265 lines
32 KiB
XML
265 lines
32 KiB
XML
<document id="8D9954C5997471059B4B3B85FC8F9029" ID-DOI="10.11646/zootaxa.5020.1.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="5222931" ID-ZooBank="882BBB9D-6E5B-4CE5-99DF-E91AC7971EB5" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629386356657" checkinUser="plazi" docAuthor="Kataev, Boris M." docDate="2021" docId="785C87DD333D002BC8F7F9EEC9CC43AD" docLanguage="en" docName="zootaxa.5020.1.2.pdf" docOrigin="Zootaxa 5020 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpalus australasiae Dejean 1829" docType="treatment" docVersion="3" lastPageNumber="46" masterDocId="8465FFA533330024C860FF99CD16402C" masterDocTitle="On some Afrotropical Harpalus, with description of two new species, and remarks on Hypharpax australis (Coleoptera: Carabidae): misidentification, mislabeling and introduction to the Australian region" masterLastPageNumber="56" masterPageNumber="31" pageNumber="45" updateTime="1699070990175" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="52B27F4DE0E7347DF8995ADF3CD05F15">On some Afrotropical Harpalus, with description of two new species, and remarks on Hypharpax australis (Coleoptera: Carabidae): misidentification, mislabeling and introduction to the Australian region</mods:title>
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<mods:namePart id="3511302DABAA8C48F70187D574EE4E0D">Kataev, Boris M.</mods:namePart>
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<mods:title id="1DF36539FA6821B23627F318EE2072F5">Zootaxa</mods:title>
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<mods:part id="D78310A95EDC03100DFC9EDBD257EA14">
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<mods:date id="84DE80E1E07B387FF09BABF041BED75B">2021</mods:date>
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<mods:number id="EBC56A55A8A06E0097799285D9386F71">2021-08-11</mods:number>
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<mods:number id="DE5A5ED7FD7A8F64433C3319F0197065">5020</mods:number>
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<treatment id="785C87DD333D002BC8F7F9EEC9CC43AD" ID-DOI="http://doi.org/10.5281/zenodo.5222945" ID-Zenodo-Dep="5222945" LSID="urn:lsid:plazi:treatment:785C87DD333D002BC8F7F9EEC9CC43AD" httpUri="http://treatment.plazi.org/id/785C87DD333D002BC8F7F9EEC9CC43AD" lastPageId="15" lastPageNumber="46" pageId="14" pageNumber="45">
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<subSubSection id="B8EF6540333D002AC8F7F9EECF5A46BD" box="[151,588,1655,1682]" pageId="14" pageNumber="45" type="nomenclature">
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<paragraph id="F04A36CB333D002AC8F7F9EECF5A46BD" blockId="14.[151,588,1655,1718]" box="[151,588,1655,1682]" pageId="14" pageNumber="45">
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<heading id="AB0281A7333D002AC8F7F9EECF5A46BD" bold="true" box="[151,588,1655,1682]" fontSize="11" level="1" pageId="14" pageNumber="45" reason="1">
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<taxonomicName id="37F54D48333D002AC8F7F9EECF5A46BD" ID-CoL="3JSYP" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" box="[151,588,1655,1682]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="australasiae">
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<emphasis id="C281EAD9333D002AC8F7F9EECF5A46BD" bold="true" box="[151,588,1655,1682]" pageId="14" pageNumber="45">
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<emphasis id="C281EAD9333D002AC8F7F9EECCB246BE" bold="true" box="[151,420,1655,1682]" italics="true" pageId="14" pageNumber="45">Harpalus australasiae</emphasis>
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<bibRefCitation id="94644B3A333D002AC9CBF9EECF5A46BD" author="Dejean, P. F. M. A." box="[427,588,1655,1681]" pageId="14" pageNumber="45" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829</bibRefCitation>
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</emphasis>
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||
</taxonomicName>
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||
</heading>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B8EF6540333D002AC8F7F902CDEF469A" box="[151,249,1691,1718]" pageId="14" pageNumber="45" type="description">
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<paragraph id="F04A36CB333D002AC8F7F902CDEF469A" blockId="14.[151,588,1655,1718]" box="[151,249,1691,1718]" pageId="14" pageNumber="45">
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(
