231 lines
28 KiB
XML
231 lines
28 KiB
XML
<document id="EC397F90DFD7050DC9CBD2F65645F543" ID-DOI="10.11646/zootaxa.3846.4.1" ID-GBIF-Dataset="317a43cd-c408-4fde-8374-357fbb9a0d53" ID-ISSN="1175-5326" ID-Zenodo-Dep="250495" ID-ZooBank="0DD5A603-D65F-4976-BBE9-94DA7110053F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467125281" checkinUser="plazi" docAuthor="Jacobsen, Niels H. G., Kuhn, Arianna L., Jackman, Todd R. & Bauer, Aaron M." docDate="2014" docId="744387D5B12FA108EBADFF46E3B6FEC5" docLanguage="en" docName="zt03846p501.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afroedura leoloensis Jacobsen, Kuhn, Jackman & Bauer, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="487" masterDocId="887AFFADB10CA12CEB3AFFD1E06AFF85" masterDocTitle="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa" masterLastPageNumber="501" masterPageNumber="451" pageNumber="486" updateTime="1698305158428" updateUser="plazi">
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<mods:titleInfo id="642F2B3BEA7F702E57568ADC5D9FB4AB">
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<mods:title id="DA2BF7518987829760B03F751C2891D3">A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa</mods:title>
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<mods:roleTerm id="7B3EC77BF402A9CB764B955784AC6D41">Author</mods:roleTerm>
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<mods:namePart id="86191A885E69F7571C1D05709AA77CEC">Jacobsen, Niels H. G.</mods:namePart>
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<mods:namePart id="C296856C0D1D3017EC60B6AAB9DB2A25">Kuhn, Arianna L.</mods:namePart>
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<mods:roleTerm id="3B55558B0AB979BEF36F6F7AC7C25A1C">Author</mods:roleTerm>
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<mods:namePart id="D810BF02A98481B23FFFE7D6939C11F3">Jackman, Todd R.</mods:namePart>
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<mods:namePart id="876884D8A31388FA4A80E3D787C5B9A3">Bauer, Aaron M.</mods:namePart>
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<mods:typeOfResource id="CD5E382F9D439E795B39BF125878BE8E">text</mods:typeOfResource>
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<mods:title id="B08916991DFCDD9B3AA1DE6EBDFC725C">Zootaxa</mods:title>
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<mods:date id="CDD8BF416F6978C3F01CD8E13789CB85">2014</mods:date>
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<mods:detail id="3B01C4123A1E159BC426B073694524C0" type="volume">
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<mods:number id="4E04B29E2955DDFD159467F2547CC11B">3846</mods:number>
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<mods:identifier id="4F37700DF49C5141C2FEBD4C68DD5DBC" type="DOI">10.11646/zootaxa.3846.4.1</mods:identifier>
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<treatment id="744387D5B12FA108EBADFF46E3B6FEC5" ID-DOI="http://doi.org/10.5281/zenodo.5620595" ID-GBIF-Taxon="119514186" ID-Zenodo-Dep="5620595" LSID="urn:lsid:plazi:treatment:744387D5B12FA108EBADFF46E3B6FEC5" httpUri="http://treatment.plazi.org/id/744387D5B12FA108EBADFF46E3B6FEC5" lastPageId="36" lastPageNumber="487" pageId="35" pageNumber="486">
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<subSubSection id="B4F06548B12FA10FEBADFF46E161FF56" pageId="35" pageNumber="486" type="nomenclature">
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<paragraph id="FC5536C3B12FA10FEBADFF46E19DFF37" blockId="35.[151,503,151,211]" box="[151,503,151,178]" pageId="35" pageNumber="486">
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<heading id="A71D81AFB12FA10FEBADFF46E19DFF37" bold="true" box="[151,503,151,178]" fontSize="11" level="1" pageId="35" pageNumber="486" reason="1">
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<emphasis id="CE9EEAD1B12FA10FEBADFF46E19DFF37" bold="true" box="[151,503,151,178]" pageId="35" pageNumber="486">
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<taxonomicName id="3BEA4D40B12FA10FEBADFF46E1F9FF34" box="[151,403,151,177]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="486" phylum="Chordata" rank="species" species="leoloensis" status="sp. nov.">
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<emphasis id="CE9EEAD1B12FA10FEBADFF46E1F9FF34" bold="true" box="[151,403,151,177]" italics="true" pageId="35" pageNumber="486">Afroedura leoloensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D5AD57AAB12FA10FEAA0FF49E19DFF37" box="[410,503,152,178]" pageId="35" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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<paragraph id="FC5536C3B12FA10FEBADFF6BE161FF56" blockId="35.[151,503,151,211]" box="[151,267,186,211]" pageId="35" pageNumber="486">
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||
(
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<figureCitation id="64D12A46B12FA10FEBA5FF6BE09BFF56" box="[159,241,186,211]" captionStart="FIGURE 10" captionStartId="34.[151,250,1912,1934]" captionTargetBox="[221,1375,195,1892]" captionTargetId="figure@34.[221,1376,195,1893]" captionTargetPageId="34" captionText="FIGURE 10. New species in the Afroedura langi group. (A) Afroedura granitica sp. nov. from Farm Lillie, Limpopo Province, South Africa (24 ° 04 ’ S, 30 ° 51 ’ E) (Photo Stuart V. Nielsen). (B) Afroedura leoloensis sp. nov. from Farm Maandagshoek, Limpopo Province, South Africa (Photo N. H. G. Jacobsen). (C) Afroedura broadleyi sp. nov. from 22 ° 58 ’ 40 ” S, 29 ° 19 ’ 57 ” E, Limpopo Province, South Africa (Photo Marius Burger)." httpUri="https://zenodo.org/record/250505/files/figure.png" pageId="35" pageNumber="486">Fig. 10</figureCitation>
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B)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B4F06548B12FA10FEBADFED3E202FEB2" pageId="35" pageNumber="486" type="reference_group">
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<paragraph id="FC5536C3B12FA10FEBADFED3E202FEB2" blockId="35.[151,723,258,311]" pageId="35" pageNumber="486">
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<taxonomicName id="3BEA4D40B12FA10FEBADFED3E1F3FE9D" box="[151,409,258,280]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="486" phylum="Chordata" rank="subSpecies" species="pondolia" subSpecies="langi">
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<emphasis id="CE9EEAD1B12FA10FEBADFED3E1F3FE9D" box="[151,409,258,280]" italics="true" pageId="35" pageNumber="486">Afroedura pondolia langi</emphasis>
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</taxonomicName>
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(part)
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<bibRefCitation id="987B4B32B12FA10FEAE7FED3E20BFE9D" author="Visser" box="[477,609,258,280]" pageId="35" pageNumber="486" refString="Visser, J. 1984 a. Diksterte sukkel nie met blyplek. Akkedisse van Suider - Afrika 14. Landbouweekblad, 4 June 1984, 60 - 61, 63." type="journal article" year="1984" yearSuffix="a">Visser 1984a</bibRefCitation>
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(fig. p. 61)
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<taxonomicName id="3BEA4D40B12FA10FEBADFEF0E152FEB2" box="[151,312,289,311]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="486" phylum="Chordata" rank="species" species="langi">
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<emphasis id="CE9EEAD1B12FA10FEBADFEF0E152FEB2" box="[151,312,289,311]" italics="true" pageId="35" pageNumber="486">Afroedura langi</emphasis>
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</taxonomicName>
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'Leolo'
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<bibRefCitation id="987B4B32B12FA10FEAB3FEF0E246FEB2" author="Jacobsen" box="[393,556,289,311]" pageId="35" pageNumber="486" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">Jacobsen 1992a</bibRefCitation>
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,
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<bibRefCitation id="987B4B32B12FA10FE90FFEF0E202FEB2" author="Jacobsen" box="[565,616,289,311]" pageId="35" pageNumber="486" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="B4F06548B12FA10FEBADFEB5E3E8FD45" pageId="35" pageNumber="486" type="materials_examined">
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<paragraph id="FC5536C3B12FA10FEBADFEB5E5E4FE25" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
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<emphasis id="CE9EEAD1B12FA10FEBADFEB5E161FEF8" bold="true" box="[151,267,356,381]" pageId="35" pageNumber="486">
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<typeStatus id="23518861B12FA10FEBADFEB5E16CFEF8" box="[151,262,356,381]" pageId="35" pageNumber="486" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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TM 81113, adult male, Farm Hendriksplaats 281KT,
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<geoCoordinate id="99DE5004B12FA10FE850FEB5E3A0FEF9" box="[874,970,356,380]" direction="south" orientation="latitude" pageId="35" pageNumber="486" precision="925" value="-24.633333">24°38' S</geoCoordinate>
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,
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<geoCoordinate id="99DE5004B12FA10FE8E2FEB5E456FEF9" box="[984,1084,356,380]" direction="east" orientation="longitude" pageId="35" pageNumber="486" precision="925" value="30.133333">30°08' E</geoCoordinate>
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, Lydenburg District, (2430CA) Mpumalanga Province,
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<collectingCountry id="84FD7653B12FA10FEA99FE56E2AAFE25" box="[419,704,391,416]" name="South Africa" pageId="35" pageNumber="486">Republic of South Africa</collectingCountry>
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, collectors R. E. Newbery and W. Petersen,
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<date id="88541003B12FA10FEF95FE56E5E0FE25" box="[1199,1418,391,416]" pageId="35" pageNumber="486" value="1985-11-15">15 November 1985</date>
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.
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</paragraph>
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<paragraph id="FC5536C3B12FA10FEBFDFE7DE3E8FD45" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
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<emphasis id="CE9EEAD1B12FA10FEBFDFE7DE12CFE40" bold="true" box="[199,326,428,453]" pageId="35" pageNumber="486">
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<typeStatus id="23518861B12FA10FEBFDFE7DE12BFE40" box="[199,321,428,453]" pageId="35" pageNumber="486" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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TM 81115, TM 81125, 81127, same data as for
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<typeStatus id="23518861B12FA10FE85DFE7DE3A6FE40" box="[871,972,428,453]" pageId="35" pageNumber="486" type="holotype">holotype</typeStatus>
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; TM 81119, 81124, Farm Maandagshoek 254KT, Sekhukhuneland District, Limpopo Province (2430CA) collector N. H. G. Jacobsen,
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<date id="88541003B12FA10FEF9DFE1EE50DFE6D" box="[1191,1383,463,488]" pageId="35" pageNumber="486" value="1980-10-25">25 October 1980</date>
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; TM 81112, 81116–81118, same locality as TM 81119, collector R.E.Newbery,
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<date id="88541003B12FA10FEF2CFE25E4D7FD89" box="[1046,1213,500,524]" pageId="35" pageNumber="486" value="1982-06-29">29 June 1982</date>
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; TM 81126, Farm Kalkfontein 367KT, Lydenburg District, Mpumalanga Provicne (2430CC), collector N. H. G. Jacobsen,
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<date id="88541003B12FA10FEE06FDC9E0B8FDD1" pageId="35" pageNumber="486" value="1981-04-23">23 April 1981</date>
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; TM 81114, 81121, 81123, 81128, same locality as TM 81126, collectors R. E. Newbery and W. Petersen,
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<date id="88541003B12FA10FEEBAFDEDE125FDFD" pageId="35" pageNumber="486" value="1985-11-16">16 November 1985</date>
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; TM 81122, Kgoloko lokasie, Sekhukhuneland District, Limpopo Province (2429DB), collector R. E. Newbery,
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<date id="88541003B12FA10FEA0CFD55E268FD19" box="[310,514,644,669]" pageId="35" pageNumber="486" value="1980-10-27">27 October 1980</date>
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; TM 81120, Farm
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<collectingCountry id="84FD7653B12FA10FE9D4FD54E365FD19" box="[750,783,645,668]" name="South Georgia and the South Sandwich Islands" pageId="35" pageNumber="486">De</collectingCountry>
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Grooteboom 373KT, Lydenburg District, Mpumalanga Province (2430CC), collector N. H. G. Jacobsen,
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<date id="88541003B12FA10FE985FD79E314FD45" box="[703,894,679,704]" pageId="35" pageNumber="486" value="1981-10-23">23 October 1981</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12FA10FEBFDFD1DE126FC8D" pageId="35" pageNumber="486" type="etymology">
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<paragraph id="FC5536C3B12FA10FEBFDFD1DE126FC8D" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
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<emphasis id="CE9EEAD1B12FA10FEBFDFD1DE124FD60" bold="true" box="[199,334,716,741]" pageId="35" pageNumber="486">Etymology.</emphasis>
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The specific epithet refers to the Leolo hills in Sekhukhuneland, Limpopo Province, where it was first discovered.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12FA10FEBFDFCC5E478FC1D" pageId="35" pageNumber="486" type="diagnosis">
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<paragraph id="FC5536C3B12FA10FEBFDFCC5E478FC1D" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
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<emphasis id="CE9EEAD1B12FA10FEBFDFCC5E12BFCA8" bold="true" box="[199,321,788,813]" pageId="35" pageNumber="486">Diagnosis.</emphasis>
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A small
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<taxonomicName id="3BEA4D40B12FA10FEA90FCC5E276FCA8" box="[426,540,788,813]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="486" phylum="Chordata" rank="genus">
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<emphasis id="CE9EEAD1B12FA10FEA90FCC5E276FCA8" box="[426,540,788,813]" italics="true" pageId="35" pageNumber="486">Afroedura</emphasis>
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</taxonomicName>
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(maximum SVL
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<quantity id="3B129B26B12FA10FE9DEFCC5E320FCA9" box="[740,842,788,813]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" pageId="35" pageNumber="486" unit="mm" value="40.5">40.5 mm</quantity>
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) differing from all other congeners by the following combination of characters: two pairs of enlarged subdigital lamellae per digit; tail faintly verticillate and flattened near base, with four subcaudal rows and 6–7 supracaudal rows per verticil; dorsal scales smooth, 87–95 scale rows at midbody; internasal scales typically absent; 31–35 precloacal pores in males.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12FA10FEBFDFC75E521F899" pageId="35" pageNumber="486" type="description">
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<paragraph id="FC5536C3B12FA10FEBFDFC75E0BCFB10" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
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<emphasis id="CE9EEAD1B12FA10FEBFDFC75E133FC38" bold="true" box="[199,345,932,957]" pageId="35" pageNumber="486">Description.</emphasis>
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(based on
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<typeStatus id="23518861B12FA10FEADCFC75E222FC38" box="[486,584,932,957]" pageId="35" pageNumber="486" type="holotype">holotype</typeStatus>
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TM 81113) Adult male; 37.0 mm SVL; 45.0 mm TailL; mass before preservation
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<quantity id="3B129B26B12FA10FEA16FC19E106FC65" box="[300,364,968,992]" metricMagnitude="-4" metricUnit="kg" metricValue="9.0" pageId="35" pageNumber="486" unit="g" value="0.9">0.9 g</quantity>
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. Body small and slender, dorsoventrally depressed; head oval, wider than the neck. Rostral approximately 2.5 times wider than high; nostril pierced between rostral, first upper labial and three nasal scales; nasorostrals in moderate contact behind rostral. Scales on snout hexagonal, flattened and much larger than scales on crown of head; nine scales between nasals and eye and 16 scales between eye and ear. Four supraciliary spines. Supralabials 10. Mental wedge-shaped, much longer than wide and in contact with two postmentals. Infralabials eight.
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</paragraph>
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<paragraph id="FC5536C3B12FA10FEBFDFB4EE3FCFAE8" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">Dorsal scales minute, more-or-less homogeneous, smooth, juxtaposed to subimbricate, rounded to slightly hexagonal. Midbody scales 89. Ventrals large, smooth and imbricate. Digits with two pairs of enlarged scansors and six enlarged inferomedian scales under the fourth toe. Precloacal pores in a continuous, almost straight row of 34. Tail broad and flattened near the base, tapering to a fine tip, faintly verticillate; caudal scales arranged in six dorsal and four ventral rows per verticil. Supracaudals subimbricate, almost rectangular; subcaudals as broad as long and imbricate. Two postcloacal spurs on either side of tail base.</paragraph>
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<paragraph id="FC5536C3B12FA10FEBFDFAA6E3F2F9A5" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
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<emphasis id="CE9EEAD1B12FA10FEBFDFAA6E17AFA15" bold="true" box="[199,272,1399,1424]" pageId="35" pageNumber="486">Color.</emphasis>
|
||
Pale brown to brown dorsally with 7–8 dark brown irregular crossbands extending from the occiput to sacrum. Crossbands with darker posterior margins and, in most specimens, a white vertebral spot just posterior to each crossband. Limbs longitudinally striped or banded. The paler areas between the stripes are spotted dark brown. Crown of head pale brown with darker spotting and other variegations. Tail with 10 blackish crossbands from base to tip. Venter pinkish; tail brownish with darker markings.
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B12FA10FEBFDF9FDE521F899" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
|
||
<emphasis id="CE9EEAD1B12FA10FEBFDF9FDE151F9C0" bold="true" box="[199,315,1580,1605]" pageId="35" pageNumber="486">Variation.</emphasis>
|
||
<typeStatus id="23518861B12FA10FEA73F9FCE1DCF9C0" box="[329,438,1581,1605]" pageId="35" pageNumber="486" type="paratype">Paratypes</typeStatus>
|
||
and other specimens agree with the
|
||
<typeStatus id="23518861B12FA10FE853F9FDE3A1F9C0" box="[873,971,1580,1605]" pageId="35" pageNumber="486" type="holotype">holotype</typeStatus>
|
||
in most features of scalation (
|
||
<tableCitation id="B1680378B12FA10FEE09F9FDE5E5F9C1" box="[1331,1423,1580,1605]" captionStart="TABLE 4" captionStartId="27.[125,205,605,626]" captionText="TABLE 4. variatiοn in key mοrphοlοgical characters in Limpοpο and Mpumalanga Prοvinces Afroedura including, fοr cοmparative purpοses, values fοr A. pondolia and A. major. Fοr size" pageId="35" pageNumber="486">Table 4</tableCitation>
|
||
). Nasorostrals in broad contact behind rostral but separated posteriorly by a single granule in TM 81120. Scales between nasals and eye 9–12, from eye to ear 16–18. Five supraciliary spines in TM 81115. Mental as long as broad in some specimens; postmentals three in TM 811141. Supralabials 8–10. Infralabials 6–10. Midbody scale rows 87–95. 0–8 enlarged inferomedian scales under fourth toe. Precloacal pores in male
|
||
<typeStatus id="23518861B12FA10FEFB3F96CE49EF950" box="[1161,1268,1725,1749]" pageId="35" pageNumber="486" type="paratype">paratypes</typeStatus>
|
||
31–35 (except for TM 81126, which has only 11), females lacking pores. Original tails 50.00–55.9% of total length. Supracaudal scales in 6 or 7 rows per tail whorl. Tail has been autotomized in 54.5% of the specimens examined (n=17).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4F06548B12FA108EBFDF8F6E424FF50" lastPageId="36" lastPageNumber="487" pageId="35" pageNumber="486" type="distribution">
|
||
<paragraph id="FC5536C3B12FA10FEBFDF8F6E369F8E0" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
|
||
<emphasis id="CE9EEAD1B12FA10FEBFDF8F6E10AF8C5" bold="true" box="[199,352,1831,1856]" pageId="35" pageNumber="486">Distribution.</emphasis>
|
||
Endemic to the Leolo Hills and outcrops above the Steelpoort River on either side of the border between Mpumalanga and Limpopo Provinces (
|
||
<figureCitation id="64D12A46B12FA10FE98BF89DE29FF8E0" box="[689,757,1868,1893]" captionStart="FIGURE 6" captionStartId="22.[151,250,1208,1230]" captionTargetBox="[182,1404,196,1182]" captionTargetId="figure@22.[178,1409,193,1187]" captionTargetPageId="22" captionText="FIGURE 6. Map of Limpopo and Mpumalanga provinces, South Africa and adjacent regions showing the distributions of previously described species of Afroedura endemic to this region and known localities of the nine new species described in this paper. For previously recognized forms an approximate outline of the range based on Bates et al. (2014) is shown; genetically sampled localities are marked by circles in Figure 4. For new species all known localities are plotted as dots (see relevant species accounts), with genetically sampled site marked by circles with a a red center, type locality marked by a star, and all other localities indicated by open circles colored to correspond to the key in the figure. The very restricted distributions of A. maripi sp. nov., A. rondavelica sp. nov., and A. rupestris sp. nov. are indicated by arrows and are labeled directly on the map." httpUri="https://zenodo.org/record/250501/files/figure.png" pageId="35" pageNumber="486">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B12FA10FEBFDF8BEE5F2F870" blockId="35.[151,1437,356,2037]" pageId="35" pageNumber="486">
|
||
<emphasis id="CE9EEAD1B12FA10FEBFDF8BEE1E1F80D" bold="true" box="[199,395,1903,1928]" pageId="35" pageNumber="486">Natural history.</emphasis>
|
||
Exclusively rupicolous, apparently limited to norite and granitic formations.
|
||
<taxonomicName id="3BEA4D40B12FA10FEE10F8BEE16DF828" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="35" pageNumber="486" phylum="Chordata" rank="species" species="leoloensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12FA10FEE10F8BEE16DF828" italics="true" pageId="35" pageNumber="486">Afroedura leoloensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12FA10FEA34F845E103F828" box="[270,361,1940,1965]" pageId="35" pageNumber="486" rank="species">
|
||
<emphasis id="CE9EEAD1B12FA10FEA34F845E103F828" bold="true" box="[270,361,1940,1965]" pageId="35" pageNumber="486">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
lives in narrow crevices under exfoliating rock. These are usually on the underside of boulders with the openings facing downwards, protecting them from rain. The species is found in Sekhukhune Mountain Bushveld (SVcb 28) and Leolo Summit Sourveld (GM 20) (
|
||
<bibRefCitation id="987B4B32B12FA10FE9C1F80DE47CF870" author="Mucina" box="[763,1046,2012,2037]" pageId="35" pageNumber="486" refString="Mucina, L. & Rutherford, M. C. (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) at an elevation of
|
||
<quantity id="3B129B26B12FA10FEFECF80DE50EF871" box="[1238,1380,2012,2036]" metricMagnitude="3" metricUnit="m" metricValue="1.5" metricValueMax="1.8" metricValueMin="1.2" pageId="35" pageNumber="486" unit="m" value="1500.0" valueMax="1800.0" valueMin="1200.0">1200–1800 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B128A108EBFDFF49E424FF50" blockId="36.[151,1436,151,320]" pageId="36" pageNumber="487">
|
||
Two eggs are laid at a time and measure 8.9–9.0 x
|
||
<quantity id="3B129B26B128A108E9C6FF46E3E9FF2A" box="[764,899,151,176]" metricMagnitude="-3" metricUnit="m" metricValue="6.25" metricValueMax="6.4" metricValueMin="6.1" pageId="36" pageNumber="487" unit="mm" value="6.25" valueMax="6.4" valueMin="6.1">6.1–6.4 mm</quantity>
|
||
with a mass of
|
||
<quantity id="3B129B26B128A108EF08FF49E406FF35" box="[1074,1132,152,176]" metricMagnitude="-4" metricUnit="kg" metricValue="2.0" pageId="36" pageNumber="487" unit="g" value="0.2">0.2 g</quantity>
|
||
. The eggs appear to be laid in midsummer and are initially soft-shelled, adhering to the rock and then hardening.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4F06548B128A108EBFDFF0EE3B6FEC5" pageId="36" pageNumber="487" type="discussion">
|
||
<paragraph id="FC5536C3B128A108EBFDFF0EE3B6FEC5" blockId="36.[151,1436,151,320]" pageId="36" pageNumber="487">
|
||
<emphasis id="CE9EEAD1B128A108EBFDFF0EE151FF7D" bold="true" box="[199,315,223,248]" pageId="36" pageNumber="487">Remarks.</emphasis>
|
||
<taxonomicName id="3BEA4D40B128A108EA79FF0EE247FF7D" box="[323,557,223,248]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="487" phylum="Chordata" rank="species" species="leoloensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B128A108EA79FF0EE247FF7D" box="[323,557,223,248]" italics="true" pageId="36" pageNumber="487">Afroedura leoloensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB128A108E90FFF31E2E7FF72" box="[565,653,224,247]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
|
||
is a member of the
|
||
<taxonomicName id="3BEA4D40B128A108E849FF31E3A6FF7D" box="[883,972,223,248]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="487" phylum="Chordata" rank="species" species="langi">
|
||
<emphasis id="CE9EEAD1B128A108E849FF31E3A6FF7D" box="[883,972,223,248]" italics="true" pageId="36" pageNumber="487">A. langi</emphasis>
|
||
</taxonomicName>
|
||
clade (
|
||
<figureCitation id="64D12A46B128A108EF1AFF0EE4E7FF7D" box="[1056,1165,223,248]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="14.[151,250,1708,1730]" captionStartId-1="15.[151,250,1791,1813]" captionTargetBox-0="[161,1431,490,1674]" captionTargetBox-1="[233,1360,205,1772]" captionTargetId-0="figure@14.[147,1432,481,1677]" captionTargetId-1="figure@15.[220,1360,193,1773]" captionTargetPageId-0="14" captionTargetPageId-1="15" captionText-0="FIGURE 1. Maximum likelihood phylogram of Afroedura relationships derived from two genes (ND 2, RAG 1). Support values are presented as maximum likelihood bootstraps (MLBS) / Bayesian posterior probabilities (BPP) / parsimony bootstraps (PBS). Only most closely related outgroup taxa are shown. Images (top to bottom) of representative Afroedura species groups: A. hawequensis, A. pondolia, A. karroica, A. loveridgei, A. haackei, A. marleyi, A. granitica sp. nov." captionText-1="FIGURE 2. Results of the 12 - gene concatenated phylogenetic analysis and 10 individual nuclear gene trees. The tree shows the topology as estimated in the two-gene (ND 2, RAG 1) concatenated analysis. Outgroup taxa (not shown) include Chondrodactylus, Pachydactylus, Rhoptropus, and Goggia. The tree has been pruned from the full dataset (72 ingroup taxa) to indicate species-level relationships only (22 ingroup taxa). Solid circles indicate> 90 MLBS / 1.0 BPP for the 12 - gene concatenated analysis at the node, and open circles indicate 70 – 90 MLBS / 9.0 – 9.9 BPP. Each individual nuclear gene not used in the two-gene concatenated analyses that was able to recover the same species relationship as in the two-gene analysis (Fig. 1) is listed on a node-by-node basis. Bold italic black names indicate> 70 MLBS for that particular gene, and gray names in normal font indicate <70 MLBS." httpUri-0="https://zenodo.org/record/250496/files/figure.png" httpUri-1="https://zenodo.org/record/250497/files/figure.png" pageId="36" pageNumber="487">Figs. 1–2</figureCitation>
|
||
; see
|
||
<emphasis id="CE9EEAD1B128A108EFFDFF0EE55EFF7D" bold="true" box="[1223,1332,223,248]" pageId="36" pageNumber="487">Remarks</emphasis>
|
||
under
|
||
<taxonomicName id="3BEA4D40B128A108EEBEFF31E094FE99" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="487" phylum="Chordata" rank="species" species="granitica" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B128A108EEBEFF31E094FE99" italics="true" pageId="36" pageNumber="487">A. granitica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB128A108EA3FFED5E135FE98" box="[261,351,260,285]" pageId="36" pageNumber="487" rank="species">
|
||
<emphasis id="CE9EEAD1B128A108EA3FFED5E135FE98" bold="true" box="[261,351,260,285]" pageId="36" pageNumber="487">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
). This species exhibits the highest number of precloacal pores (31–35) not only in its clade, but in the genus as a whole, and may thus be distinguished from its congeners.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |