274 lines
33 KiB
XML
274 lines
33 KiB
XML
<document id="C50B4D2E39920CAA621F72742ED83763" ID-DOI="10.11646/zootaxa.3846.4.1" ID-GBIF-Dataset="317a43cd-c408-4fde-8374-357fbb9a0d53" ID-ISSN="1175-5326" ID-Zenodo-Dep="250495" ID-ZooBank="0DD5A603-D65F-4976-BBE9-94DA7110053F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467125281" checkinUser="plazi" docAuthor="Jacobsen, Niels H. G., Kuhn, Arianna L., Jackman, Todd R. & Bauer, Aaron M." docDate="2014" docId="744387D5B128A109EBADFE45E3F0FB25" docLanguage="en" docName="zt03846p501.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afroedura broadleyi Jacobsen, Kuhn, Jackman & Bauer, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="488" masterDocId="887AFFADB10CA12CEB3AFFD1E06AFF85" masterDocTitle="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa" masterLastPageNumber="501" masterPageNumber="451" pageNumber="487" updateTime="1698305158428" updateUser="plazi">
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<mods:title id="0F3D1F3C2F29D213FD5FAEBCA35265CF">A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa</mods:title>
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<mods:namePart id="B70BBD4A8FD73664A1E7FAD5AC00F4E9">Jacobsen, Niels H. G.</mods:namePart>
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<mods:namePart id="43C51C0055A8DE2C4852785527AD47AF">Kuhn, Arianna L.</mods:namePart>
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<mods:namePart id="50977D379D122B9347425BBE180FA401">Jackman, Todd R.</mods:namePart>
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<mods:roleTerm id="35CB9B556B0C031CFAA9825F3410EDB2">Author</mods:roleTerm>
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<mods:namePart id="2C076AE41212AC688C38B8ED948ADD99">Bauer, Aaron M.</mods:namePart>
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<mods:typeOfResource id="63E81A96B4BA41939A70644734F7217A">text</mods:typeOfResource>
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<mods:title id="8850A96AC913B1BFB78D088607B985D3">Zootaxa</mods:title>
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<mods:part id="518E24216B92889888703B22A0077285">
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<mods:date id="6685455E5A78D6641D722158214815A9">2014</mods:date>
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<mods:detail id="1CEB7EEAB99BBEB262502D97356EB743" type="volume">
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<mods:number id="60B8BAEAD4D96097BD6D4E6EF9F62465">3846</mods:number>
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<mods:detail id="70085D231B3C9D4E7D33DAB62A793F95" type="issue">
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<mods:number id="F02FB2EC43081B492F798E5F86F1FCE8">4</mods:number>
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<mods:identifier id="8C18D1C9DF5D0007DC345518E430C72E" type="DOI">10.11646/zootaxa.3846.4.1</mods:identifier>
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<mods:identifier id="B3681F12DC072522357BD285AC0DC6E8" type="GBIF-Dataset">317a43cd-c408-4fde-8374-357fbb9a0d53</mods:identifier>
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<mods:identifier id="0A8FBE3F584365C107D3893D39F0715F" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="199C8B524C27E2A75918BE2C7735E985" type="Zenodo-Dep">250495</mods:identifier>
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<treatment id="744387D5B128A109EBADFE45E3F0FB25" ID-DOI="http://doi.org/10.5281/zenodo.5620597" ID-GBIF-Taxon="119514192" ID-Zenodo-Dep="5620597" LSID="urn:lsid:plazi:treatment:744387D5B128A109EBADFE45E3F0FB25" httpUri="http://treatment.plazi.org/id/744387D5B128A109EBADFE45E3F0FB25" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487">
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<subSubSection id="B4F06548B128A108EBADFE45E161FE4A" pageId="36" pageNumber="487" type="nomenclature">
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<paragraph id="FC5536C3B128A108EBADFE45E19BFE2B" blockId="36.[151,497,404,463]" box="[151,497,404,430]" pageId="36" pageNumber="487">
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<heading id="A71D81AFB128A108EBADFE45E19BFE2B" bold="true" box="[151,497,404,430]" fontSize="11" level="1" pageId="36" pageNumber="487" reason="1">
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<emphasis id="CE9EEAD1B128A108EBADFE45E19BFE2B" bold="true" box="[151,497,404,430]" pageId="36" pageNumber="487">
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<taxonomicName id="3BEA4D40B128A108EBADFE45E1E6FE2B" box="[151,396,404,430]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="487" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
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<emphasis id="CE9EEAD1B128A108EBADFE45E1E6FE2B" bold="true" box="[151,396,404,430]" italics="true" pageId="36" pageNumber="487">Afroedura broadleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D5AD57AAB128A108EAA9FE45E19BFE2B" box="[403,497,404,430]" pageId="36" pageNumber="487" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="FC5536C3B128A108EBADFE67E161FE4A" blockId="36.[151,497,404,463]" box="[151,267,438,463]" pageId="36" pageNumber="487">
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(
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<figureCitation id="64D12A46B128A108EBA5FE67E09AFE4A" box="[159,240,438,463]" captionStart="FIGURE 10" captionStartId="34.[151,250,1912,1934]" captionTargetBox="[221,1375,195,1892]" captionTargetId="figure@34.[221,1376,195,1893]" captionTargetPageId="34" captionText="FIGURE 10. New species in the Afroedura langi group. (A) Afroedura granitica sp. nov. from Farm Lillie, Limpopo Province, South Africa (24 ° 04 ’ S, 30 ° 51 ’ E) (Photo Stuart V. Nielsen). (B) Afroedura leoloensis sp. nov. from Farm Maandagshoek, Limpopo Province, South Africa (Photo N. H. G. Jacobsen). (C) Afroedura broadleyi sp. nov. from 22 ° 58 ’ 40 ” S, 29 ° 19 ’ 57 ” E, Limpopo Province, South Africa (Photo Marius Burger)." httpUri="https://zenodo.org/record/250505/files/figure.png" pageId="36" pageNumber="487">Fig. 10</figureCitation>
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C)
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</paragraph>
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</subSubSection>
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||
<subSubSection id="B4F06548B128A108EBADFE2FE22CFDB6" pageId="36" pageNumber="487" type="reference_group">
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<paragraph id="FC5536C3B128A108EBADFE2FE22CFDB6" blockId="36.[151,913,510,563]" pageId="36" pageNumber="487">
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<taxonomicName id="3BEA4D40B128A108EBADFE2FE152FD91" box="[151,312,510,532]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="487" phylum="Chordata" rank="species" species="langi">
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<emphasis id="CE9EEAD1B128A108EBADFE2FE152FD91" box="[151,312,510,532]" italics="true" pageId="36" pageNumber="487">Afroedura langi</emphasis>
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</taxonomicName>
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'Soutpansberg'
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<bibRefCitation id="987B4B32B128A108EAEDFE2FE210FD91" author="Jacobsen" box="[471,634,510,532]" pageId="36" pageNumber="487" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">Jacobsen 1992a</bibRefCitation>
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,
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<bibRefCitation id="987B4B32B128A108E9BEFE2FE2D0FD91" author="Jacobsen" box="[644,698,510,532]" pageId="36" pageNumber="487" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
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;
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<bibRefCitation id="987B4B32B128A108E9FEFE2FE3FBFD91" author="Kirchhof" box="[708,913,510,533]" pageId="36" pageNumber="487" refString="Kirchhof, S., Kramer, M., Linden, J. & Richter, K. (2010) The reptile species assemblage of the Soutpansberg (Limpopo Province, South Africa) and its characterisitcs. Salamandra, 46, 147 - 166." type="journal article" year="2010">
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Kirchhof
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<emphasis id="CE9EEAD1B128A108E81FFDD1E338FD91" box="[805,850,510,533]" italics="true" pageId="36" pageNumber="487">et al</emphasis>
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||
. 2010
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</bibRefCitation>
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||
<taxonomicName id="3BEA4D40B128A108EBA7FDCCE16EFDB6" box="[157,260,541,563]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="487" phylum="Chordata" rank="genus">
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<emphasis id="CE9EEAD1B128A108EBA7FDCCE16EFDB6" box="[157,260,541,563]" italics="true" pageId="36" pageNumber="487">Afroedura</emphasis>
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</taxonomicName>
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'Matlala'
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<bibRefCitation id="987B4B32B128A108EA5DFDCCE260FDB6" author="Jacobsen" box="[359,522,541,563]" pageId="36" pageNumber="487" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">Jacobsen 1992a</bibRefCitation>
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,
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<bibRefCitation id="987B4B32B128A108E929FDCCE22CFDB6" author="Jacobsen" box="[531,582,541,563]" pageId="36" pageNumber="487" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B4F06548B128A108EBADFD8EE32FFC1D" pageId="36" pageNumber="487" type="materials_examined">
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<paragraph id="FC5536C3B128A108EBADFD8EE4E6FD18" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">
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<emphasis id="CE9EEAD1B128A108EBADFD8EE161FDFD" bold="true" box="[151,267,607,632]" pageId="36" pageNumber="487">
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<typeStatus id="23518861B128A108EBADFD8EE16CFDFD" box="[151,262,607,632]" pageId="36" pageNumber="487" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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TM 81316, adult male, Farm Peover 772MS,
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<geoCoordinate id="99DE5004B128A108E80EFD8EE3F3FDFD" box="[820,921,607,632]" direction="south" orientation="latitude" pageId="36" pageNumber="487" precision="925" value="-22.983334">22°59' S</geoCoordinate>
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,
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<geoCoordinate id="99DE5004B128A108E890FDB1E47BFDF2" box="[938,1041,608,632]" direction="east" orientation="longitude" pageId="36" pageNumber="487" precision="925" value="29.733334">29°44' E</geoCoordinate>
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, Soutpansberg District (2229DC), Limpopo Province,
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<collectingCountry id="84FD7653B128A108EA4CFD55E2FEFD18" box="[374,660,644,669]" name="South Africa" pageId="36" pageNumber="487">Republic of South Africa</collectingCountry>
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, collector R. E. Newbery,
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<date id="88541003B128A108E880FD55E4EDFD18" box="[954,1159,644,669]" pageId="36" pageNumber="487" value="1985-11-06">6 November 1985</date>
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.
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</paragraph>
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<paragraph id="FC5536C3B128A108EBFDFD76E4B2FD60" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">
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<emphasis id="CE9EEAD1B128A108EBFDFD76E12CFD45" bold="true" box="[199,326,679,704]" pageId="36" pageNumber="487">
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<typeStatus id="23518861B128A108EBFDFD76E12BFD45" box="[199,321,679,704]" pageId="36" pageNumber="487" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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TM 81317–81322, 81341, same data as for
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<typeStatus id="23518861B128A108E85BFD76E3ADFD45" box="[865,967,679,704]" pageId="36" pageNumber="487" type="holotype">holotype</typeStatus>
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; TM 81338-81340, Farm Leek 769MS, Soutpansberg District (2229DC), Limpopo Province, collector R. E. Newbery
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<date id="88541003B128A108EF3CFD1DE4B9FD60" box="[1030,1235,716,741]" pageId="36" pageNumber="487" value="1985-11-06">6 November 1985</date>
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.
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</paragraph>
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<paragraph id="FC5536C3B128A108EBFDFD3EE32FFC1D" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">
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<emphasis id="CE9EEAD1B128A108EBFDFD3EE241FC8D" bold="true" box="[199,555,751,776]" pageId="36" pageNumber="487">Additional material examined</emphasis>
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(all localities in Limpopo Province). TM 81323-81326, Farm Outlook 789MS, Soutpansberg District (2229DD); TM 81327–81329, 81330-81336, Farm Newgate 802MS, Soutpansberg District (2229DD); TM 81337, Mutshenzheni, Sibasa District (2230CD); TM 81300-81307, 81309-81314, Farm Leipzig 264LR, Bochum District (2328BB); TM 81288–81299, 81308, Farm Urk 10LS, Bochum District (2329AA); TM 81188–81207 Matlala Mountain, Seshego District (2329CC).
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B128A108EBFDFC75E38AFC66" pageId="36" pageNumber="487" type="etymology">
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<paragraph id="FC5536C3B128A108EBFDFC75E38AFC66" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">
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<emphasis id="CE9EEAD1B128A108EBFDFC75E124FC38" bold="true" box="[199,334,932,957]" pageId="36" pageNumber="487">Etymology.</emphasis>
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The species is named for our friend and colleague Dr. Donald G. Broadley in honor of his many contributions to African herpetology and in celebration of his 80th birthday.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B128A108EBFDFC3FE5ECFBF6" pageId="36" pageNumber="487" type="diagnosis">
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<paragraph id="FC5536C3B128A108EBFDFC3FE5ECFBF6" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">
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<emphasis id="CE9EEAD1B128A108EBFDFC3FE12BFB82" bold="true" box="[199,321,1006,1031]" pageId="36" pageNumber="487">Diagnosis.</emphasis>
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A mid-sized
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<taxonomicName id="3BEA4D40B128A108EAD0FC3FE236FB82" box="[490,604,1006,1031]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="36" pageNumber="487" phylum="Chordata" rank="genus">
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<emphasis id="CE9EEAD1B128A108EAD0FC3FE236FB82" box="[490,604,1006,1031]" italics="true" pageId="36" pageNumber="487">Afroedura</emphasis>
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</taxonomicName>
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(maximum SVL 56.0 mm) differing from all other congeners by the following combination of characters: two pairs of enlarged subdigital lamellae per digit; tail moderately verticillate (semi-verticillate) and flattened near base, with four subcaudal rows and 6–8 supracaudal rows per verticil; dorsal scales smooth, 88–106 scale rows at midbody; internasal scales typically absent; 8–12 precloacal pores in males.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B128A109EBFDFBAFE438FE98" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487" type="description">
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||
<paragraph id="FC5536C3B128A108EBFDFBAFE412FAEA" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">
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<emphasis id="CE9EEAD1B128A108EBFDFBAFE133FB12" bold="true" box="[199,345,1150,1175]" pageId="36" pageNumber="487">Description.</emphasis>
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(based on
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<typeStatus id="23518861B128A108EAE3FBAFE251FB12" box="[473,571,1150,1175]" pageId="36" pageNumber="487" type="holotype">holotype</typeStatus>
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TM 81316) Adult male;
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<quantity id="3B129B26B128A108E85AFBAFE3ADFB12" box="[864,967,1150,1175]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" pageId="36" pageNumber="487" unit="mm" value="45.5">45.5 mm</quantity>
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SVL; 49.0 mm TailL (part regenerated); mass before preservation
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<quantity id="3B129B26B128A108EA85FB72E197FB3E" box="[447,509,1187,1211]" metricMagnitude="-3" metricUnit="kg" metricValue="2.3" pageId="36" pageNumber="487" unit="g" value="2.3">2.3 g</quantity>
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. Moderately dorsoventrally depressed; head oval, wider than neck; limbs robust with stout digits; tail depressed, long and tapered, distal portion regenerating. Rostral twice as broad as high; nostril pierced between rostral, first upper labial and three nasals; nasorostrals large and in contact behind rostral. Scales on snout large, heterogeneous and flattened, decreasing in size posteriorly, smallest on crown of head; 10 scales from nasals to eye and 21 scales from eye to ear; three supraciliary spines. Supralabials nine. Mental narrow, wedge-shaped, longer than wide and in contact with two postmentals. Infralabials seven.
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</paragraph>
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<paragraph id="FC5536C3B128A108EBFDFAABE2A3F9C2" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">Dorsal scales relatively homogeneous, rounded,smooth, slightly flattened and juxtaposed; midbody scales 91. Ventral scales larger, imbricate and almost hexagonal. Digits with two pairs of enlarged scansors and one enlarged inferomedian scale under the 4th toe. Precloacal pores 11, arranged in a shallow curve, separated (five and six) by a poreless scale. Tail semi-verticillate with scales arranged in whorls with six dorsal and four ventral rows per verticil. Supracaudals rectangular with a rounded posterior margin; subcaudals squarish with rounded posterior margins. Two to three cloacal spurs at base of tail.</paragraph>
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<paragraph id="FC5536C3B128A108EBFDF983E43EF8C6" blockId="36.[151,1437,607,2003]" pageId="36" pageNumber="487">
|
||
<emphasis id="CE9EEAD1B128A108EBFDF983E17AF9EE" bold="true" box="[199,272,1618,1643]" pageId="36" pageNumber="487">Color.</emphasis>
|
||
Pale grey to buff or pinkish brown above with 6–7 dark brown to blackish, very wavy and irregular crossbands extending from occiput to sacrum. Crossbands darkest posteriorly, incompletely edged with white and with a median white spot. Crown of head heavily spotted or mottled with dark brown; a forward pointing darker ‘V’ extending along the canthus rostralis, whereas a dark streak runs from the nostril through the middle of the eye to above the ear, joining the occipital band. Limbs spotted with white and dark brown. Original tail banded with 10 regular dark brown to blackish crossbars edged with black posteriorly, each followed by a narrow band of white. Venter pale whitish pink; underside of tail mottled and spotted with white and brown.
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B128A109EBFDF89FE438FE98" blockId="36.[151,1437,607,2003]" lastBlockId="37.[151,1437,151,1184]" lastPageId="37" lastPageNumber="488" pageId="36" pageNumber="487">
|
||
<emphasis id="CE9EEAD1B128A108EBFDF89FE151F8E2" bold="true" box="[199,315,1870,1895]" pageId="36" pageNumber="487">Variation.</emphasis>
|
||
<typeStatus id="23518861B128A108EA73F881E1DCF8ED" box="[329,438,1872,1896]" pageId="36" pageNumber="487" type="paratype">Paratypes</typeStatus>
|
||
and other specimens agree with the
|
||
<typeStatus id="23518861B128A108E853F89FE3A1F8E2" box="[873,971,1870,1895]" pageId="36" pageNumber="487" type="holotype">holotype</typeStatus>
|
||
in most features of scalation (
|
||
<tableCitation id="B1680378B128A108EE09F89FE5E5F8E2" box="[1331,1423,1870,1895]" captionStart="TABLE 4" captionStartId="27.[125,205,605,626]" captionText="TABLE 4. variatiοn in key mοrphοlοgical characters in Limpοpο and Mpumalanga Prοvinces Afroedura including, fοr cοmparative purpοses, values fοr A. pondolia and A. major. Fοr size" pageId="36" pageNumber="487">Table 4</tableCitation>
|
||
). Nasorostrals separated by a single granular scale in four (of 75) specimens examined. Scales between nasals and eye 10–13, from eye to ear 15–22. Mental from as long to twice as long as broad; one postmental in 18 of
|
||
<specimenCount id="EAECFD4AB128A108EEBAF846E17AF856" pageId="36" pageNumber="487" type="generic">70 specimens</specimenCount>
|
||
. Supralabials 7–10. Infralabials 6–9. Postmentals usually two, but occasionally 1 or 3 (18 and 2 of 72 examined, respectively). Midbody scale rows 88–106. 0–11 enlarged inferomedian scales under the fourth toe. Precloacal pores in males 8–12 (mostly 9–11) often with a median gap; females lacking pores, although exhibiting a row of somewhat enlarged scales in this position. Original tails 50.0–55.5% of total length. Supracaudal scales in 6–7 (rarely 8) rows per tail whorl. 44.7% of specimens have regenerated tails (n=76).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4F06548B129A109EBFDFEF6E5F7FDB8" pageId="37" pageNumber="488" type="distribution">
|
||
<paragraph id="FC5536C3B129A109EBFDFEF6E136FEE0" blockId="37.[151,1437,151,1184]" pageId="37" pageNumber="488">
|
||
<emphasis id="CE9EEAD1B129A109EBFDFEF6E10AFEC5" bold="true" box="[199,352,295,320]" pageId="37" pageNumber="488">Distribution.</emphasis>
|
||
Isolated populations are found on the Soutpansberg, Blouberg and Matlala inselberg, Limpopo Province (
|
||
<figureCitation id="64D12A46B129A109EA30FE9DE124FEE0" box="[266,334,332,357]" captionStart="FIGURE 6" captionStartId="22.[151,250,1208,1230]" captionTargetBox="[182,1404,196,1182]" captionTargetId="figure@22.[178,1409,193,1187]" captionTargetPageId="22" captionText="FIGURE 6. Map of Limpopo and Mpumalanga provinces, South Africa and adjacent regions showing the distributions of previously described species of Afroedura endemic to this region and known localities of the nine new species described in this paper. For previously recognized forms an approximate outline of the range based on Bates et al. (2014) is shown; genetically sampled localities are marked by circles in Figure 4. For new species all known localities are plotted as dots (see relevant species accounts), with genetically sampled site marked by circles with a a red center, type locality marked by a star, and all other localities indicated by open circles colored to correspond to the key in the figure. The very restricted distributions of A. maripi sp. nov., A. rondavelica sp. nov., and A. rupestris sp. nov. are indicated by arrows and are labeled directly on the map." httpUri="https://zenodo.org/record/250501/files/figure.png" pageId="37" pageNumber="488">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B129A109EBFDFEBEE4C8FE70" blockId="37.[151,1437,151,1184]" pageId="37" pageNumber="488">
|
||
<emphasis id="CE9EEAD1B129A109EBFDFEBEE1EDFE0D" bold="true" box="[199,391,367,392]" pageId="37" pageNumber="488">Natural history.</emphasis>
|
||
The species is nocturnal. It frequents crevices and fissures between rocks as well as under flakes on Waterberg sandstone outcrops, although the Matlala population occurs on granites. May be solitary or can occur in small (presumably family) groups in Soutpansberg Mountain Bushveld (SVcb 21) and Soutpansberg Summit Sourveld (GM 28) (
|
||
<bibRefCitation id="987B4B32B129A109EAEEFE0DE361FE70" author="Mucina" box="[468,779,476,501]" pageId="37" pageNumber="488" refString="Mucina, L. & Rutherford, M. C. (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) at elevations of
|
||
<quantity id="3B129B26B129A109E8F1FE0DE40CFE71" box="[971,1126,476,500]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.7" metricValueMin="1.0" pageId="37" pageNumber="488" unit="m" value="1350.0" valueMax="1700.0" valueMin="1000.0">1000–1700 m</quantity>
|
||
a.s.l.
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B129A109EBADFDD1E5F7FDB8" blockId="37.[151,1437,151,1184]" pageId="37" pageNumber="488">
|
||
Two eggs at a time are laid during midsummer.The eggs adhere to the roof of the crevice before hardening. Communal nesting has been observed in the Soutpansberg where more than
|
||
<specimenCount id="EAECFD4AB129A109E8D7FDF5E429FDB8" box="[1005,1091,548,573]" pageId="37" pageNumber="488" type="egg">20 eggs</specimenCount>
|
||
have been found under a rock.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4F06548B129A109EBFDFD96E3F0FB25" pageId="37" pageNumber="488" type="discussion">
|
||
<paragraph id="FC5536C3B129A109EBFDFD96E0DFFC91" blockId="37.[151,1437,151,1184]" pageId="37" pageNumber="488">
|
||
<emphasis id="CE9EEAD1B129A109EBFDFD96E151FDE5" bold="true" box="[199,315,583,608]" pageId="37" pageNumber="488">Remarks.</emphasis>
|
||
<taxonomicName id="3BEA4D40B129A109EA7CFD96E244FDE5" box="[326,558,583,608]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="broadleyi" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B129A109EA7CFD96E244FDE5" box="[326,558,583,608]" italics="true" pageId="37" pageNumber="488">Afroedura broadleyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB129A109E901FD96E2F3FDE5" box="[571,665,583,608]" pageId="37" pageNumber="488" rank="species">
|
||
<emphasis id="CE9EEAD1B129A109E901FD96E2F3FDE5" bold="true" box="[571,665,583,608]" pageId="37" pageNumber="488">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shares a suite of characters with
|
||
<taxonomicName id="3BEA4D40B129A109EF17FD99E4E6FDE5" box="[1069,1164,583,608]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="langi">
|
||
<emphasis id="CE9EEAD1B129A109EF17FD99E4E6FDE5" box="[1069,1164,583,608]" italics="true" pageId="37" pageNumber="488">A. langi</emphasis>
|
||
</taxonomicName>
|
||
and four other species described herein as new. In comparison with other members of this group it differs most obviously in male precloacal pore counts, being much lower than
|
||
<taxonomicName id="3BEA4D40B129A109E993FD41E352FD2D" box="[681,824,655,680]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="leoloensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B129A109E993FD41E352FD2D" box="[681,824,655,680]" italics="true" pageId="37" pageNumber="488">A. leoloensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB129A109E804FD5EE3F2FD2D" box="[830,920,655,680]" pageId="37" pageNumber="488" rank="species">
|
||
<emphasis id="CE9EEAD1B129A109E804FD5EE3F2FD2D" bold="true" box="[830,920,655,680]" pageId="37" pageNumber="488">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and having non-overlapping pore ranges with
|
||
<emphasis id="CE9EEAD1B129A109EBADFD65E165FD4E" box="[151,271,692,717]" italics="true" pageId="37" pageNumber="488">
|
||
<taxonomicName id="3BEA4D40B129A109EBADFD65E098FD48" box="[151,242,692,717]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="langi">A. langi</taxonomicName>
|
||
,
|
||
<taxonomicName id="3BEA4D40B129A109EBC4FD65E165FD4E" box="[254,271,692,715]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="pienaari" status="sp. nov.">A</taxonomicName>
|
||
</emphasis>
|
||
.
|
||
<emphasis id="CE9EEAD1B129A109EA24FD64E117FD49" box="[286,381,693,716]" italics="true" pageId="37" pageNumber="488">pienaari</emphasis>
|
||
<taxonomicNameLabel id="D5AD57AAB129A109EABEFD65E1B4FD48" box="[388,478,692,717]" pageId="37" pageNumber="488" rank="species">
|
||
<emphasis id="CE9EEAD1B129A109EABEFD65E1B4FD48" bold="true" box="[388,478,692,717]" pageId="37" pageNumber="488">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="3BEA4D40B129A109E92FFD65E2F6FD49" box="[533,668,692,716]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="granitica" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B129A109E92FFD65E24CFD4E" box="[533,550,692,715]" italics="true" pageId="37" pageNumber="488">A</emphasis>
|
||
.
|
||
<emphasis id="CE9EEAD1B129A109E90FFD64E2F6FD49" box="[565,668,693,716]" italics="true" pageId="37" pageNumber="488">granitica</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB129A109E999FD65E297FD48" box="[675,765,692,717]" pageId="37" pageNumber="488" rank="species">
|
||
<emphasis id="CE9EEAD1B129A109E999FD65E297FD48" bold="true" box="[675,765,692,717]" pageId="37" pageNumber="488">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(more pores) and
|
||
<taxonomicName id="3BEA4D40B129A109E8F6FD65E4FFFD48" box="[972,1173,692,717]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="waterbergensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B129A109E8F6FD65E4FFFD48" box="[972,1173,692,717]" italics="true" pageId="37" pageNumber="488">A. waterbergensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB129A109EFA7FD65E49DFD48" box="[1181,1271,692,717]" pageId="37" pageNumber="488" rank="species">
|
||
<emphasis id="CE9EEAD1B129A109EFA7FD65E49DFD48" bold="true" box="[1181,1271,692,717]" pageId="37" pageNumber="488">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(fewer pores). In addition it is substantially larger than
|
||
<taxonomicName id="3BEA4D40B129A109E967FD09E286FD75" box="[605,748,727,752]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="leoloensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B129A109E967FD09E286FD75" box="[605,748,727,752]" italics="true" pageId="37" pageNumber="488">A. leoloensis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="CE9EEAD1B129A109E9C9FD06E331FD75" bold="true" box="[755,859,727,752]" pageId="37" pageNumber="488">
|
||
<taxonomicNameLabel id="D5AD57AAB129A109E9C9FD06E327FD75" box="[755,845,727,752]" pageId="37" pageNumber="488" rank="species">sp. nov.</taxonomicNameLabel>
|
||
,
|
||
</emphasis>
|
||
<emphasis id="CE9EEAD1B129A109E858FD09E3A8FD75" box="[866,962,727,752]" italics="true" pageId="37" pageNumber="488">
|
||
<taxonomicName id="3BEA4D40B129A109E858FD09E3D7FD75" box="[866,957,727,752]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="langi">A. langi</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3BEA4D40B129A109E8C3FD09E4A8FD75" box="[1017,1218,727,752]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="waterbergensis" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B129A109E8C3FD09E4A8FD75" box="[1017,1218,727,752]" italics="true" pageId="37" pageNumber="488">A. waterbergensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB129A109EFF3FD06E549FD75" box="[1225,1315,727,752]" pageId="37" pageNumber="488" rank="species">
|
||
<emphasis id="CE9EEAD1B129A109EFF3FD06E549FD75" bold="true" box="[1225,1315,727,752]" pageId="37" pageNumber="488">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(see
|
||
<tableCitation id="B1680378B129A109EE64FD06E0C2FC91" captionStart="TABLE 4" captionStartId="27.[125,205,605,626]" captionText="TABLE 4. variatiοn in key mοrphοlοgical characters in Limpοpο and Mpumalanga Prοvinces Afroedura including, fοr cοmparative purpοses, values fοr A. pondolia and A. major. Fοr size" pageId="37" pageNumber="488">Table 4</tableCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B129A109EBFDFCCEE3F0FB25" blockId="37.[151,1437,151,1184]" pageId="37" pageNumber="488">
|
||
Three populations are known, each of which appears to differ slightly from the others. However, there is significant overlap in morphological features and Jacobsen’s (1990, 1992a) initial recognition of two putative species (‘Soutpansberg’ and ‘Matlala’) cannot be substantiated given available data. Unfortunately, molecular data were available only from Soutpansberg samples. The Soutpansberg form is separated from the others by
|
||
<taxonomicName id="3BEA4D40B129A109EEBBFC5DE09BFC4D" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="37" pageNumber="488" phylum="Chordata" rank="species" species="pienaari">A. pienaari</taxonomicName>
|
||
along the Waterpoort Gap, whereas those from Matlala are separated by
|
||
<quantity id="3B129B26B129A109EF72FC61E4F9FC4D" box="[1096,1171,943,968]" metricMagnitude="4" metricUnit="m" metricValue="7.0" pageId="37" pageNumber="488" unit="km" value="70.0">70 km</quantity>
|
||
from the nearest other population (on the Blouberg). The Matlala and Blouberg forms have the precloacal pores in a continuous row, only exceptionally subdivided by a poreless scale, as is frequently the case in Soutpansberg populations. Precloacal pores on Matlala and in the Blouberg are mostly eight, nine or 10 and on the Soutpansberg nine, 11 or 12. Individuals from the eastern Blouberg tend to be smaller and have a mean midbody scale count of 91.46 + 2.37 (n = 13), which is considerably lower than that from Soutpansberg individuals—96.91 + 4.32 (n = 22), western Blouberg—97.73 + 3.49 (n = 15) and Matlala—99.5 + 4.19 (n = 18).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |