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<document id="232DFE5BFEE941F01167E4CBE8878D4A" ID-DOI="10.26028/cybium/2016-402-003" ID-ISSN="2101-0315" ID-Zenodo-Dep="13493776" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="guilherme" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="guilherme" checkinTime="1723045288427" checkinUser="felipe" docAuthor="Meunier, François J., Eustache, René-Paul, Dutheil, Didier &amp; Cavin, Lionel" docDate="2016" docId="76442E58922B69271301FAA4CA7F8955" docLanguage="en" docName="Cybium.40.2.121-132.pdf" docOrigin="Cybium 40 (2)" docSource="http://dx.doi.org/10.26028/cybium/2016-402-003" docStyle="DocumentStyle:A45444AC9C9E9900C6D8D41D5B4A30BB.1:Cybium.2013-2018.journal_article.type2" docStyleId="A45444AC9C9E9900C6D8D41D5B4A30BB" docStyleName="Cybium.2013-2018.journal_article.type2" docStyleVersion="1" docTitle="Adrianaichthys Meunier, Eustache, Dutheil &amp; Cavin, 2016, gen. nov." docType="treatment" docVersion="3" lastPageNumber="122" masterDocId="8A7D5620922A6925170AFFFBCB5A8A30" masterDocTitle="Histology of ganoid scales from the early Late Cretaceous of the Kem Kem beds, SE Morocco: systematic and evolutionary implications" masterLastPageNumber="132" masterPageNumber="121" pageNumber="122" updateTime="1724942774381" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="8B074911F874F356F042CB7406C92E78">Histology of ganoid scales from the early Late Cretaceous of the Kem Kem beds, SE Morocco: systematic and evolutionary implications</mods:title>
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<mods:namePart id="1AFF0B43BEF28D53BD83D0F6FC10A7EC">Meunier, François J.</mods:namePart>
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<paragraph id="FE529F4E922B69241301FAA4CE478F48" blockId="1.[1035,1309,1375,1400]" box="[1035,1309,1375,1400]" pageId="1" pageNumber="122">
<heading id="A51A2822922B69241301FAA4CE478F48" bold="true" box="[1035,1309,1375,1400]" centered="true" fontSize="9" level="2" pageId="1" pageNumber="122" reason="2">
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<taxonomicName id="39EDE4CD922B69241301FAA4CFEE8F47" authorityName="Meunier &amp; Eustache &amp; Dutheil &amp; Cavin" authorityYear="2016" box="[1035,1204,1375,1399]" class="Actinopterygii" family="Semionotidae" genus="Adrianaichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="1" pageNumber="122" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="CC99435C922B69241301FAA4CFEE8F47" bold="true" box="[1035,1204,1375,1399]" italics="true" pageId="1" pageNumber="122">Adrianaichthys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7AAFE27922B692413B3FA9BCE478F48" box="[1209,1309,1376,1400]" pageId="1" pageNumber="122" rank="genus">gen. nov.</taxonomicNameLabel>
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<paragraph id="FE529F4E922B69241472FA5BCF8F8FE8" blockId="1.[850,1494,1439,1944]" pageId="1" pageNumber="122">
<emphasis id="CC99435C922B69241472FA5BCF578F87" box="[888,1037,1440,1463]" italics="true" pageId="1" pageNumber="122">
<typeStatus id="215621EC922B69241472FA5BC8F48F87" box="[888,942,1440,1463]" pageId="1" pageNumber="122">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="39EDE4CD922B6924133CFA64CF8F8FE8" authority="(Forey, Lopez-Arbarello &amp; MacLeod, 2011)" baseAuthorityName="Forey, Lopez-Arbarello &amp; MacLeod" baseAuthorityYear="2011" class="Actinopterygii" family="Semionotidae" genus="Adrianaichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="1" pageNumber="122" phylum="Chordata" rank="species" species="pankowskii">
<emphasis id="CC99435C922B6924133CFA64CE2D8F87" box="[1078,1399,1439,1463]" italics="true" pageId="1" pageNumber="122">Adrianaichthys pankowskii</emphasis>
(
<bibRefCitation id="9A7CE2BF922B69241283FA5BCF948FE8" author="FOREY P. L. &amp; LOPEZ-ARBARELLO A. &amp; MACLEOD N." pageId="1" pageNumber="122" pagination="1 - 12" refId="ref7814" refString="FOREY P. L., LOPEZ-ARBARELLO A. &amp; MACLEOD N., 2011. - A new species of Lepidotes (Actinopterygii: Semiontiformes) from the Cenomanian (Upper Cretaceous) of Morocco. Palaeont. Electron., 14 (1); 7 A: 1 - 12." type="journal article" year="2011">Forey, López-Arbarello &amp; MacLeod, 2011</bibRefCitation>
)
</taxonomicName>
</paragraph>
<paragraph id="FE529F4E922B69241472FA1BCF458C28" blockId="1.[850,1494,1439,1944]" pageId="1" pageNumber="122">
<emphasis id="CC99435C922B69241472FA1BC8BE8FC7" box="[888,996,1504,1527]" italics="true" pageId="1" pageNumber="122">Diagnosis</emphasis>
. As for the
<typeStatus id="215621EC922B6924137EFA1BCFF88FC8" box="[1140,1186,1504,1528]" pageId="1" pageNumber="122">type</typeStatus>
and only known species, see
<bibRefCitation id="9A7CE2BF922B69241458F9FBCF418C28" author="FOREY P. L. &amp; LOPEZ-ARBARELLO A. &amp; MACLEOD N." box="[850,1051,1536,1560]" pageId="1" pageNumber="122" pagination="1 - 12" refId="ref7814" refString="FOREY P. L., LOPEZ-ARBARELLO A. &amp; MACLEOD N., 2011. - A new species of Lepidotes (Actinopterygii: Semiontiformes) from the Cenomanian (Upper Cretaceous) of Morocco. Palaeont. Electron., 14 (1); 7 A: 1 - 12." type="journal article" year="2011">
Forey
<emphasis id="CC99435C922B6924149DF9FBC89F8C28" box="[919,965,1536,1560]" italics="true" pageId="1" pageNumber="122">et al</emphasis>
. (2011)
</bibRefCitation>
.
</paragraph>
<paragraph id="FE529F4E922B69241472F9DBCE0B8C88" blockId="1.[850,1494,1439,1944]" pageId="1" pageNumber="122">
<emphasis id="CC99435C922B69241472F9DBC8AA8C08" box="[888,1008,1568,1592]" italics="true" pageId="1" pageNumber="122">Etymology</emphasis>
. In honour of Adriana López-Arbarello, palaeoichthyologist at the Bayerische Staatssammlung für Paläontologie und Geologie, München, who undertook the difficult task to untangle the phylogeny and systematics of ginglymodian fish, and
<emphasis id="CC99435C922B6924135FF95BCFF38C88" box="[1109,1193,1696,1720]" italics="true" pageId="1" pageNumber="122">ichthys</emphasis>
(fish in Greek).
</paragraph>
<paragraph id="FE529F4E922B69271472F93BCA7F8955" blockId="1.[850,1494,1439,1944]" lastBlockId="2.[170,813,255,869]" lastPageId="2" lastPageNumber="123" pageId="1" pageNumber="122">
<emphasis id="CC99435C922B69241472F93BC88D8CE8" box="[888,983,1728,1752]" italics="true" pageId="1" pageNumber="122">Remarks</emphasis>
. In their original description of
<emphasis id="CC99435C922B69241246F93BCE0C8CE8" box="[1356,1366,1728,1752]" italics="true" pageId="1" pageNumber="122"></emphasis>
<taxonomicName id="39EDE4CD922B6924125CF93BCFFF8CC8" authority=", Forey et al. (2011)" authorityName="Forey" authorityYear="2011" class="Actinopterygii" family="Lepidotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="1" pageNumber="122" phylum="Chordata" rank="species" species="pankowskii">
<emphasis id="CC99435C922B6924125CF93BC8968CC8" italics="true" pageId="1" pageNumber="122">Lepidotes” pankowskii</emphasis>
,
<bibRefCitation id="9A7CE2BF922B692414D0F91BCFFF8CC8" author="FOREY P. L. &amp; LOPEZ-ARBARELLO A. &amp; MACLEOD N." box="[986,1189,1760,1784]" pageId="1" pageNumber="122" pagination="1 - 12" refId="ref7814" refString="FOREY P. L., LOPEZ-ARBARELLO A. &amp; MACLEOD N., 2011. - A new species of Lepidotes (Actinopterygii: Semiontiformes) from the Cenomanian (Upper Cretaceous) of Morocco. Palaeont. Electron., 14 (1); 7 A: 1 - 12." type="journal article" year="2011">
Forey
<emphasis id="CC99435C922B6924132AF91ACF0A8CC8" box="[1056,1104,1760,1784]" italics="true" pageId="1" pageNumber="122">et al</emphasis>
. (2011)
</bibRefCitation>
</taxonomicName>
pointed out the heterogeneous status of the genus “
<emphasis id="CC99435C922B69241351F8FBCF8E8D28" box="[1115,1236,1792,1816]" italics="true" pageId="1" pageNumber="122">
<taxonomicName id="39EDE4CD922B69241351F8FBCF908D28" box="[1115,1226,1792,1816]" class="Actinopterygii" family="Lepidotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="1" pageNumber="122" phylum="Chordata" rank="genus">Lepidotes</taxonomicName>
</emphasis>
, which was at that time in urgent need of revision.
<bibRefCitation id="9A7CE2BF922B69241380F8DBCEC48D08" author="LOPEZ-ARBARELLO A." box="[1162,1438,1824,1848]" pageId="1" pageNumber="122" refId="ref8622" refString="LOPEZ-ARBARELLO A., 2012. - Phylogenetic interrelationships of Ginglymodian fishes (Actinopterygii: neopterygii). PLoS ONE, 7 (7): e 39370." type="journal volume" year="2012">López-Arbarello (2012)</bibRefCitation>
, in a revision of several ginglymodian taxa, did not include this species in her phylogenetic analysis, but suggested that “
<taxonomicName id="39EDE4CD922B692412B9F89AC8908DA8" class="Actinopterygii" family="Lepidotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="1" pageNumber="122" phylum="Chordata" rank="species" species="pankowskii">
<emphasis id="CC99435C922B692412B9F89ACE938D48" box="[1459,1481,1889,1912]" italics="true" pageId="1" pageNumber="122">L.</emphasis>
<emphasis id="CC99435C922B69241458F87BC8908DA8" box="[850,970,1920,1944]" italics="true" pageId="1" pageNumber="122">pankowskii</emphasis>
</taxonomicName>
is most probably a lepisosteiform incertae genus. In subsequent articles dealing with this species, the genus
<taxonomicName id="39EDE4CD9228692717A0FEDBCA4C8B08" box="[170,278,288,312]" class="Actinopterygii" family="Lepidotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="2" pageNumber="123" phylum="Chordata" rank="genus">
<emphasis id="CC99435C9228692717A0FEDBCA4C8B08" box="[170,278,288,312]" italics="true" pageId="2" pageNumber="123">Lepidotes</emphasis>
</taxonomicName>
was generally quoted between inverted comma (
<bibRefCitation id="9A7CE2BF9228692717B8FEBAC9FF8B68" author="BERMUDEZ-ROCHAS D. D. &amp; POYATO-ARIZA F. J." box="[178,677,320,345]" pageId="2" pageNumber="123" pagination="265 - 285" refId="ref6787" refString="BERMUDEZ-ROCHAS D. D. &amp; POYATO-ARIZA F. J., 2015. - A new semionotiform actinopterygian fish from the Mesozoic of Spain and its phylogenetic implications. J. Syst. Palaeont., 13 (4): 265 - 285." type="journal article" year="2015">Bermúdez-Rochas and Poyato-Ariza, 2015</bibRefCitation>
;
<bibRefCitation id="9A7CE2BF9228692715B8FEBACA728B49" author="BENYOUCEF M. &amp; LANG E. &amp; CAVIN L. &amp; MEBARKI K. &amp; ADACI M. &amp; BENSALAH M." pageId="2" pageNumber="123" pagination="44 - 55" refId="ref6730" refString="BENYOUCEF M., LANG E., CAVIN L., MEBARKI K., ADACI M. &amp; BENSALAH M., 2015. - Overabundance of piscivorous dinosaurs (Theropoda: Spinosauridae) in the mid-Cretaceous of North Africa: The Algerian dilemma. Cretaceous Res., 55: 44 - 55." type="journal article" year="2015">
Benyoucef
<emphasis id="CC99435C9228692717A0FE99CB808B49" box="[170,218,353,377]" italics="true" pageId="2" pageNumber="123">et al</emphasis>
., 2015
</bibRefCitation>
) or preceded by a question mark (
<bibRefCitation id="9A7CE2BF9228692715AFFE9ACBBC8BAA" author="CAVIN L. &amp; BOUDAD L. &amp; TONG H. &amp; LANG E. &amp; TABOUELLE J. &amp; VULLO R." pageId="2" pageNumber="123" refId="ref7289" refString="CAVIN L., BOUDAD L., TONG H., LANG E., TABOUELLE J. &amp; VULLO R., 2015. - Taxonomic composition and trophic structure of the continental bony fish assemblage from the Early Late Cretaceous of Southeastern Morocco. PLoS ONE, 10 (5): e 0125786." type="journal volume" year="2015">
Cavin
<emphasis id="CC99435C9228692715E4FE99C8448B49" box="[750,798,353,377]" italics="true" pageId="2" pageNumber="123">et al</emphasis>
., 2015
</bibRefCitation>
). In order to clarify this situation, a new genus name is coined. Among other characters,
<taxonomicName id="39EDE4CD922869271528FE58C99F8B8B" authorityName="Meunier &amp; Eustache &amp; Dutheil &amp; Cavin" authorityYear="2016" box="[546,709,419,443]" class="Actinopterygii" family="Semionotidae" genus="Adrianaichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="2" pageNumber="123" phylum="Chordata" rank="genus" status="gen. nov.">
<emphasis id="CC99435C922869271528FE58C99F8B8B" box="[546,709,419,443]" italics="true" pageId="2" pageNumber="123">Adrianaichthys</emphasis>
</taxonomicName>
<taxonomicNameLabel id="D7AAFE279228692715C6FE58C8778B8A" box="[716,813,419,443]" pageId="2" pageNumber="123" rank="genus">gen. nov.</taxonomicNameLabel>
differs from
<taxonomicName id="39EDE4CD922869271625FE3FCACD8BEC" box="[303,407,452,476]" class="Actinopterygii" family="Lepidotidae" genus="Lepidotes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="2" pageNumber="123" phylum="Chordata" rank="genus">
<emphasis id="CC99435C922869271625FE3FCACD8BEC" box="[303,407,452,476]" italics="true" pageId="2" pageNumber="123">Lepidotes</emphasis>
</taxonomicName>
as defined by
<bibRefCitation id="9A7CE2BF922869271524FE3FC8768BEC" author="LOPEZ-ARBARELLO A." box="[558,812,452,476]" pageId="2" pageNumber="123" refId="ref8622" refString="LOPEZ-ARBARELLO A., 2012. - Phylogenetic interrelationships of Ginglymodian fishes (Actinopterygii: neopterygii). PLoS ONE, 7 (7): e 39370." type="journal volume" year="2012">López-Arbarello (2012)</bibRefCitation>
by its first anterior infraorbital bone not deeper than the more posterior anterior infraorbitals, three pairs of extrascapulars (versus one) and eight suborbitals, four of which lying in front of the orbit. According to
<bibRefCitation id="9A7CE2BF922869271515FDBCC873886F" author="LOPEZ-ARBARELLO A." box="[543,809,583,607]" pageId="2" pageNumber="123" refId="ref8622" refString="LOPEZ-ARBARELLO A., 2012. - Phylogenetic interrelationships of Ginglymodian fishes (Actinopterygii: neopterygii). PLoS ONE, 7 (7): e 39370." type="journal volume" year="2012">López-Arbarello (2012)</bibRefCitation>
,
<taxonomicName id="39EDE4CD9228692717A0FD93CA8B88B0" baseAuthorityName="Forey, Lopez-Arbarello &amp; MacLeod" baseAuthorityYear="2011" box="[170,465,616,640]" class="Actinopterygii" family="Semionotidae" genus="Adrianaichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="2" pageNumber="123" phylum="Chordata" rank="species" species="pankowskii">
<emphasis id="CC99435C9228692717A0FD93CA8B88B0" box="[170,465,616,640]" italics="true" pageId="2" pageNumber="123">Adrianaichthys pankowskii</emphasis>
</taxonomicName>
shows characters of
<taxonomicName id="39EDE4CD9228692715BEFD93CBAD8890" authorityName="Hay" authorityYear="1929" class="Actinopterygii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="2" pageNumber="123" phylum="Chordata" rank="order">Lepisosteiformes</taxonomicName>
, such as scales with rostro-caudal articulation through anterior dorsal and ventral processes, co-ossified vomers, three or more pairs of extrascapulars, closed circumorbital bone ring, and large supraorbital bone longest ventrally, contacting more than one infraorbital bone. We include here
<taxonomicName id="39EDE4CD92286927141FFCF7CA7C8974" baseAuthorityName="Forey, Lopez-Arbarello &amp; MacLeod" baseAuthorityYear="2011" class="Actinopterygii" family="Semionotidae" genus="Adrianaichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="2" pageNumber="123" phylum="Chordata" rank="species" species="pankowskii">
<emphasis id="CC99435C92286927141FFCF7CA7C8974" italics="true" pageId="2" pageNumber="123">A. pankowskii</emphasis>
</taxonomicName>
in the
<taxonomicName id="39EDE4CD92286927167EFCD7C96D8974" authorityName="Hay" authorityYear="1929" box="[372,567,812,836]" class="Actinopterygii" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepisosteiformes" pageId="2" pageNumber="123" phylum="Chordata" rank="order">Lepisosteiformes</taxonomicName>
, pending a revision of the species.
</paragraph>
</subSubSection>
</treatment>
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