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<figureCitation id="68CE2A4E333D002AC8C0F902CDE7469A" box="[160,241,1691,1718]" captionStart="FIGURES 29–32" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[180,1402,189,1896]" captionTargetId="figure-16@13.[172,1415,181,1901]" captionTargetPageId="13" captionText="FIGURES 29–32. Harpalus spp., habitus. 29–31, H. fuscoaeneus Dejean, 1929 (29, S Africa; 30, lectotype; 31, holotype of H. nanus Dejean, 1829); 32, H. australasiae Dejean, 1829 (holotype). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222947" httpUri="https://zenodo.org/record/5222947/files/figure.png" pageId="14" pageNumber="45">Fig. 32</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="B8EF6540333D002AC8F7F979CF5446D4" box="[151,578,1760,1784]" pageId="14" pageNumber="45" type="reference_group">
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||
<paragraph id="F04A36CB333D002AC8F7F979CF5446D4" blockId="14.[151,578,1760,1784]" box="[151,578,1760,1784]" pageId="14" pageNumber="45">
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||
<treatmentCitationGroup id="D0E511E5333D002AC8F7F979CF5446D4" box="[151,578,1760,1784]" pageId="14" pageNumber="45">
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||
<treatmentCitation id="715410DA333D002AC8F7F979CF2846D4" author="Dejean, P. F. M. A." box="[151,574,1760,1784]" page="386" pageId="14" pageNumber="45" year="1829">
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||
<taxonomicName id="37F54D48333D002AC8F7F979CF2846D4" ID-CoL="3JSYP" authority="Dejean, 1829: 386" authorityName="Dejean" authorityPageNumber="386" authorityYear="1829" box="[151,574,1760,1784]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="australasiae">
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<emphasis id="C281EAD9333D002AC8F7F979CC6C46D4" box="[151,378,1760,1784]" italics="true" pageId="14" pageNumber="45">Harpalus australasiae</emphasis>
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<bibRefCitation id="94644B3A333D002AC9E1F979CF2846D4" author="Dejean, P. F. M. A." box="[385,574,1760,1784]" pageId="14" pageNumber="45" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829: 386</bibRefCitation>
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||
</taxonomicName>
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||
</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B8EF6540333D002AC8F7F8BFC80B4748" pageId="14" pageNumber="45" type="materials_examined">
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||
<paragraph id="F04A36CB333D002AC8F7F8BFC80B4748" blockId="14.[151,1437,1830,2037]" pageId="14" pageNumber="45">
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||
<emphasis id="C281EAD9333D002AC8F7F8BFCCAA476C" bold="true" box="[151,444,1830,1856]" pageId="14" pageNumber="45">Type material examined.</emphasis>
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<materialsCitation id="409D3C96333D002AC9A2F8BFC80B4748" collectingDate="1978-09-26" collectionCode="MNHN" collectorName="P. M. Tohus" country="Netherlands" location="Dejean's" pageId="14" pageNumber="45" specimenCount="2" specimenCount-female="2" typeStatus="holotype">
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<typeStatus id="2F4E8869333D002AC9A2F8BFCF3C476C" box="[450,554,1830,1856]" pageId="14" pageNumber="45" type="holotype">Holotype</typeStatus>
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of
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<taxonomicName id="37F54D48333D002ACA2CF8BECFE1476C" authorityName="Dejean" authorityYear="1829" box="[588,759,1830,1856]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="australasiae">
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<emphasis id="C281EAD9333D002ACA2CF8BECFE1476C" box="[588,759,1830,1856]" italics="true" pageId="14" pageNumber="45">H. australasiae</emphasis>
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</taxonomicName>
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:
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<specimenCount id="E6F3FD42333D002ACB65F8BECE0C476D" box="[773,794,1831,1857]" pageId="14" pageNumber="45" type="female">♀</specimenCount>
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||
, labeled “
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||
<specimenCount id="E6F3FD42333D002ACBEBF8BFCE88476C" box="[907,926,1830,1856]" pageId="14" pageNumber="45" type="female">♀</specimenCount>
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||
”, “australasiae m. in Nov.
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<collectingCountry id="88E2765B333D002ACCA5F8BFC834476C" box="[1221,1314,1830,1856]" name="Netherlands" pageId="14" pageNumber="45">Holland</collectingCountry>
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||
” [
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||
<location id="F52A6010333D002ACD57F8BFC88A476C" LSID="urn:lsid:plazi:treatment:785C87DD333D002BC8F7F9EEC9CC43AD:F52A6010333D002ACD57F8BFC88A476C" box="[1335,1436,1830,1856]" country="Netherlands" name="Dejean's" pageId="14" pageNumber="45">Dejean’s</location>
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handwriting], “TYPE”, and “
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<taxonomicName id="37F54D48333D002AC9BCF8D3CE0A4748" authority="Dej." authorityName="Dej." box="[476,796,1866,1892]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="australasiae">Hypharpax australasiae Dej.</taxonomicName>
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||
examined
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<date id="844B100B333D002ACBF6F8D3C9034748" box="[918,1045,1866,1892]" pageId="14" pageNumber="45" value="1978-09-26">
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||
<collectingDate id="940FE9E3333D002ACBF6F8D3C9034748" box="[918,1045,1866,1892]" pageId="14" pageNumber="45" value="1978-09-26">26.IX.1978</collectingDate>
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||
</date>
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||
<collectorName id="5D00531D333D002ACC7CF8D2C98B4748" box="[1052,1181,1866,1893]" pageId="14" pageNumber="45">P.M. Tohus</collectorName>
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||
” (
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<collectionCode id="96E4AE0E333D002ACCD5F8D3C8074748" box="[1205,1297,1866,1892]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="14" pageNumber="45">MNHN</collectionCode>
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||
).
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||
</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B8EF6540333D002BC8A7F8F7C9CC43AD" lastPageId="15" lastPageNumber="46" pageId="14" pageNumber="45" type="discussion">
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||
<paragraph id="F04A36CB333D002BC8A7F8F7CE3C4131" blockId="14.[151,1437,1830,2037]" lastBlockId="15.[151,1437,151,897]" lastPageId="15" lastPageNumber="46" pageId="14" pageNumber="45">
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<emphasis id="C281EAD9333D002AC8A7F8F7CC2D47A4" bold="true" box="[199,315,1902,1928]" pageId="14" pageNumber="45">Remarks.</emphasis>
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The description of
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<taxonomicName id="37F54D48333D002ACA7EF8F6CFDA47A4" box="[542,716,1902,1928]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="species" species="australasiae">
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<emphasis id="C281EAD9333D002ACA7EF8F6CFDA47A4" box="[542,716,1902,1928]" italics="true" pageId="14" pageNumber="45">H. australasiae</emphasis>
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</taxonomicName>
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is based on
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<specimenCount id="E6F3FD42333D002ACB3FF8F6CECA47A4" box="[863,988,1902,1929]" pageId="14" pageNumber="45" type="female">one female</specimenCount>
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from
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<collectingCountry id="88E2765B333D002ACC43F8F7C99C47A4" box="[1059,1162,1902,1928]" name="Australia" pageId="14" pageNumber="45">Australia</collectingCountry>
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(“
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<collectingCountry id="88E2765B333D002ACCC5F8F7C89E47A5" box="[1189,1416,1902,1929]" name="Australia" pageId="14" pageNumber="45">Nouvelle-Hollande</collectingCountry>
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”). The systematic position of this species has been changed repeatedly.
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<bibRefCitation id="94644B3A333D002ACBEDF80AC90D4780" author="Bates, H. W." box="[909,1051,1938,1965]" pageId="14" pageNumber="45" pagination="233 - 246" refId="ref15164" refString="Bates, H. W. (1874) On the geodephagous Coleoptera of New Zealand. Annals and Magazine of Natural History, Series 4, 13, 233 - 246 + 270 - 277. https: // doi. org / 10.1080 / 00222937408680859" type="journal article" year="1874">Bates (1874)</bibRefCitation>
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referred it to the genus
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<taxonomicName id="37F54D48333D002ACD7EF80BCC5147FD" authority="MacLeay, 1825" authorityName="MacLeay" authorityYear="1825" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="C281EAD9333D002ACD7EF80BC88A4780" box="[1310,1436,1938,1964]" italics="true" pageId="14" pageNumber="45">Hypharpax</emphasis>
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MacLeay, 1825
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</taxonomicName>
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of the subtribe
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<taxonomicName id="37F54D48333D002AC99FF82FCFBF47FC" baseAuthorityName="Noonan" baseAuthorityYear="1973" box="[511,681,1974,2000]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="subTribe" subTribe="Anisodactylina">Anisodactylina</taxonomicName>
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and the subsequent authors also treated this species as a member of this subtribe but belonging to differing genera:
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<taxonomicName id="37F54D48333D002ACABAF843C95547D8" authority="Chaudoir, 1843" authorityName="Chaudoir" authorityYear="1843" box="[730,1091,2010,2036]" class="Insecta" family="Carabidae" genus="Diaphoromerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="45" phylum="Arthropoda" rank="genus">
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<emphasis id="C281EAD9333D002ACABAF843CE9C47D8" box="[730,906,2010,2036]" italics="true" pageId="14" pageNumber="45">Diaphoromerus</emphasis>
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Chaudoir, 1843
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</taxonomicName>
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||
(
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<bibRefCitation id="94644B3A333D002ACC34F843C81447D8" author="Chaudoir, M. de" box="[1108,1282,2010,2036]" pageId="14" pageNumber="45" pagination="475 - 517" refId="ref15468" refString="Chaudoir, M. de (1878) Les harpaliens de l'Australie d'apres la collection de M. le Comte de Castelnau et la mienne. Annali del Museo Civico di Storia Naturale di Genova, 12, 475 - 517." type="journal article" year="1878">Chaudoir 1878</bibRefCitation>
|
||
,
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||
<bibRefCitation id="94644B3A333D002ACD6FF843C88647D8" author="Csiki, E." box="[1295,1424,2010,2036]" pageId="14" pageNumber="45" pagination="1023 - 1278" refId="ref15628" refString="Csiki, E. (1932) Carabidae: Harpalinae VI. Pars 121. In: Junk, W. & Schenkling, S. (Eds.), Coleopterorum Catalogus. Junk, Berlin, pp. 1023 - 1278." type="book chapter" year="1932">Csiki 1932</bibRefCitation>
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||
),
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<taxonomicName id="37F54D48333C002BC8F7FF0ECF2C409D" authority="Perty, 1830" authorityName="Perty" authorityYear="1830" box="[151,570,151,177]" class="Insecta" family="Carabidae" genus="Notiobia" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">
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||
<emphasis id="C281EAD9333C002BC8F7FF0ECDEF409D" box="[151,249,151,177]" italics="true" pageId="15" pageNumber="46">Notiobia</emphasis>
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||
Perty, 1830 (
|
||
<bibRefCitation id="94644B3A333C002BC9F3FF0ECF27409D" author="Noonan, G. R." box="[403,561,151,177]" pageId="15" pageNumber="46" pagination="266 - 480" refId="ref16823" refString="Noonan, G. R. (1973) The Anisodactylines (Insecta: Coleoptera: Carabinae: Harpalini): classification, evolution, and zoogeography. Quaestiones Entomologicae, 9, 266 - 480." type="journal article" year="1973">Noonan 1973</bibRefCitation>
|
||
)
|
||
</taxonomicName>
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, and
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<taxonomicName id="37F54D48333C002BCA19FF0ECFE1409D" authorityName="MacLeay" authorityYear="1825" box="[633,759,151,177]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">
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||
<emphasis id="C281EAD9333C002BCA19FF0ECFE1409D" box="[633,759,151,177]" italics="true" pageId="15" pageNumber="46">Hypharpax</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="94644B3A333C002BCB67FF0ECE84409D" author="Broun, T." box="[775,914,151,177]" pageId="15" pageNumber="46" refId="ref15309" refString="Broun, T. (1880) Manual of the New Zealand Coleoptera. Part I. Government Printer, Wellington, XIX + 651 + VIII pp. https: // doi. org / 10.5962 / bhl. title. 32505" type="book" year="1880">Broun 1880</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="94644B3A333C002BCBFFFF0EC95A409D" author="Thomson, G. M." box="[927,1100,151,177]" pageId="15" pageNumber="46" refId="ref17051" refString="Thomson, G. M. (1922) The naturalisation of animals and plants in New Zealand. Cambridge University Press, London, 607 pp. https: // doi. org / 10.5962 / bhl. title. 28093" type="book" year="1922">Thomson 1922</bibRefCitation>
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||
,
|
||
<bibRefCitation id="94644B3A333C002BCC39FF0EC9E2409D" author="Hudson, G. V." box="[1113,1268,151,177]" pageId="15" pageNumber="46" pagination="353 - 399" refId="ref15856" refString="Hudson, G. V. (1923) An index of New Zealand beetles. Transactions and Proceedings of the New Zealand Institute, 54, 353 - 399. [1922]" type="journal article" year="1923">Hudson 1923</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="94644B3A333C002BCD60FF0EC88E409D" author="Pilgrim, R. L. C." box="[1280,1432,151,177]" pageId="15" pageNumber="46" pagination="837 - 847" refId="ref16903" refString="Pilgrim, R. L. C. (1963) Anisodactylus binotatus Fabr., a carabid beetle new to New Zealand, and a review of the exotic carabid fauna. Pacific Insects, 5, 837 - 847." type="journal article" year="1963">Pilgrim 1963</bibRefCitation>
|
||
, Moore in
|
||
<bibRefCitation id="94644B3A333C002BC969FF22CCCF40F9" author="Moore, B. P. & Weir, T. A. & Pyke, J. E." box="[265,473,187,213]" pageId="15" pageNumber="46" refId="ref16710" refString="Moore, B. P., Weir, T. A. & Pyke, J. E. (1987) Carabidae. In: Lawrence, J. F., Weir, T. A. & Pyke, J. E. (Eds.), Coleoptera: Archostemata, Myxophaga and Adephaga. Zoological catalogue of Australia. Vol. 4. Australian Government Publishing Service, Canberra, pp. 20 - 320." type="book" year="1987">
|
||
Moore
|
||
<emphasis id="C281EAD9333C002BC93CFF22CC8040F9" box="[348,406,187,213]" italics="true" pageId="15" pageNumber="46">et al.</emphasis>
|
||
1987
|
||
</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="94644B3A333C002BC986FF22CF8A40F9" author="Townsend, J. I." box="[486,668,187,213]" pageId="15" pageNumber="46" pagination="7 - 13" refId="ref17093" refString="Townsend, J. I. (1994) Carabidae of the Manawatu - Horowhenua lowlands as revealed by collections from coastal flood debris. New Zealand Entomologist, 17, 7 - 13. https: // doi. org / 10.1080 / 00779962.1994.9721978" type="journal article" year="1994">Townsend 1994</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="94644B3A333C002BCAC9FF22CEE540F9" author="Larochelle, A & Lariviere, M. - C." box="[681,1011,187,213]" pageId="15" pageNumber="46" refId="ref16461" refString="Larochelle, A, & Lariviere, M. - C. (2001) Carabidae (Insecta: Coleoptera): catalogue. Fauna of New Zealand. Vol. 43. Manaaki Whenua Press, Linkoln, Canburry, 281 pp." type="book" year="2001">Larochelle & Larivière 2001</bibRefCitation>
|
||
), until
|
||
<bibRefCitation id="94644B3A333C002BCC24FF22C88A40F9" author="Larochelle, A. & Lariviere, M. - C." box="[1092,1436,186,213]" pageId="15" pageNumber="46" refId="ref16505" refString="Larochelle, A. & Lariviere, M. - C. (2005) Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae). Fauna of New Zealand. Vol. 53. Manaaki Whenua Press, Linkoln, Canburry, 160 pp." type="book" year="2005">Larochelle & Larivière (2005)</bibRefCitation>
|
||
incorporated it again in the genus
|
||
<taxonomicName id="37F54D48333C002BCA76FF46CF9640D5" authorityName="Latreille" authorityYear="1802" box="[534,640,223,249]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C281EAD9333C002BCA76FF46CF9640D5" box="[534,640,223,249]" italics="true" pageId="15" pageNumber="46">Harpalus</emphasis>
|
||
</taxonomicName>
|
||
based mainly on the biseriate vestiture of male pro- and mesotarsi and the aedeagus with apical orifice strongly shifted to the left.
|
||
</paragraph>
|
||
<paragraph id="F04A36CB333C002BC8A7FEBEC9CC43AD" blockId="15.[151,1437,151,897]" pageId="15" pageNumber="46">
|
||
I examined the
|
||
<typeStatus id="2F4E8869333C002BC91BFEBECCCB416D" box="[379,477,295,321]" pageId="15" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="37F54D48333C002BCA67FEBECFA3416D" box="[519,693,295,321]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="C281EAD9333C002BCA67FEBECFA3416D" box="[519,693,295,321]" italics="true" pageId="15" pageNumber="46">H. australasiae</emphasis>
|
||
</taxonomicName>
|
||
at MNHN (
|
||
<figureCitation id="68CE2A4E333C002BCB24FEBECE8F416D" box="[836,921,295,321]" captionStart="FIGURES 29–32" captionStartId="13.[151,264,1925,1949]" captionTargetBox="[180,1402,189,1896]" captionTargetId="figure-16@13.[172,1415,181,1901]" captionTargetPageId="13" captionText="FIGURES 29–32. Harpalus spp., habitus. 29–31, H. fuscoaeneus Dejean, 1929 (29, S Africa; 30, lectotype; 31, holotype of H. nanus Dejean, 1829); 32, H. australasiae Dejean, 1829 (holotype). Scale bars = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5222947" httpUri="https://zenodo.org/record/5222947/files/figure.png" pageId="15" pageNumber="46">Fig. 32</figureCitation>
|
||
), which well corresponds to
|
||
<bibRefCitation id="94644B3A333C002BCC86FEBEC88A416D" author="Dejean, P. F. M. A." box="[1254,1436,295,321]" pageId="15" pageNumber="46" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean’s (1829)</bibRefCitation>
|
||
original description. It is indeed a female of
|
||
<emphasis id="C281EAD9333C002BCAEEFED2CE044149" box="[654,786,331,357]" italics="true" pageId="15" pageNumber="46">
|
||
<taxonomicName id="37F54D48333C002BCAEEFED2CFE24149" authorityName="Latreille" authorityYear="1802" box="[654,756,331,357]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">Harpalus</taxonomicName>
|
||
—
|
||
</emphasis>
|
||
like all
|
||
<taxonomicName id="37F54D48333C002BCB07FED2CEC74149" authorityName="Latreille" authorityYear="1802" box="[871,977,331,357]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C281EAD9333C002BCB07FED2CEC74149" box="[871,977,331,357]" italics="true" pageId="15" pageNumber="46">Harpalus</emphasis>
|
||
</taxonomicName>
|
||
, its paraglossae are setose as opposed to the glabrous paraglossae of
|
||
<taxonomicName id="37F54D48333C002BC9BCFEF6CF4C41A5" authorityName="MacLeay" authorityYear="1825" box="[476,602,367,393]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C281EAD9333C002BC9BCFEF6CF4C41A5" box="[476,602,367,393]" italics="true" pageId="15" pageNumber="46">Hypharpax</emphasis>
|
||
</taxonomicName>
|
||
and other Australian members of
|
||
<taxonomicName id="37F54D48333C002BCB87FEF6C98241A5" baseAuthorityName="Noonan" baseAuthorityYear="1973" box="[999,1172,367,393]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="subTribe" subTribe="Anisodactylina">Anisodactylina</taxonomicName>
|
||
. The characters of this female, however, contradict the re-description and illustrations by
|
||
<bibRefCitation id="94644B3A333C002BCBE3FE0AC9CD4181" author="Larochelle, A. & Lariviere, M. - C." box="[899,1243,402,429]" pageId="15" pageNumber="46" refId="ref16505" refString="Larochelle, A. & Lariviere, M. - C. (2005) Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae). Fauna of New Zealand. Vol. 53. Manaaki Whenua Press, Linkoln, Canburry, 160 pp." type="book" year="2005">Larochelle & Larivière (2005)</bibRefCitation>
|
||
. In the examined
|
||
<typeStatus id="2F4E8869333C002BC8F7FE2ECDEA41FD" box="[151,252,439,465]" pageId="15" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
, the pronotum is more or less equally narrowed apically and basally, with basal angles clearly obtusangular, rounded at apex, and the elytra are more oval; the proportions: HWmax/PWmax 0.63 HWmin/PWmax 0.52; HWmax/HWmin 1.22; PWmax/PL 1.43; PWmax/PWmin 1.24; EL/EW 1.44; EL/PL 2.58; EW/PWmax 1.26; the body length
|
||
<quantity id="370D9B2E333C002BC943FDBACC6B4211" box="[291,381,547,573]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="15" pageNumber="46" unit="mm" value="6.2">6.2 mm</quantity>
|
||
. In these and other characters, including coloration and metafemur with three setae along posterior margin, this female is very similar to specimens of
|
||
<taxonomicName id="37F54D48333C002BCB75FDDECEA9424D" authorityName="Dejean" authorityYear="1929" box="[789,959,583,609]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333C002BCB75FDDECEA9424D" box="[789,959,583,609]" italics="true" pageId="15" pageNumber="46">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37F54D48333C002BCB95FDDEC97F424C" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[1013,1129,583,608]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="asemus">
|
||
<emphasis id="C281EAD9333C002BCB95FDDEC97F424C" box="[1013,1129,583,608]" italics="true" pageId="15" pageNumber="46">H. asemus</emphasis>
|
||
</taxonomicName>
|
||
. Such elytral features of
|
||
<taxonomicName id="37F54D48333C002BCDE1FDDECC3742A9" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="C281EAD9333C002BCDE1FDDECC3742A9" italics="true" pageId="15" pageNumber="46">H. australasiae</emphasis>
|
||
</taxonomicName>
|
||
as comparatively long parascutellar striole and absence of discal pore on interval 3 occasionally occur in both
|
||
<taxonomicName id="37F54D48333C002BC88CFD16CC814285" authorityName="Dejean" authorityYear="1929" box="[236,407,655,681]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="fuscoaeneus">
|
||
<emphasis id="C281EAD9333C002BC88CFD16CC814285" box="[236,407,655,681]" italics="true" pageId="15" pageNumber="46">H. fuscoaeneus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="37F54D48333C002BC9ACFD16CF564284" authorityName="Basilew. P. Basilewsky" authorityYear="1947" box="[460,576,655,680]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="asemus">
|
||
<emphasis id="C281EAD9333C002BC9ACFD16CF564284" box="[460,576,655,680]" italics="true" pageId="15" pageNumber="46">H. asemus</emphasis>
|
||
</taxonomicName>
|
||
. The status of
|
||
<taxonomicName id="37F54D48333C002BCA84FD16CE994285" box="[740,911,655,681]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="C281EAD9333C002BCA84FD16CE994285" box="[740,911,655,681]" italics="true" pageId="15" pageNumber="46">H. australasiae</emphasis>
|
||
</taxonomicName>
|
||
needs further study, but it is quite possible that this taxon is conspecific with one of these two species and its
|
||
<typeStatus id="2F4E8869333C002BCB22FD2ACEB242E1" box="[834,932,691,717]" pageId="15" pageNumber="46" type="holotype">holotype</typeStatus>
|
||
was either mislabeled or brought to
|
||
<collectingCountry id="88E2765B333C002BCD56FD2AC88B42E1" box="[1334,1437,691,717]" name="Australia" pageId="15" pageNumber="46">Australia</collectingCountry>
|
||
from Africa. Unfortunately, the more accurate determination based only on
|
||
<specimenCount id="E6F3FD42333C002BCB93FD4EC96642DD" box="[1011,1136,727,753]" pageId="15" pageNumber="46" type="female">one female</specimenCount>
|
||
is now impossible. As for
|
||
<taxonomicName id="37F54D48333C002BC8F7FD62CFF54339" authority="sensu Larochelle & Lariviere (2005)" authorityName="sensu Larochelle & Lariviere" authorityYear="2005" box="[151,739,762,789]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="C281EAD9333C002BC8F7FD62CC554339" box="[151,323,763,789]" italics="true" pageId="15" pageNumber="46">H. australasiae</emphasis>
|
||
sensu
|
||
<bibRefCitation id="94644B3A333C002BC9EFFD62CFF54339" author="Larochelle, A. & Lariviere, M. - C." box="[399,739,762,789]" pageId="15" pageNumber="46" refId="ref16505" refString="Larochelle, A. & Lariviere, M. - C. (2005) Harpalini (Insecta: Coleoptera: Carabidae: Harpalinae). Fauna of New Zealand. Vol. 53. Manaaki Whenua Press, Linkoln, Canburry, 160 pp." type="book" year="2005">Larochelle & Larivière (2005)</bibRefCitation>
|
||
</taxonomicName>
|
||
, I think that their interpretation is actually based on specimens of
|
||
<taxonomicName id="37F54D48333C002BC8D6FC86CCCF4315" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" box="[182,473,799,825]" class="Insecta" family="Carabidae" genus="Harpalus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="parvulus">
|
||
<emphasis id="C281EAD9333C002BC8D6FC86CC2D4315" box="[182,315,799,825]" italics="true" pageId="15" pageNumber="46">H. parvulus</emphasis>
|
||
<bibRefCitation id="94644B3A333C002BC923FC86CCCF4315" author="Dejean, P. F. M. A." box="[323,473,799,825]" pageId="15" pageNumber="46" refId="ref15672" refString="Dejean, P. F. M. A. (1829) Species general des coleopteres, de la collection de M. le Comte Dejean. Tome quatrieme. Mequignon- Marvis, Paris, vii + 520 pp." type="book" year="1829">Dejean, 1829</bibRefCitation>
|
||
</taxonomicName>
|
||
(see below). My work with the museum collections revealed that some of the former records of
|
||
<taxonomicName id="37F54D48333C002BC971FCDACCA84371" authorityName="sensu Larochelle & Lariviere" authorityYear="2005" box="[273,446,835,861]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="australasiae">
|
||
<emphasis id="C281EAD9333C002BC971FCDACCA84371" box="[273,446,835,861]" italics="true" pageId="15" pageNumber="46">H. australasiae</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="88E2765B333C002BCA64FCDACF7D4371" box="[516,619,835,861]" name="Australia" pageId="15" pageNumber="46">Australia</collectingCountry>
|
||
and
|
||
<collectingCountry id="88E2765B333C002BCAC3FCDACE2C4371" box="[675,826,835,861]" name="New Zealand" pageId="15" pageNumber="46">New Zealand</collectingCountry>
|
||
should apparently be referred to
|
||
<taxonomicName id="37F54D48333C002BCCD4FCDACC2943AD" authority="(Dejean, 1929)" baseAuthorityName="Dejean" baseAuthorityYear="1929" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="species" species="australis">
|
||
<emphasis id="C281EAD9333C002BCCD4FCDAC88A4371" box="[1204,1436,835,861]" italics="true" pageId="15" pageNumber="46">Hypharpax australis</emphasis>
|
||
(Dejean, 1929)
|
||
</taxonomicName>
|
||
and some other species of this genus. The revision of
|
||
<taxonomicName id="37F54D48333C002BCBFFFCFEC90B43AD" authorityName="MacLeay" authorityYear="1825" box="[927,1053,871,897]" class="Insecta" family="Carabidae" genus="Hypharpax" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="46" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="C281EAD9333C002BCBFFFCFEC90B43AD" box="[927,1053,871,897]" italics="true" pageId="15" pageNumber="46">Hypharpax</emphasis>
|
||
</taxonomicName>
|
||
is much needed.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |