricValueMax="3.2004" metricValueMin="3.0734" pageId="12" pageNumber="43" unit="in" value="1.2349999999999999" valueMax="1.26" valueMin="1.21">1.21–1.26 in</quantity>
|
||
female.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B8EF6540333F0028C8A7F8BBCE6047E1" pageId="12" pageNumber="43" type="distribution">
|
||
<paragraph id="F04A36CB333F0028C8A7F8BBCE034711" blockId="12.[151,1437,602,1997]" box="[199,789,1826,1853]" pageId="12" pageNumber="43">
|
||
<emphasis id="C281EAD9333F0028C8A7F8BBCC774710" bold="true" box="[199,353,1826,1852]" pageId="12" pageNumber="43">Distribution.</emphasis>
|
||
Widely distributed in Southern Africa.
|
||
</paragraph>
|
||
<paragraph id="F04A36CB333F0028C8A7F8DFCE6047E1" blockId="12.[151,1437,602,1997]" pageId="12" pageNumber="43">
|
||
<emphasis id="C281EAD9333F0028C8A7F8DFCC74474C" bold="true" box="[199,354,1862,1888]" pageId="12" pageNumber="43">Comparison.</emphasis>
|
||
In external characters and male genitalia,
|
||
<taxonomicName id="37F54D48333F0028CB22F8DECEF8474D" box="[834,1006,1863,1889]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333F0028CB22F8DECEF8474D" box="[834,1006,1863,1889]" italics="true" pageId="12" pageNumber="43">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
is very similar to
|
||
<taxonomicName id="37F54D48333F0028CCDDF8DEC824474C" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[1213,1330,1863,1888]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="asemus">
|
||
<emphasis id="C281EAD9333F0028CCDDF8DEC824474C" box="[1213,1330,1863,1888]" italics="true" pageId="12" pageNumber="43">H. asemus</emphasis>
|
||
</taxonomicName>
|
||
differing slightly from it in shape of the median lobe and in, on average, smaller body size. In
|
||
<taxonomicName id="37F54D48333F0028CC04F8F2C80647A9" box="[1124,1296,1899,1925]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333F0028CC04F8F2C80647A9" box="[1124,1296,1899,1925]" italics="true" pageId="12" pageNumber="43">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
, the median lobe is bent ventrally at the middle (more distally than in
|
||
<taxonomicName id="37F54D48333F0028CB40F816CE834784" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[800,917,1935,1960]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="43" phylum="Arthropoda" rank="species" species="asemus">
|
||
<emphasis id="C281EAD9333F0028CB40F816CE834784" box="[800,917,1935,1960]" italics="true" pageId="12" pageNumber="43">H. asemus</emphasis>
|
||
</taxonomicName>
|
||
) and its terminal lamella narrower. These two species seem to be vicariants descending from one ancestor form.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="A48A6643333E0029C8F7F81CC90C47ED" ID-DOI="http://doi.org/10.5281/zenodo.5222947" ID-Zenodo-Dep="5222947" httpUri="https://zenodo.org/record/5222947/files/figure.png" pageId="13" pageNumber="44" startId="13.[151,264,1925,1949]" targetBox="[180,1402,189,1896]" targetPageId="13">
|
||
<paragraph id="F04A36CB333E0029C8F7F81CC90C47ED" blockId="13.[151,1437,1925,1985]" pageId="13" pageNumber="44">
|
||
<emphasis id="C281EAD9333E0029C8F7F81CCC4247B1" bold="true" box="[151,340,1925,1949]" pageId="13" pageNumber="44">FIGURES 29–32.</emphasis>
|
||
<taxonomicName id="37F54D48333E0029C93AF81CCCFA47B1" box="[346,492,1925,1949]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="C281EAD9333E0029C93AF81CCCAC47B1" box="[346,442,1925,1949]" italics="true" pageId="13" pageNumber="44">Harpalus</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
, habitus. 29–31,
|
||
<taxonomicName id="37F54D48333E0029CAF7F81FCEA947B1" authority="Dejean, 1929" authorityName="Dejean" authorityYear="1929" box="[663,959,1925,1949]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333E0029CAF7F81FCE2447B1" box="[663,818,1925,1949]" italics="true" pageId="13" pageNumber="44">H. fuscoaeneus</emphasis>
|
||
Dejean, 1929
|
||
</taxonomicName>
|
||
(29, S Africa; 30, lectotype; 31, holotype of
|
||
<taxonomicName id="37F54D48333E0029CDE4F81FCC7247ED" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="nanus">
|
||
<emphasis id="C281EAD9333E0029CDE4F81FCDC247ED" italics="true" pageId="13" pageNumber="44">H. nanus</emphasis>
|
||
<bibRefCitation id="94644B3A333E0029C8BAF830CC7247ED" author="Dejean, P. F. M. A." box="[218,356,1961,1985]" pageId="13" pageNumber="44" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829</bibRefCitation>
|
||
</taxonomicName>
|
||
); 32,
|
||
<taxonomicName id="37F54D48333E0029C9FDF833CFD147ED" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" box="[413,711,1961,1985]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="44" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="C281EAD9333E0029C9FDF833CF2F47ED" box="[413,569,1961,1985]" italics="true" pageId="13" pageNumber="44">H. australasiae</emphasis>
|
||
<bibRefCitation id="94644B3A333E0029CA5FF830CFD147ED" author="Dejean, P. F. M. A." box="[575,711,1961,1985]" pageId="13" pageNumber="44" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829</bibRefCitation>
|
||
</taxonomicName>
|
||
(holotype). Scale bars = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="A48A6643333D002AC8F7FB8CC861447D" ID-DOI="http://doi.org/10.5281/zenodo.5222949" ID-Zenodo-Dep="5222949" httpUri="https://zenodo.org/record/5222949/files/figure.png" pageId="14" pageNumber="45" startId="14.[151,264,1045,1069]" targetBox="[158,1426,188,1013]" targetPageId="14">
|
||
<paragraph id="F04A36CB333D002AC8F7FB8CC861447D" blockId="14.[151,1436,1045,1105]" pageId="14" pageNumber="45">
|
||
<emphasis id="C281EAD9333D002AC8F7FB8CCC424401" bold="true" box="[151,340,1045,1069]" pageId="14" pageNumber="45">FIGURES 33–36.</emphasis>
|
||
<taxonomicName id="37F54D48333D002AC939FB8CCFD14401" authority="Dejean, 1929" authorityName="Dejean" authorityYear="1929" box="[345,711,1045,1069]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333D002AC939FB8CCF2D4401" box="[345,571,1045,1069]" italics="true" pageId="14" pageNumber="45">Harpalus fuscoaeneus</emphasis>
|
||
Dejean, 1929
|
||
</taxonomicName>
|
||
(Cape of Good Hope). 33, gonocoxite; 34, laterotergite, gonosubcoxite, and gonocoxite; 35, 36, median lobe of aedeagus. Lateral (33, 35), ventral (34) and dorsal (36) views. Scale bar = 0. 5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="B8EF6540333D002AC8A7FB19CC5F460A" pageId="14" pageNumber="45" type="discussion">
|
||
<paragraph id="F04A36CB333D002AC8A7FB19CC6F4529" blockId="14.[151,1437,1151,1574]" pageId="14" pageNumber="45">
|
||
<emphasis id="C281EAD9333D002AC8A7FB19CC2D44B6" bold="true" box="[199,315,1152,1178]" pageId="14" pageNumber="45">Remarks.</emphasis>
|
||
<bibRefCitation id="94644B3A333D002AC922FBE6CCF244B6" author="Dejean, P. F. M. A." box="[322,484,1151,1178]" pageId="14" pageNumber="45" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean (1829)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="37F54D48333D002ACA3EFB19CE1F44B6" authorityName="Dejean" authorityYear="1929" box="[606,777,1152,1178]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333D002ACA3EFB19CE1F44B6" box="[606,777,1152,1178]" italics="true" pageId="14" pageNumber="45">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="3231F829333D002ACB2FFB19CE9F44B6" box="[847,905,1152,1178]" country="South Africa" name="Western Cape" pageId="14" pageNumber="45">Cape</collectingRegion>
|
||
of Good Hope (“cap de Bonne-Espérance”) on the basis of more than
|
||
<specimenCount id="E6F3FD42333D002AC9FCFB3DCF204491" box="[412,566,1187,1214]" pageId="14" pageNumber="45" type="generic">one specimen</specimenCount>
|
||
(the ranges of measurements are given). The single available specimen with Dejean’s original labels at MNHN is designated here as
|
||
<typeStatus id="2F4E8869333D002ACB63FB5ECE7C44CD" box="[771,874,1223,1249]" pageId="14" pageNumber="45" type="lectotype">lectotype</typeStatus>
|
||
(
|
||
<figureCitation id="68CE2A4E333D002ACB18FB5ECEDD44CE" box="[888,971,1223,1250]" captionStart="FIGURES 29–32" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[180,1402,189,1896]" captionTargetId="figure-16@13.[172,1415,181,1901]" captionTargetPageId="13" captionText="FIGURES 29–32. Harpalus spp., habitus. 29–31, H. fuscoaeneus Dejean, 1929 (29, S Africa; 30, lectotype; 31, holotype of H. nanus Dejean, 1829); 32, H. australasiae Dejean, 1829 (holotype). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222947" httpUri="https://zenodo.org/record/5222947/files/figure.png" pageId="14" pageNumber="45">Fig. 30</figureCitation>
|
||
). Its characteristics well correspond to the original description.
|
||
</paragraph>
|
||
<paragraph id="F04A36CB333D002AC8A7FA96CC5F460A" blockId="14.[151,1437,1151,1574]" pageId="14" pageNumber="45">
|
||
The species is variable in coloration and shape of pronotum.
|
||
<bibRefCitation id="94644B3A333D002ACB0CFA96C9284506" author="Basilewsky, P." box="[876,1086,1295,1322]" pageId="14" pageNumber="45" pagination="1 - 333" refId="ref14741" refString="Basilewsky, P. (1951) Revision generale des Harpalinae d'Afrique et de Madagascar (Coleoptera Carabidae). Deuxieme partie. Annales du Musee Royal du Congo Belge, Serie in 8 °, Sciences Zoologiques, 9, 1 - 333, pls. I - VI." type="journal article" year="1951">Basilewsky (1951)</bibRefCitation>
|
||
treated
|
||
<taxonomicName id="37F54D48333D002ACCF7FA96C85C4505" authorityName="Dejean" authorityYear="1829" box="[1175,1354,1295,1321]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="nanus">
|
||
<emphasis id="C281EAD9333D002ACCF7FA96C85C4505" box="[1175,1354,1295,1321]" italics="true" pageId="14" pageNumber="45">Harpalus nanus</emphasis>
|
||
</taxonomicName>
|
||
, which was described from
|
||
<specimenCount id="E6F3FD42333D002AC91BFAADCCE14561" box="[379,503,1331,1358]" pageId="14" pageNumber="45" type="female">one female</specimenCount>
|
||
(
|
||
<figureCitation id="68CE2A4E333D002ACA67FAAACF4F4562" box="[519,601,1331,1358]" captionStart="FIGURES 29–32" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[180,1402,189,1896]" captionTargetId="figure-16@13.[172,1415,181,1901]" captionTargetPageId="13" captionText="FIGURES 29–32. Harpalus spp., habitus. 29–31, H. fuscoaeneus Dejean, 1929 (29, S Africa; 30, lectotype; 31, holotype of H. nanus Dejean, 1829); 32, H. australasiae Dejean, 1829 (holotype). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222947" httpUri="https://zenodo.org/record/5222947/files/figure.png" pageId="14" pageNumber="45">Fig. 31</figureCitation>
|
||
) also from
|
||
<collectingRegion id="3231F829333D002ACABCFAADCE004562" box="[732,790,1332,1358]" country="South Africa" name="Western Cape" pageId="14" pageNumber="45">Cape</collectingRegion>
|
||
of Good Hope (“cap de Bonne-Espérance”), as synonym of
|
||
<taxonomicName id="37F54D48333D002AC8D6FAC1CC74455E" authorityName="Dejean" authorityYear="1929" box="[182,354,1368,1394]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333D002AC8D6FAC1CC74455E" box="[182,354,1368,1394]" italics="true" pageId="14" pageNumber="45">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
, but its status needs further study because this female is rather distinct in having pronotum with very fine and short setae along the middle portion of basal edge; in addition, it has elytra with short parascutellar (abbreviate) striole and without discal pore on interval 3 and dorsum dark brown, without green or copper luster. The body length (
|
||
<quantity id="370D9B2E333D002AC900FA5ACCAC45F2" box="[352,442,1475,1502]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" pageId="14" pageNumber="45" unit="mm" value="5.9">5.9 mm</quantity>
|
||
) and proportions (HWmax/PWmax 0.67; HWmin/PWmax 0.52; HWmax/HWmin 1.27; PWmax/PL 1.44; EL/EW 1.48; EL/PL 2.64; EW/PWmax 1.24) are within variation of other examined specimens of
|
||
<taxonomicName id="37F54D48333D002AC8F7F995CC54460A" authorityName="Dejean" authorityYear="1929" box="[151,322,1548,1574]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333D002AC8F7F995CC54460A" box="[151,322,1548,1574]" italics="true" pageId="14" pageNumber="45">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |