292 lines
34 KiB
XML
292 lines
34 KiB
XML
<document id="B87157EC75C9A5B2CCE39017107B542A" ID-DOI="10.11646/zootaxa.3846.4.1" ID-GBIF-Dataset="317a43cd-c408-4fde-8374-357fbb9a0d53" ID-ISSN="1175-5326" ID-Zenodo-Dep="250495" ID-ZooBank="0DD5A603-D65F-4976-BBE9-94DA7110053F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467125281" checkinUser="plazi" docAuthor="Jacobsen, Niels H. G., Kuhn, Arianna L., Jackman, Todd R. & Bauer, Aaron M." docDate="2014" docId="744387D5B12CA10DEBADFA50E17AF8AD" docLanguage="en" docName="zt03846p501.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afroedura granitica Jacobsen, Kuhn, Jackman & Bauer, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="484" masterDocId="887AFFADB10CA12CEB3AFFD1E06AFF85" masterDocTitle="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa" masterLastPageNumber="501" masterPageNumber="451" pageNumber="483" updateTime="1698305158428" updateUser="plazi">
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<mods:titleInfo id="B4611113B7FB5CA01B381C173B675C8B">
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<mods:title id="A84C52989304626A9CCDF9F77F8A538C">A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa</mods:title>
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</mods:titleInfo>
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<mods:name id="5F64BEEE1F8DA8A7A3F8E87F1258A393" type="personal">
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<mods:roleTerm id="60EF0E5CE74233A5E474B9BD284AD928">Author</mods:roleTerm>
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<mods:namePart id="58B1291AAAAD9C46B73C0FD804F94F2D">Jacobsen, Niels H. G.</mods:namePart>
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<mods:roleTerm id="8686C9E660349934402A31EED49B8F6A">Author</mods:roleTerm>
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<mods:namePart id="A48802ACC2927A5DA0047F786689B10E">Kuhn, Arianna L.</mods:namePart>
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<mods:name id="72D9B95938C326832EED857757CF8719" type="personal">
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<mods:roleTerm id="30E6EDA24CA13B078B99F7853A43705B">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="363348BB4C676E4116B5F5220C77A6FD">Jackman, Todd R.</mods:namePart>
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</mods:name>
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<mods:name id="5B6344EB77FB09F233B903CC938BF367" type="personal">
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<mods:roleTerm id="59ACADC173F8760AABF8ED944B4AF7E7">Author</mods:roleTerm>
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<mods:namePart id="56BFBDE3ED39B4BAC2110E8A8582F52F">Bauer, Aaron M.</mods:namePart>
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<mods:typeOfResource id="A4B8030BC416EB700BB69A897EEE7FBA">text</mods:typeOfResource>
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<mods:title id="863C38397E4FF566B3FF283B6C6AAE0A">Zootaxa</mods:title>
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<mods:part id="671B087C55A92DA2A73BEF98291165F8">
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<mods:date id="AA87DBD2EC3112AD9DBF6DDC26759809">2014</mods:date>
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<mods:detail id="A1B5B494045670AA75974083E30942FD" type="volume">
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<mods:number id="4916ECA7CB1E5B5F728DCC10DA86F2E5">3846</mods:number>
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</mods:detail>
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<mods:detail id="CDCF694D877566B634CCB1F63D47BF8D" type="issue">
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<mods:number id="45A83DEA0DA6B5EFC6BC1B4EF29BB4D9">4</mods:number>
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<mods:start id="AA72CB358C2EA6DA05E3409FE7433DC5">451</mods:start>
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<mods:classification id="A51CB9557F1ECBD0549B60E7ED4E3FF9">journal article</mods:classification>
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<mods:identifier id="8B74548B69D2FE3082C45DB262D6D41F" type="DOI">10.11646/zootaxa.3846.4.1</mods:identifier>
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<mods:identifier id="F3876C621F03B7C88B41B9B0B4B6B426" type="GBIF-Dataset">317a43cd-c408-4fde-8374-357fbb9a0d53</mods:identifier>
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<mods:identifier id="9EB0363153930840AF3FEF117422B92A" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="5D748C18CFA430EEB6D78881EDB8E975" type="Zenodo-Dep">250495</mods:identifier>
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<mods:identifier id="0A581113F929EE187FDC06C03845EE2C" type="ZooBank">0DD5A603-D65F-4976-BBE9-94DA7110053F</mods:identifier>
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</mods:mods>
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<treatment id="744387D5B12CA10DEBADFA50E17AF8AD" ID-DOI="http://doi.org/10.5281/zenodo.5620593" ID-GBIF-Taxon="119514190" ID-Zenodo-Dep="5620593" LSID="urn:lsid:plazi:treatment:744387D5B12CA10DEBADFA50E17AF8AD" httpUri="http://treatment.plazi.org/id/744387D5B12CA10DEBADFA50E17AF8AD" lastPageId="33" lastPageNumber="484" pageId="32" pageNumber="483">
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<subSubSection id="B4F06548B12CA10CEBADFA50E167FA39" pageId="32" pageNumber="483" type="nomenclature">
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<paragraph id="FC5536C3B12CA10CEBADFA50E187FA1E" blockId="32.[151,493,1409,1469]" box="[151,493,1409,1435]" pageId="32" pageNumber="483">
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<heading id="A71D81AFB12CA10CEBADFA50E187FA1E" bold="true" box="[151,493,1409,1435]" fontSize="11" level="1" pageId="32" pageNumber="483" reason="1">
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<emphasis id="CE9EEAD1B12CA10CEBADFA50E187FA1E" bold="true" box="[151,493,1409,1435]" pageId="32" pageNumber="483">
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<taxonomicName id="3BEA4D40B12CA10CEBADFA50E1E3FA1E" box="[151,393,1409,1435]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="granitica" status="sp. nov.">
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<emphasis id="CE9EEAD1B12CA10CEBADFA50E1E3FA1E" bold="true" box="[151,393,1409,1435]" italics="true" pageId="32" pageNumber="483">Afroedura granitica</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D5AD57AAB12CA10CEAAAFA50E187FA1E" box="[400,493,1409,1435]" pageId="32" pageNumber="483" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="FC5536C3B12CA10CEBADFA75E167FA39" blockId="32.[151,493,1409,1469]" box="[151,269,1444,1469]" pageId="32" pageNumber="483">
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(
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<figureCitation id="64D12A46B12CA10CEBA5FA75E085FA39" box="[159,239,1444,1469]" captionStart="FIGURE 10" captionStartId="34.[151,250,1912,1934]" captionTargetBox="[221,1375,195,1892]" captionTargetId="figure@34.[221,1376,195,1893]" captionTargetPageId="34" captionText="FIGURE 10. New species in the Afroedura langi group. (A) Afroedura granitica sp. nov. from Farm Lillie, Limpopo Province, South Africa (24 ° 04 ’ S, 30 ° 51 ’ E) (Photo Stuart V. Nielsen). (B) Afroedura leoloensis sp. nov. from Farm Maandagshoek, Limpopo Province, South Africa (Photo N. H. G. Jacobsen). (C) Afroedura broadleyi sp. nov. from 22 ° 58 ’ 40 ” S, 29 ° 19 ’ 57 ” E, Limpopo Province, South Africa (Photo Marius Burger)." httpUri="https://zenodo.org/record/250505/files/figure.png" pageId="32" pageNumber="483">Fig. 10</figureCitation>
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A)
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12CA10CEBADFA3DE20EF987" box="[151,612,1516,1538]" pageId="32" pageNumber="483" type="reference_group">
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<paragraph id="FC5536C3B12CA10CEBADFA3DE20EF987" blockId="32.[151,612,1516,1538]" box="[151,612,1516,1538]" pageId="32" pageNumber="483">
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<taxonomicName id="3BEA4D40B12CA10CEBADFA3DE152F987" box="[151,312,1516,1538]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="483" phylum="Chordata" rank="species" species="langi">
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<emphasis id="CE9EEAD1B12CA10CEBADFA3DE152F987" box="[151,312,1516,1538]" italics="true" pageId="32" pageNumber="483">Afroedura langi</emphasis>
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</taxonomicName>
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'Lillie'
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<bibRefCitation id="987B4B32B12CA10CEABFFA3DE24DF987" author="Jacobsen" box="[389,551,1516,1538]" pageId="32" pageNumber="483" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">Jacobsen 1992a</bibRefCitation>
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,
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<bibRefCitation id="987B4B32B12CA10CE90BFA3DE20EF987" author="Jacobsen" box="[561,612,1516,1538]" pageId="32" pageNumber="483" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12CA10CEBADF9FFE361F936" pageId="32" pageNumber="483" type="materials_examined">
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<paragraph id="FC5536C3B12CA10CEBADF9FFE369F9EE" blockId="32.[151,1436,1582,2003]" pageId="32" pageNumber="483">
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<emphasis id="CE9EEAD1B12CA10CEBADF9FFE161F9C2" bold="true" box="[151,267,1582,1607]" pageId="32" pageNumber="483">
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<typeStatus id="23518861B12CA10CEBADF9FFE16CF9C2" box="[151,262,1582,1607]" pageId="32" pageNumber="483" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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TM 81187, adult male, Farm Lillie 148KT,
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<geoCoordinate id="99DE5004B12CA10CE820F9FEE314F9C2" box="[794,894,1583,1607]" direction="south" orientation="latitude" pageId="32" pageNumber="483" precision="925" value="-24.066668">24°04' S</geoCoordinate>
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,
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<geoCoordinate id="99DE5004B12CA10CE8B5F9FFE39FF9C3" box="[911,1013,1582,1607]" direction="east" orientation="longitude" pageId="32" pageNumber="483" precision="925" value="30.85">30°51' E</geoCoordinate>
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, Letaba district (2430BB), Northern Province, collector R. E. Newbery,
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<date id="88541003B12CA10CE91EF982E294F9EE" box="[548,766,1618,1643]" pageId="32" pageNumber="483" value="1985-11-11">11 November 1985</date>
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.
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</paragraph>
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<paragraph id="FC5536C3B12CA10CEBFDF9A7E361F936" blockId="32.[151,1436,1582,2003]" pageId="32" pageNumber="483">
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<emphasis id="CE9EEAD1B12CA10CEBFDF9A7E12CF90A" bold="true" box="[199,326,1654,1679]" pageId="32" pageNumber="483">
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<typeStatus id="23518861B12CA10CEBFDF9A7E12BF90A" box="[199,321,1654,1679]" pageId="32" pageNumber="483" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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TM 81239-81240, 81243–81247, same data as for
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<typeStatus id="23518861B12CA10CE8AFF9A7E391F90A" box="[917,1019,1654,1679]" pageId="32" pageNumber="483" type="holotype">holotype</typeStatus>
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; TM 81241–81242, same locality as for
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<typeStatus id="23518861B12CA10CEB84F94BE149F936" box="[190,291,1690,1715]" pageId="32" pageNumber="483" type="holotype">holotype</typeStatus>
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, collector N. H. G. Jacobsen,
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<date id="88541003B12CA10CE954F94AE36DF936" box="[622,775,1691,1715]" pageId="32" pageNumber="483" value="1980-05-17">17 May 1980</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12CA10CEBFDF96FE5F1F952" box="[199,1435,1726,1751]" pageId="32" pageNumber="483" type="etymology">
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<paragraph id="FC5536C3B12CA10CEBFDF96FE5F1F952" blockId="32.[151,1436,1582,2003]" box="[199,1435,1726,1751]" pageId="32" pageNumber="483">
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<emphasis id="CE9EEAD1B12CA10CEBFDF96FE124F952" bold="true" box="[199,334,1726,1751]" pageId="32" pageNumber="483">Etymology.</emphasis>
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The specific epithet refers to the decomposing granite hills to which this gecko appears restricted.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12CA10CEBFDF933E26DF80E" pageId="32" pageNumber="483" type="diagnosis">
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<paragraph id="FC5536C3B12CA10CEBFDF933E26DF80E" blockId="32.[151,1436,1582,2003]" pageId="32" pageNumber="483">
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<emphasis id="CE9EEAD1B12CA10CEBFDF933E12BF97E" bold="true" box="[199,321,1762,1787]" pageId="32" pageNumber="483">Diagnosis.</emphasis>
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A medium to large
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<taxonomicName id="3BEA4D40B12CA10CE91EF933E2FCF97E" box="[548,662,1762,1787]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="32" pageNumber="483" phylum="Chordata" rank="genus">
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<emphasis id="CE9EEAD1B12CA10CE91EF933E2FCF97E" box="[548,662,1762,1787]" italics="true" pageId="32" pageNumber="483">Afroedura</emphasis>
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</taxonomicName>
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(maximum SVL 58.0 mm) differing from all other congeners by the following combination of characters: two pairs of enlarged subdigital lamellae per digit; tail not obviously verticillate and weakly flattened, with four subcaudal rows and seven supracaudal rows per verticil; dorsal scales smooth, 99–107 scale rows at midbody; internasal scales typically absent, a deep furrow between nasorostrals; 13–16 precloacal pores in males.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12CA10DEBFDF847E161FBDD" lastPageId="33" lastPageNumber="484" pageId="32" pageNumber="483" type="description">
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<paragraph id="FC5536C3B12CA10DEBFDF847E2B6FEBA" blockId="32.[151,1436,1582,2003]" lastBlockId="33.[151,1437,151,1832]" lastPageId="33" lastPageNumber="484" pageId="32" pageNumber="483">
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<emphasis id="CE9EEAD1B12CA10CEBFDF847E133F82A" bold="true" box="[199,345,1942,1967]" pageId="32" pageNumber="483">Description.</emphasis>
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(based on
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<typeStatus id="23518861B12CA10CEADCF847E222F82A" box="[486,584,1942,1967]" pageId="32" pageNumber="483" type="holotype">holotype</typeStatus>
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TM 81187) Adult male; 55.0 mm SVL; 62.0 mm TailL; mass before preservation 3.0 g. Dorsoventrally depressed; head ovate, distinctly wider than neck. Forelimbs small, hindlimbs robust. Rostral more than twice as wide as high; nostril pierced between rostral, first upper labial and three nasal scales; nasorostrals in broad contact behind rostral, with a deep furrow between them. Scales on snout rounded and juxtaposed, much larger than granules on crown of head; ten scales between nasals and eye and 18 scales between eye and ear; three supraciliary spines; supralabials nine. Mental wedge-shaped, much longer than wide and in contact with a single postmental. Infralabials seven.
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</paragraph>
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<paragraph id="FC5536C3B12DA10DEBFDFE9DE173FDB8" blockId="33.[151,1437,151,1832]" pageId="33" pageNumber="484">Dorsal granules uniform, smooth, rounded and juxtaposed. Midbody scales 100. Ventrals lozenge-shaped and imbricate. Digits with two pairs of scansors and four enlarged inferomedian scales under the 4th toe. Fifteen continuous precloacal pores arranged in a curved row. Tail flattened proximally and thickening about one verticil length posterior to the cloaca. Tail inconspicuously verticillate and tapering, with caudal scales in seven dorsal and four ventral rows per verticil; supracaudals subimbricate, squarish to rectangular, with a rounded posterior margin; subcaudals large, squarish to hexagonal with rounded posterior margins, subimbricate. Two postcloacal spurs at base of tail.</paragraph>
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<paragraph id="FC5536C3B12DA10DEBFDFD96E560FCBD" blockId="33.[151,1437,151,1832]" pageId="33" pageNumber="484">
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<emphasis id="CE9EEAD1B12DA10DEBFDFD96E17AFDE5" bold="true" box="[199,272,583,608]" pageId="33" pageNumber="484">Color.</emphasis>
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The crown of the head is pale brown with dark brown spots and stripes that are also found on the snout. A dark brown stripe extends from the nostril through the eye, and above the ear to communicate with the occipital bar. Pale gray-brown to brown dorsally with six wavy dark brown irregular crossbars between occiput and sacrum. Crossbars are edged with black posteriorly and in their posterior indentations have white or whitish spots, particularly along the vertebral column. Variable pale and dark spots and blotches present between crossbars. The limbs have a variable dark brown reticulate pattern. Original tails with at least 10 dark brown to blackish crossbands with half bands and speckling in the intervening areas. Venter white, subcaudals speckled.
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</paragraph>
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<paragraph id="FC5536C3B12DA10DEBFDFC95E161FBDD" blockId="33.[151,1437,151,1832]" pageId="33" pageNumber="484">
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<emphasis id="CE9EEAD1B12DA10DEBFDFC95E151FCD8" bold="true" box="[199,315,836,861]" pageId="33" pageNumber="484">Variation.</emphasis>
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<typeStatus id="23518861B12DA10DEA73FC94E1DCFCD8" box="[329,438,837,861]" pageId="33" pageNumber="484" type="paratype">Paratypes</typeStatus>
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and other specimens agree with the
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<typeStatus id="23518861B12DA10DE853FC95E3A1FCD8" box="[873,971,836,861]" pageId="33" pageNumber="484" type="holotype">holotype</typeStatus>
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in most features of scalation (
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<tableCitation id="B1680378B12DA10DEE09FC95E5E5FCD9" box="[1331,1423,836,861]" captionStart="TABLE 4" captionStartId="27.[125,205,605,626]" captionText="TABLE 4. variatiοn in key mοrphοlοgical characters in Limpοpο and Mpumalanga Prοvinces Afroedura including, fοr cοmparative purpοses, values fοr A. pondolia and A. major. Fοr size" pageId="33" pageNumber="484">Table 4</tableCitation>
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).
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<typeStatus id="23518861B12DA10DEBADFCB9E16EFC05" box="[151,260,872,896]" pageId="33" pageNumber="484" type="paratype">Paratypes</typeStatus>
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TM 81244 and TM 81246 with nasorostrals separated by a single granular scale. Scales between nasals and eye 10–12, from eye to ear 18–22. Postmentals 1–2. Supralabials 7–10. Midbody scale rows 99–107. No enlarged inferomedian scales under the fourth toe in TM 81243. Precloacal pores in male
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<typeStatus id="23518861B12DA10DEF45FC61E481FC4D" box="[1151,1259,944,968]" pageId="33" pageNumber="484" type="paratype">paratypes</typeStatus>
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13–16, females lacking pores but female
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<typeStatus id="23518861B12DA10DEA8AFC04E27AFC68" box="[432,528,981,1005]" pageId="33" pageNumber="484" type="paratype">paratype</typeStatus>
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TM 81243 with a row of elongate scales in precloacal position. Postcloacal spurs 1–2, absent in some females. Original tails 53.18–58.3% of total length. Degree of distinctness of tail whorls variable; faint, but clearly evident in some specimens. Tail has been autotomized in only 20% of the specimens examined.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B12DA10DEBFDFBB5E2BCFAD1" pageId="33" pageNumber="484" type="distribution">
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<paragraph id="FC5536C3B12DA10DEBFDFBB5E098FB25" blockId="33.[151,1437,151,1832]" pageId="33" pageNumber="484">
|
||
<emphasis id="CE9EEAD1B12DA10DEBFDFBB5E10AFBF8" bold="true" box="[199,352,1124,1149]" pageId="33" pageNumber="484">Distribution.</emphasis>
|
||
Restricted to granite hills and outcrops in the eastern Lowveld near Mica, Limpopo Province (
|
||
<figureCitation id="64D12A46B12DA10DEBA5FB56E08EFB25" box="[159,228,1159,1184]" captionStart="FIGURE 6" captionStartId="22.[151,250,1208,1230]" captionTargetBox="[182,1404,196,1182]" captionTargetId="figure@22.[178,1409,193,1187]" captionTargetPageId="22" captionText="FIGURE 6. Map of Limpopo and Mpumalanga provinces, South Africa and adjacent regions showing the distributions of previously described species of Afroedura endemic to this region and known localities of the nine new species described in this paper. For previously recognized forms an approximate outline of the range based on Bates et al. (2014) is shown; genetically sampled localities are marked by circles in Figure 4. For new species all known localities are plotted as dots (see relevant species accounts), with genetically sampled site marked by circles with a a red center, type locality marked by a star, and all other localities indicated by open circles colored to correspond to the key in the figure. The very restricted distributions of A. maripi sp. nov., A. rondavelica sp. nov., and A. rupestris sp. nov. are indicated by arrows and are labeled directly on the map." httpUri="https://zenodo.org/record/250501/files/figure.png" pageId="33" pageNumber="484">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B12DA10DEBFDFB7DE2BCFAD1" blockId="33.[151,1437,151,1832]" pageId="33" pageNumber="484">
|
||
<emphasis id="CE9EEAD1B12DA10DEBFDFB7DE1EEFB40" bold="true" box="[199,388,1196,1221]" pageId="33" pageNumber="484">Natural history.</emphasis>
|
||
Apparently restricted to decomposing granite outcrops and hillsides on the farm Lillie 148KT in veld
|
||
<typeStatus id="23518861B12DA10DEBD7FB00E177FB6C" box="[237,285,1233,1257]" pageId="33" pageNumber="484">type</typeStatus>
|
||
SVl 3 Granite Lowveld (
|
||
<bibRefCitation id="987B4B32B12DA10DE905FB1EE313FB6D" author="Mucina" box="[575,889,1231,1256]" pageId="33" pageNumber="484" refString="Mucina, L. & Rutherford, M. C. (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) at elevations of
|
||
<quantity id="3B129B26B12DA10DEF05FB1EE4D5FB6D" box="[1087,1215,1231,1256]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="33" pageNumber="484" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">600–800 m</quantity>
|
||
a.s.l. These geckos inhabit crevices formed by exfoliating sheets on the underside of boulders, as well as crevices between rocks. They are occasionally found occupying the same retreats as large flat rock scorpions (
|
||
<taxonomicName id="3BEA4D40B12DA10DEF27FAC6E575FAB5" box="[1053,1311,1303,1328]" class="Arachnida" family="Hormuridae" genus="Hadogenes" higherTaxonomySource="CoL" kingdom="Animalia" order="Scorpiones" pageId="33" pageNumber="484" phylum="Arthropoda" rank="species" species="troglodytes">
|
||
<emphasis id="CE9EEAD1B12DA10DEF27FAC6E575FAB5" box="[1053,1311,1303,1328]" italics="true" pageId="33" pageNumber="484">Hadogenes troglodytes</emphasis>
|
||
</taxonomicName>
|
||
). The very similar
|
||
<taxonomicName id="3BEA4D40B12DA10DEBD6FAEDE12FFAD0" box="[236,325,1340,1365]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="langi">
|
||
<emphasis id="CE9EEAD1B12DA10DEBD6FAEDE12FFAD0" box="[236,325,1340,1365]" italics="true" pageId="33" pageNumber="484">A. langi</emphasis>
|
||
</taxonomicName>
|
||
occurs on small schistose outcrops.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4F06548B12DA10DEBFDFA8EE17AF8AD" pageId="33" pageNumber="484" type="discussion">
|
||
<paragraph id="FC5536C3B12DA10DEBFDFA8EE17AF8AD" blockId="33.[151,1437,151,1832]" pageId="33" pageNumber="484">
|
||
<emphasis id="CE9EEAD1B12DA10DEBFDFA8EE151FAFD" bold="true" box="[199,315,1375,1400]" pageId="33" pageNumber="484">Remarks.</emphasis>
|
||
<taxonomicName id="3BEA4D40B12DA10DEA7DFA8EE246FAFD" box="[327,556,1375,1400]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="granitica" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DEA7DFA8EE246FAFD" box="[327,556,1375,1400]" italics="true" pageId="33" pageNumber="484">Afroedura granitica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DE90DFA8EE2FCFAFD" box="[567,662,1375,1400]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DE90DFA8EE2FCFAFD" bold="true" box="[567,662,1375,1400]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shares a suite of characters with
|
||
<taxonomicName id="3BEA4D40B12DA10DEF16FAB1E4E0FAFD" box="[1068,1162,1375,1400]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="langi">
|
||
<emphasis id="CE9EEAD1B12DA10DEF16FAB1E4E0FAFD" box="[1068,1162,1375,1400]" italics="true" pageId="33" pageNumber="484">A. langi</emphasis>
|
||
</taxonomicName>
|
||
and four other species described herein as new. The affinities of all but one of these,
|
||
<taxonomicName id="3BEA4D40B12DA10DE865FA55E39AFA18" box="[863,1008,1412,1437]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="leoloensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DE865FA55E39AFA18" box="[863,1008,1412,1437]" italics="true" pageId="33" pageNumber="484">A. leoloensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DE8C2FA55E439FA18" box="[1016,1107,1412,1437]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DE8C2FA55E439FA18" bold="true" box="[1016,1107,1412,1437]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
have been confirmed by our molecular phylogeny (
|
||
<figureCitation id="64D12A46B12DA10DEAADFA76E1B1FA45" box="[407,475,1447,1472]" captionStart="FIGURE 1" captionStartId="14.[151,250,1708,1730]" captionTargetBox="[161,1431,490,1674]" captionTargetId="figure@14.[147,1432,481,1677]" captionTargetPageId="14" captionText="FIGURE 1. Maximum likelihood phylogram of Afroedura relationships derived from two genes (ND 2, RAG 1). Support values are presented as maximum likelihood bootstraps (MLBS) / Bayesian posterior probabilities (BPP) / parsimony bootstraps (PBS). Only most closely related outgroup taxa are shown. Images (top to bottom) of representative Afroedura species groups: A. hawequensis, A. pondolia, A. karroica, A. loveridgei, A. haackei, A. marleyi, A. granitica sp. nov." httpUri="https://zenodo.org/record/250496/files/figure.png" pageId="33" pageNumber="484">Fig. 1</figureCitation>
|
||
). In comparison with other members of this group
|
||
<taxonomicName id="3BEA4D40B12DA10DEF11FA79E4D9FA45" box="[1067,1203,1448,1472]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="granitica" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DEF11FA79E4D9FA45" box="[1067,1203,1448,1472]" italics="true" pageId="33" pageNumber="484">A. granitica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DEF86FA76E572FA45" box="[1212,1304,1447,1472]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DEF86FA76E572FA45" bold="true" box="[1212,1304,1447,1472]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(maximum SVL
|
||
<quantity id="3B129B26B12DA10DEBE8FA1DE14BFA61" box="[210,289,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="33" pageNumber="484" unit="mm" value="58.0">58 mm</quantity>
|
||
) is substantially larger than
|
||
<taxonomicName id="3BEA4D40B12DA10DE961FA1CE2D9FA60" box="[603,691,1484,1509]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="langi">
|
||
<emphasis id="CE9EEAD1B12DA10DE961FA1CE206FA61" box="[603,620,1485,1508]" italics="true" pageId="33" pageNumber="484">A</emphasis>
|
||
.
|
||
<emphasis id="CE9EEAD1B12DA10DE943FA1DE2D9FA60" box="[633,691,1484,1509]" italics="true" pageId="33" pageNumber="484">langi</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BEA4D40B12DA10DE9D2FA1CE3DAFA60" box="[744,944,1484,1509]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="waterbergensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DE9D2FA1CE3DAFA60" box="[744,944,1484,1509]" italics="true" pageId="33" pageNumber="484">A. waterbergensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DE88DFA1DE47AFA60" box="[951,1040,1484,1509]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DE88DFA1DE47AFA60" bold="true" box="[951,1040,1484,1509]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(both reaching only
|
||
<quantity id="3B129B26B12DA10DEFCDFA1DE52DFA61" box="[1271,1351,1484,1509]" metricMagnitude="-2" metricUnit="m" metricValue="4.6" pageId="33" pageNumber="484" unit="mm" value="46.0">46 mm</quantity>
|
||
) and
|
||
<taxonomicName id="3BEA4D40B12DA10DEEBEFA1CE16DF98D" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="leoloensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DEEBEFA1CE16DF98D" italics="true" pageId="33" pageNumber="484">A. leoloensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DEA2AFA3EE106F98D" box="[272,364,1519,1544]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DEA2AFA3EE106F98D" bold="true" box="[272,364,1519,1544]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(
|
||
<quantity id="3B129B26B12DA10DEA47FA3EE18EF98D" box="[381,484,1519,1544]" metricMagnitude="-2" metricUnit="m" metricValue="4.05" pageId="33" pageNumber="484" unit="mm" value="40.5">40.5 mm</quantity>
|
||
maximum SVL). In comparison with the last of these species it also has far fewer precloacal pores (13–16
|
||
<emphasis id="CE9EEAD1B12DA10DEA96F9C4E19EF9A9" box="[428,500,1557,1580]" italics="true" pageId="33" pageNumber="484">versus</emphasis>
|
||
typically 31–35), whereas compared to
|
||
<taxonomicName id="3BEA4D40B12DA10DE883F9C4E4E8F9A8" box="[953,1154,1556,1581]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="waterbergensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DE883F9C4E4E8F9A8" box="[953,1154,1556,1581]" italics="true" pageId="33" pageNumber="484">A. waterbergensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DEFB0F9C5E48EF9A8" box="[1162,1252,1556,1581]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DEFB0F9C5E48EF9A8" bold="true" box="[1162,1252,1556,1581]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(4–7 pores) and
|
||
<taxonomicName id="3BEA4D40B12DA10DEBADF9E9E175F9D5" box="[151,287,1591,1616]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DEBADF9E9E175F9D5" box="[151,287,1591,1616]" italics="true" pageId="33" pageNumber="484">A. broadleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DEA1CF9E6E115F9D5" box="[294,383,1591,1616]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DEA1CF9E6E115F9D5" bold="true" box="[294,383,1591,1616]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(8–12 pores) it has more. In addition, in comparison with the superficially similar
|
||
<taxonomicName id="3BEA4D40B12DA10DEE2EF9E9E506F9D5" box="[1300,1388,1591,1616]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="langi">
|
||
<emphasis id="CE9EEAD1B12DA10DEE2EF9E9E506F9D5" box="[1300,1388,1591,1616]" italics="true" pageId="33" pageNumber="484">A. langi</emphasis>
|
||
</taxonomicName>
|
||
, the nasorostrals of
|
||
<taxonomicName id="3BEA4D40B12DA10DEA79F98CE1A0F9F1" box="[323,458,1629,1652]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="granitica" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DEA79F98CE13EF9F1" box="[323,340,1629,1652]" italics="true" pageId="33" pageNumber="484">A</emphasis>
|
||
.
|
||
<emphasis id="CE9EEAD1B12DA10DEA59F98CE1A0F9F1" box="[355,458,1629,1652]" italics="true" pageId="33" pageNumber="484">granitica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DEAEBF98DE241F9F0" box="[465,555,1628,1653]" pageId="33" pageNumber="484" rank="species">
|
||
<emphasis id="CE9EEAD1B12DA10DEAEBF98DE241F9F0" bold="true" box="[465,555,1628,1653]" pageId="33" pageNumber="484">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
form a deep furrow between them (versus weakly raised nasorostrals forming a shallow furrow).
|
||
<taxonomicName id="3BEA4D40B12DA10DEA4BF9AEE239F91D" box="[369,595,1663,1688]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="granitica" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DEA4BF9AEE239F91D" box="[369,595,1663,1688]" italics="true" pageId="33" pageNumber="484">Afroedura granitica</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CE9EEAD1B12DA10DE966F9AEE2DDF91D" bold="true" box="[604,695,1663,1688]" pageId="33" pageNumber="484">
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DE966F9AEE2DDF91D" box="[604,695,1663,1688]" pageId="33" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is most similar in the basic diagnostic features used herein to
|
||
<taxonomicName id="3BEA4D40B12DA10DEEBEF951E09CF939" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="33" pageNumber="484" phylum="Chordata" rank="species" species="pienaari" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B12DA10DEEBEF951E09CF939" italics="true" pageId="33" pageNumber="484">A. pienaari</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CE9EEAD1B12DA10DEBC6F975E13CF938" bold="true" box="[252,342,1700,1725]" pageId="33" pageNumber="484">
|
||
<taxonomicNameLabel id="D5AD57AAB12DA10DEBC6F975E13CF938" box="[252,342,1700,1725]" pageId="33" pageNumber="484" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, to which it is rather distantly related. It may be distinguished by its larger size (58
|
||
<emphasis id="CE9EEAD1B12DA10DEFC5F974E52DF939" box="[1279,1351,1701,1724]" italics="true" pageId="33" pageNumber="484">versus</emphasis>
|
||
<quantity id="3B129B26B12DA10DEE77F975E5F6F939" box="[1357,1436,1700,1725]" metricMagnitude="-2" metricUnit="m" metricValue="5.3" pageId="33" pageNumber="484" unit="mm" value="53.0">53 mm</quantity>
|
||
maximum SVL and even greater mean differences;
|
||
<tableCitation id="B1680378B12DA10DE9D3F916E32AF965" box="[745,832,1735,1760]" captionStart="TABLE 4" captionStartId="27.[125,205,605,626]" captionText="TABLE 4. variatiοn in key mοrphοlοgical characters in Limpοpο and Mpumalanga Prοvinces Afroedura including, fοr cοmparative purpοses, values fοr A. pondolia and A. major. Fοr size" pageId="33" pageNumber="484">Table 4</tableCitation>
|
||
), lower number of male precloacal pores (mean ~14
|
||
<emphasis id="CE9EEAD1B12DA10DEBADF93CE0B5F881" box="[151,223,1773,1796]" italics="true" pageId="33" pageNumber="484">versus</emphasis>
|
||
~16), and higher number of eye to ear scales (mean of 19.10 versus 16.20), although there is overlap in all characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |