265 lines
32 KiB
XML
265 lines
32 KiB
XML
<document id="AF632827DA8FBF0C7049E38A7699FA5E" ID-DOI="10.11646/zootaxa.3846.4.1" ID-GBIF-Dataset="317a43cd-c408-4fde-8374-357fbb9a0d53" ID-ISSN="1175-5326" ID-Zenodo-Dep="250495" ID-ZooBank="0DD5A603-D65F-4976-BBE9-94DA7110053F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460467125281" checkinUser="plazi" docAuthor="Jacobsen, Niels H. G., Kuhn, Arianna L., Jackman, Todd R. & Bauer, Aaron M." docDate="2014" docId="744387D5B11BA135EBADFAAEE369FEC5" docLanguage="en" docName="zt03846p501.pdf" docOrigin="Zootaxa 3846 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Afroedura rupestris Jacobsen, Kuhn, Jackman & Bauer, 2014, sp. nov." docType="treatment" docVersion="9" lastPageNumber="475" masterDocId="887AFFADB10CA12CEB3AFFD1E06AFF85" masterDocTitle="A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa" masterLastPageNumber="501" masterPageNumber="451" pageNumber="474" updateTime="1698305158428" updateUser="plazi">
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<mods:titleInfo id="C145DDFA1666EB3BD988880843794435">
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<mods:title id="3D956686F9A610A8BDB28C5975D08915">A phylogenetic analysis of the southern African gecko genus Afroedura Loveridge (Squamata: Gekkonidae), with the description of nine new species from Limpopo and Mpumalanga provinces of South Africa</mods:title>
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</mods:titleInfo>
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<mods:name id="B5BB8CBBD5F65AC8F9BD72D634102ED3" type="personal">
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<mods:roleTerm id="580D38CB4E132960B42A427765828C3A">Author</mods:roleTerm>
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<mods:namePart id="122B063E8158C50A12A0E6676BF80E8A">Jacobsen, Niels H. G.</mods:namePart>
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<mods:roleTerm id="77485A2479D0098EA462252EAD024EB2">Author</mods:roleTerm>
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<mods:namePart id="B5806B2E04B6B404520C1687E26D3A5E">Kuhn, Arianna L.</mods:namePart>
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<mods:name id="B1D49EC3C756DCE8EA8F9833BDDCBAEA" type="personal">
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<mods:roleTerm id="36464DD7D73C2764F6AE551830F93FCE">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="5F15B472A265AC742D2E21697EB281DD">Jackman, Todd R.</mods:namePart>
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</mods:name>
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<mods:name id="D3318FCA9AA88ED030C31ED711753B3E" type="personal">
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<mods:roleTerm id="180CDBB249FDC512D34E83AF7BA6BE56">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="765B72BF0DB3BDFC6AABBB2C7FD741A9">Bauer, Aaron M.</mods:namePart>
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<mods:typeOfResource id="7B6ED57BC7346611BF0CE78A09BBC8B7">text</mods:typeOfResource>
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<mods:title id="E4CFF97D3F402D60005357948FBC5634">Zootaxa</mods:title>
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<mods:part id="BA450E7F34E0958744504F50EDAEBE10">
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<mods:date id="6EA123D4695BF112BA7A1800B0CA08C1">2014</mods:date>
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<mods:detail id="C49F171549B3EF657761DA33B78F6A5C" type="volume">
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<mods:number id="7D11A0D5C2AC41EE91D63B3B015E1D47">3846</mods:number>
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<mods:detail id="98A2BB728583C6D68FCAB77441039F22" type="issue">
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<mods:number id="975FB643D940B52C6B86BF15DB7D99CF">4</mods:number>
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<mods:classification id="5EBC38916ACE71EE9043A6746A7D94C4">journal article</mods:classification>
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<mods:identifier id="6C76330F01635F8D656BBE771607F3CB" type="DOI">10.11646/zootaxa.3846.4.1</mods:identifier>
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<mods:identifier id="00E1ED24E98843AC57F96D041414A8E0" type="GBIF-Dataset">317a43cd-c408-4fde-8374-357fbb9a0d53</mods:identifier>
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<mods:identifier id="CAC1821B392F0443A2EBAB2BC0765EF0" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="F218D8671235DA9217566997B2618568" type="Zenodo-Dep">250495</mods:identifier>
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<mods:identifier id="1676CE7AAB6B90AA2F564FFD94258C33" type="ZooBank">0DD5A603-D65F-4976-BBE9-94DA7110053F</mods:identifier>
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</mods:mods>
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<treatment id="744387D5B11BA135EBADFAAEE369FEC5" ID-DOI="http://doi.org/10.5281/zenodo.5620569" ID-GBIF-Taxon="119514209" ID-Zenodo-Dep="5620569" LSID="urn:lsid:plazi:treatment:744387D5B11BA135EBADFAAEE369FEC5" httpUri="http://treatment.plazi.org/id/744387D5B11BA135EBADFAAEE369FEC5" lastPageId="25" lastPageNumber="475" pageId="23" pageNumber="474">
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<subSubSection id="B4F06548B11BA13BEBADFAAEE14AFA3E" pageId="23" pageNumber="474" type="nomenclature">
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<paragraph id="FC5536C3B11BA13BEBADFAAEE180FA1F" blockId="23.[151,490,1407,1467]" box="[151,490,1407,1434]" pageId="23" pageNumber="474">
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<heading id="A71D81AFB11BA13BEBADFAAEE180FA1F" bold="true" box="[151,490,1407,1434]" fontSize="11" level="1" pageId="23" pageNumber="474" reason="1">
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<emphasis id="CE9EEAD1B11BA13BEBADFAAEE180FA1F" bold="true" box="[151,490,1407,1434]" pageId="23" pageNumber="474">
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<taxonomicName id="3BEA4D40B11BA13BEBADFAAEE1ECFA1C" box="[151,390,1407,1433]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="rupestris" status="sp. nov.">
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<emphasis id="CE9EEAD1B11BA13BEBADFAAEE1ECFA1C" bold="true" box="[151,390,1407,1433]" italics="true" pageId="23" pageNumber="474">Afroedura rupestris</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="D5AD57AAB11BA13BEAB6FA51E180FA1F" box="[396,490,1408,1434]" pageId="23" pageNumber="474" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="FC5536C3B11BA13BEBADFA73E14AFA3E" blockId="23.[151,490,1407,1467]" box="[151,288,1442,1467]" pageId="23" pageNumber="474">
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||
(
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<figureCitation id="64D12A46B11BA13BEBA5FA73E08BFA3E" box="[159,225,1442,1467]" captionStart="FIGURE 7" captionStartId="24.[151,250,839,861]" captionTargetBox="[151,1436,373,817]" captionTargetId="figure@24.[151,1436,373,818]" captionTargetPageId="24" captionText="FIGURE 7. Afroedura rupestris sp. nov. (A) Farm Perkoe, Limpopo Province, South Africa (Photo N. H. G. Jacobsen). (B) Abel Erasmus Pass, Limpopo Province, South Africa (Photo Marius Burger)." httpUri="https://zenodo.org/record/250502/files/figure.png" pageId="23" pageNumber="474">Fig. 7</figureCitation>
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A–B)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B4F06548B11BA13BEBADFA3BE292F985" box="[151,760,1514,1536]" pageId="23" pageNumber="474" type="reference_group">
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<paragraph id="FC5536C3B11BA13BEBADFA3BE292F985" blockId="23.[151,760,1514,1536]" box="[151,760,1514,1536]" pageId="23" pageNumber="474">
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<taxonomicName id="3BEA4D40B11BA13BEBADFA3BE106F985" authority="Abel" authorityName="Abel" box="[151,364,1514,1536]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="474" phylum="Chordata" rank="species" species="multiporis">
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<emphasis id="CE9EEAD1B11BA13BEBADFA3BE106F985" box="[151,364,1514,1536]" italics="true" pageId="23" pageNumber="474">Afroedura multiporis</emphasis>
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</taxonomicName>
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‘Abel Erasmus’
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<bibRefCitation id="987B4B32B11BA13BE923FA3BE2D1F985" author="Jacobsen" box="[537,699,1514,1536]" pageId="23" pageNumber="474" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">Jacobsen 1992a</bibRefCitation>
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,
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<bibRefCitation id="987B4B32B11BA13BE9FFFA3BE292F985" author="Jacobsen" box="[709,760,1514,1536]" pageId="23" pageNumber="474" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
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</paragraph>
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</subSubSection>
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||
<subSubSection id="B4F06548B11BA13BEBADF9FCE25BF953" pageId="23" pageNumber="474" type="materials_examined">
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||
<paragraph id="FC5536C3B11BA13BEBADF9FCE43AF9EF" blockId="23.[151,1436,1581,2038]" pageId="23" pageNumber="474">
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||
<emphasis id="CE9EEAD1B11BA13BEBADF9FCE161F9C3" bold="true" box="[151,267,1581,1606]" pageId="23" pageNumber="474">
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<typeStatus id="23518861B11BA13BEBADF9FCE16CF9C3" box="[151,262,1581,1606]" pageId="23" pageNumber="474" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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TM 81233, adult male, Farm Perkeo 223KT,
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<geoCoordinate id="99DE5004B11BA13BE80FF9FFE3E5F9C3" box="[821,911,1582,1606]" direction="south" orientation="latitude" pageId="23" pageNumber="474" precision="925" value="-24.45">24°27'S</geoCoordinate>
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,
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<geoCoordinate id="99DE5004B11BA13BE89AF9FCE391F9C3" box="[928,1019,1581,1606]" direction="east" orientation="longitude" pageId="23" pageNumber="474" precision="925" value="30.583334">30°35'E</geoCoordinate>
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, Sekukhuneland District (2430BC), Limpopo Province,
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<collectingCountry id="84FD7653B11BA13BEA4CF980E2FEF9EF" box="[374,660,1617,1642]" name="South Africa" pageId="23" pageNumber="474">Republic of South Africa</collectingCountry>
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, collector R.E. Newbery,
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<date id="88541003B11BA13BE889F983E426F9EF" box="[947,1100,1618,1642]" pageId="23" pageNumber="474" value="1982-06-28">28 June 1982</date>
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.
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</paragraph>
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<paragraph id="FC5536C3B11BA13BEBFDF9A4E41EF90B" blockId="23.[151,1436,1581,2038]" box="[199,1140,1653,1678]" pageId="23" pageNumber="474">
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<emphasis id="CE9EEAD1B11BA13BEBFDF9A4E12CF90B" bold="true" box="[199,326,1653,1678]" pageId="23" pageNumber="474">
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<typeStatus id="23518861B11BA13BEBFDF9A4E12BF90B" box="[199,321,1653,1678]" pageId="23" pageNumber="474" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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TM 81224–81228, 81230–81232, 81234–81235, same data as
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<typeStatus id="23518861B11BA13BEF30F9A4E41AF90B" box="[1034,1136,1653,1678]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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<paragraph id="FC5536C3B11BA13BEBFDF948E25BF953" blockId="23.[151,1436,1581,2038]" pageId="23" pageNumber="474">
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<emphasis id="CE9EEAD1B11BA13BEBFDF948E1DCF937" bold="true" box="[199,438,1689,1714]" pageId="23" pageNumber="474">Additional material</emphasis>
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: TM 47504, 52120, 81229, Bourk’s Luck, Pilgrim’s Rest District, Limpopo Province,
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<collectingCountry id="84FD7653B11BA13BEBADF96CE1DBF953" box="[151,433,1725,1750]" name="South Africa" pageId="23" pageNumber="474">Republic of South Africa</collectingCountry>
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(2430DB).
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B11BA13BEBFDF930E17AF898" pageId="23" pageNumber="474" type="etymology">
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<paragraph id="FC5536C3B11BA13BEBFDF930E17AF898" blockId="23.[151,1436,1581,2038]" pageId="23" pageNumber="474">
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<emphasis id="CE9EEAD1B11BA13BEBFDF930E124F97F" bold="true" box="[199,334,1761,1786]" pageId="23" pageNumber="474">Etymology.</emphasis>
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The species name refers to the rupicolous nature of this (and most) members of the genus
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<taxonomicName id="3BEA4D40B11BA13BEBADF8D4E163F89B" box="[151,265,1797,1822]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="474" phylum="Chordata" rank="genus">
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<emphasis id="CE9EEAD1B11BA13BEBADF8D4E163F89B" box="[151,265,1797,1822]" italics="true" pageId="23" pageNumber="474">Afroedura</emphasis>
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</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="B4F06548B11BA13BEBFDF8F8E366F82B" pageId="23" pageNumber="474" type="diagnosis">
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<paragraph id="FC5536C3B11BA13BEBFDF8F8E366F82B" blockId="23.[151,1436,1581,2038]" pageId="23" pageNumber="474">
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<emphasis id="CE9EEAD1B11BA13BEBFDF8F8E12BF8C7" bold="true" box="[199,321,1833,1858]" pageId="23" pageNumber="474">Diagnosis.</emphasis>
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A large
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<taxonomicName id="3BEA4D40B11BA13BEA86F8F8E244F8C7" box="[444,558,1833,1858]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="23" pageNumber="474" phylum="Chordata" rank="genus">
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<emphasis id="CE9EEAD1B11BA13BEA86F8F8E244F8C7" box="[444,558,1833,1858]" italics="true" pageId="23" pageNumber="474">Afroedura</emphasis>
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</taxonomicName>
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(to
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<quantity id="3B129B26B11BA13BE957F8F8E2AFF8C4" box="[621,709,1833,1858]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="23" pageNumber="474" unit="mm" value="62.0">62 mm</quantity>
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SVL) differing from all other congeners by the following combination of characters: two pairs of enlarged subdigital lamellae per digit; tail faintly verticillate with three subcaudal rows and usually six supracaudal rows per verticil; dorsal scales bluntly keeled to trihedral; usually a single internasal scale; 20–24 precloacal pores in males.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="B4F06548B11BA134EBFDF868E5E5F935" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474" type="description">
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||
<paragraph id="FC5536C3B11BA134EBFDF868E2CDFEC5" blockId="23.[151,1436,1581,2038]" lastBlockId="24.[151,1436,151,320]" lastPageId="24" lastPageNumber="475" pageId="23" pageNumber="474">
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<emphasis id="CE9EEAD1B11BA13BEBFDF868E133F857" bold="true" box="[199,345,1977,2002]" pageId="23" pageNumber="474">Description.</emphasis>
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(based on
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<typeStatus id="23518861B11BA13BEAEDF868E253F857" box="[471,569,1977,2002]" pageId="23" pageNumber="474" type="holotype">holotype</typeStatus>
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||
TM 81233) Adult male; SVL 52.0 mm; tail 61.0 mm; mass in life
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<quantity id="3B129B26B11BA13BEE04F86BE511F857" box="[1342,1403,1978,2002]" metricMagnitude="-3" metricUnit="kg" metricValue="3.4" pageId="23" pageNumber="474" unit="g" value="3.4">3.4 g</quantity>
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. A slender elongated, lightly pigmented gecko with a rounded crown to the head. Rostral twice as broad as high, heptagonal; nostril pierced between rostral, first upper labial and three nasal scales; nasorostrals separated by a granular scale behind rostral. Scales on snout larger than on crown with some keeled scales along canthus rostralis and above and slightly anterior to the eyes; 2–3 supraciliary spines. Scales between the nasals and anterior margin of eye 12 and from posterior margin of eye to ear 20. Supralabials nine. Mental wedge-shaped, pentagonal, longer than broad; postmentals two; infralabials eight.
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</paragraph>
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<caption id="A895664BB114A134EBADFC96E3CFFCF9" httpUri="https://zenodo.org/record/250502/files/figure.png" pageId="24" pageNumber="475" targetBox="[151,1436,373,817]" targetPageId="24">
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||
<paragraph id="FC5536C3B114A134EBADFC96E3CFFCF9" blockId="24.[151,1436,839,892]" pageId="24" pageNumber="475">
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||
<emphasis id="CE9EEAD1B114A134EBADFC96E17FFCD9" bold="true" box="[151,277,839,861]" pageId="24" pageNumber="475">FIGURE 7.</emphasis>
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||
<taxonomicName id="3BEA4D40B114A134EA24FC96E182FCD8" box="[286,488,839,861]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="rupestris" status="sp. nov.">
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<emphasis id="CE9EEAD1B114A134EA24FC96E182FCD8" box="[286,488,839,861]" italics="true" pageId="24" pageNumber="475">Afroedura rupestris</emphasis>
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||
</taxonomicName>
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||
<taxonomicNameLabel id="D5AD57AAB114A134EACBFC96E22EFCD9" box="[497,580,839,860]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="CE9EEAD1B114A134EACBFC96E22EFCD9" bold="true" box="[497,580,839,860]" pageId="24" pageNumber="475">sp. nov.</emphasis>
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||
</taxonomicNameLabel>
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(A) Farm Perkoe, Limpopo Province, South Africa (Photo N.H.G. Jacobsen). (B) Abel Erasmus Pass, Limpopo Province, South Africa (Photo Marius Burger).
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||
</paragraph>
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||
</caption>
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||
<paragraph id="FC5536C3B114A134EBFDFC72E25DFB11" blockId="24.[151,1437,931,2036]" pageId="24" pageNumber="475">Dorsals granular, heterogeneous, smaller paravertebrally, larger dorsolaterally, bluntly keeled to rounded, conical and juxtaposed. Scales rows at midbody 87. Ventral scales smooth, larger than dorsals, imbricate. Digits slender with two pairs of enlarged scansors and some enlarged inferomedian scales under the fourth toe. Other digits lacking enlarged inferomedian scales. Precloacal pores in a continuous row of 24, arranged in a chevron; three postcloacal spurs on each side of tail base. Tail long, somewhat flattened proximally; caudal scales arranged in poorly developed verticils with six dorsal and three ventral rows per verticil; dorsal scales smaller than ventrals and subimbricate; ventrals imbricate.</paragraph>
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||
<paragraph id="FC5536C3B114A134EBFDFB4EE1FBFA15" blockId="24.[151,1437,931,2036]" pageId="24" pageNumber="475">
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||
<emphasis id="CE9EEAD1B114A134EBFDFB4EE17AFB3D" bold="true" box="[199,272,1183,1208]" pageId="24" pageNumber="475">Color.</emphasis>
|
||
Olive-brown above with six, rarely seven, blackish wavy and irregular crossbands extending from occiput to sacrum. Pale whitish spots sometimes present at posterior margin of crossbands but may be absent. Limbs pale olive-brown with reticulate darker patterns on the thighs or with wavy stripes and spots. Crown of head paler, mostly without distinct markings. A darkish stripe from nostril to the anterior margin of eye. Two stripes extend from posterior margin of eye to above ear opening. In some individuals a single stripe extends through the eye and is continuous with the occipital bar. Tail with up to 10 crossbars. Body venter whitish, tail venter brownish with darker speckling.
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B114A134EBFDFA4AE5E5F935" blockId="24.[151,1437,931,2036]" pageId="24" pageNumber="475">
|
||
<emphasis id="CE9EEAD1B114A134EBFDFA4AE151FA31" bold="true" box="[199,315,1435,1460]" pageId="24" pageNumber="475">Variation.</emphasis>
|
||
<typeStatus id="23518861B114A134EA73FA4DE1DCFA31" box="[329,438,1436,1460]" pageId="24" pageNumber="475" type="paratype">Paratypes</typeStatus>
|
||
and other specimens agree with the
|
||
<typeStatus id="23518861B114A134E853FA4AE3A1FA31" box="[873,971,1435,1460]" pageId="24" pageNumber="475" type="holotype">holotype</typeStatus>
|
||
in most features of scalation (
|
||
<tableCitation id="B1680378B114A134EE09FA4AE5E5FA31" box="[1331,1423,1435,1460]" captionStart="TABLE 4" captionStartId="27.[125,205,605,626]" captionText="TABLE 4. variatiοn in key mοrphοlοgical characters in Limpοpο and Mpumalanga Prοvinces Afroedura including, fοr cοmparative purpοses, values fοr A. pondolia and A. major. Fοr size" pageId="24" pageNumber="475">Table 4</tableCitation>
|
||
). Single internasal typically present but two in
|
||
<typeStatus id="23518861B114A134E9B5FA11E285FA5D" box="[655,751,1472,1496]" pageId="24" pageNumber="475" type="paratype">paratype</typeStatus>
|
||
TM 81232 and nasorostrals in contact (no internasals) in TM 81234. Scales from eye to ear 16–19. Supraciliary spines four and scales on snout not keeled except above and anterior to the eye in TM 81232. Rostral and mental scales variable in relative proportion and shape. Postmentals generally two, but three in TM 47504, 52120 and 81234 and one in TM 81232. Female specimens lack precloacal pores; males typically with 20–26 pores with the exception of TM 81231 and TM 81225 which have 12, with a series of poreless scales separating left and right pore series. Postcloacal spurs 1–4. Caudal verticils with 6–7 subcaudal rows each. Tail 51.76–54.31% of total length. 45.45 % of individuals (n = 11) exhibit caudal autotomy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4F06548B114A134EBFDF96AE14BF80D" pageId="24" pageNumber="475" type="distribution">
|
||
<paragraph id="FC5536C3B114A134EBFDF96AE26DF97D" blockId="24.[151,1437,931,2036]" pageId="24" pageNumber="475">
|
||
<emphasis id="CE9EEAD1B114A134EBFDF96AE10AF951" bold="true" box="[199,352,1723,1748]" pageId="24" pageNumber="475">Distribution.</emphasis>
|
||
Apparently limited to the Eastern Escarpment of Limpopo Province, between the Abel Erasmus Pass and Bourke’s Luck (
|
||
<figureCitation id="64D12A46B114A134EA8EF90EE192F97D" box="[436,504,1759,1784]" captionStart="FIGURE 6" captionStartId="22.[151,250,1208,1230]" captionTargetBox="[182,1404,196,1182]" captionTargetId="figure@22.[178,1409,193,1187]" captionTargetPageId="22" captionText="FIGURE 6. Map of Limpopo and Mpumalanga provinces, South Africa and adjacent regions showing the distributions of previously described species of Afroedura endemic to this region and known localities of the nine new species described in this paper. For previously recognized forms an approximate outline of the range based on Bates et al. (2014) is shown; genetically sampled localities are marked by circles in Figure 4. For new species all known localities are plotted as dots (see relevant species accounts), with genetically sampled site marked by circles with a a red center, type locality marked by a star, and all other localities indicated by open circles colored to correspond to the key in the figure. The very restricted distributions of A. maripi sp. nov., A. rondavelica sp. nov., and A. rupestris sp. nov. are indicated by arrows and are labeled directly on the map." httpUri="https://zenodo.org/record/250501/files/figure.png" pageId="24" pageNumber="475">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="FC5536C3B114A134EBFDF8D2E14BF80D" blockId="24.[151,1437,931,2036]" pageId="24" pageNumber="475">
|
||
<emphasis id="CE9EEAD1B114A134EBFDF8D2E1E0F899" bold="true" box="[199,394,1795,1820]" pageId="24" pageNumber="475">Natural history.</emphasis>
|
||
An exclusively rupicolous species living in crevices between rocks and under flakes of exfoliating rock along sandstone cliffs in Ohrigstad Mountain Bushveld (SVcb 26) (
|
||
<bibRefCitation id="987B4B32B114A134EF79F8F6E510F8C5" author="Mucina" box="[1091,1402,1831,1856]" pageId="24" pageNumber="475" refString="Mucina, L. & Rutherford, M. C. (2006) The Vegetation of South Africa, Lesotho and Swaziland. Strelitzia, 19. South African National Biodiversity Institute, Pretoria, 807 pp." type="book" year="2006">Mucina & Rutherford 2006</bibRefCitation>
|
||
) at altitudes between
|
||
<quantity id="3B129B26B114A134EA59F89AE19BF8E6" box="[355,497,1867,1892]" metricMagnitude="3" metricUnit="m" metricValue="1.025" metricValueMax="1.2" metricValueMin="0.85" pageId="24" pageNumber="475" unit="m" value="1025.0" valueMax="1200.0" valueMin="850.0">850–1200 m</quantity>
|
||
a.s.l. (
|
||
<figureCitation id="64D12A46B114A134E907F89AE2EAF8E1" box="[573,640,1867,1892]" captionStart="FIGURE 8" captionStartId="25.[151,250,1382,1404]" captionTargetBox="[151,1436,375,1360]" captionTargetId="figure@25.[151,1436,375,1360]" captionTargetPageId="25" captionText="FIGURE 8. Habitats of new species of South African Afroedura spp. (A) General view of Abel Erasmus Pass, Limpopo, habitat of A. rupestris sp. nov. (B) Habitat of A. maripi sp. nov. near the summit of Mariepskop, Mpumalanga Province in heavy mist. (C) Habitat of A. granitica sp. nov. at its type locality on Farm Lillie, Limpopo Province. (D) Habitat of A. pienaari sp. nov. at Waterpoort on the northern edge of the soutpansberg, Limpopo Province. Photos A and C: Stuart V. Nielsen, Photos B and D: Aaron M. Bauer." httpUri="https://zenodo.org/record/250503/files/figure.png" pageId="24" pageNumber="475">Fig. 8</figureCitation>
|
||
A). Also found in kloofs and on rocky Blackreef quartzite outcrops on top of cliffs.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="B4F06548B114A135EBFDF842E369FEC5" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475" type="discussion">
|
||
<paragraph id="FC5536C3B114A135EBFDF842E369FEC5" blockId="24.[151,1437,931,2036]" lastBlockId="25.[151,1436,151,320]" lastPageId="25" lastPageNumber="476" pageId="24" pageNumber="475">
|
||
<emphasis id="CE9EEAD1B114A134EBFDF842E151F829" bold="true" box="[199,315,1939,1964]" pageId="24" pageNumber="475">Remarks.</emphasis>
|
||
<taxonomicName id="3BEA4D40B114A134EA79F842E248F82E" box="[323,546,1939,1964]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="rupestris" status="sp. nov.">
|
||
<emphasis id="CE9EEAD1B114A134EA79F842E248F82E" box="[323,546,1939,1964]" italics="true" pageId="24" pageNumber="475">Afroedura rupestris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="D5AD57AAB114A134E910F842E2EFF829" box="[554,645,1939,1964]" pageId="24" pageNumber="475" rank="species">
|
||
<emphasis id="CE9EEAD1B114A134E910F842E2EFF829" bold="true" box="[554,645,1939,1964]" pageId="24" pageNumber="475">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
shares with
|
||
<taxonomicName id="3BEA4D40B114A134E82CF845E3E7F829" box="[790,909,1939,1964]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="haackei">
|
||
<emphasis id="CE9EEAD1B114A134E82CF845E3E7F829" box="[790,909,1939,1964]" italics="true" pageId="24" pageNumber="475">A. haackei</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="CE9EEAD1B114A134E8A7F845E45DF829" box="[925,1079,1939,1964]" italics="true" pageId="24" pageNumber="475">
|
||
<taxonomicName id="3BEA4D40B114A134E8A7F845E459F829" box="[925,1075,1939,1964]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="multiporis">A. multiporis</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="3BEA4D40B114A134EF4BF845E4BEF82E" box="[1137,1236,1940,1963]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="24" pageNumber="475" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="CE9EEAD1B114A134EF4BF845E4BEF82E" box="[1137,1236,1940,1963]" italics="true" pageId="24" pageNumber="475">A. major</emphasis>
|
||
</taxonomicName>
|
||
bluntly keeled to trihedral dorsal scales and although not sampled in our molecular phylogeny, presumably belongs to the same clade as these taxa (
|
||
<bibRefCitation id="987B4B32B114A134EA01F80AE199F871" author="Jacobsen" box="[315,499,2011,2036]" pageId="24" pageNumber="475" refString="Jacobsen, N. H. G. (1992 a) Flat geckos (genus Afroedura) in the Transvaal. Journal of the Herpetological Association of Africa, 40, 22 - 25. http: // dx. doi. org / 10.1080 / 04416651.1992.9650313" type="journal article" year="1992" yearSuffix="a">Jacobsen 1992a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="987B4B32B114A134E93BF80DE257F871" author="Jacobsen" box="[513,573,2012,2036]" pageId="24" pageNumber="475" refString="Jacobsen, N. H. G. (1997) Family Gekkonidae, Genus Afroedura. In: Van Wyk, J. H. (Ed.), Proceedings of the FitzSimons Commemorative Symposium, South African Lizards: 50 years of Progress and Third H. A. A. Symposium on African Herpetology, Transvaal Museum, Pretoria, South Africa, 11 - 15 October 1993. Herpetological Association of Africa, Stellenbosch, pp. 34 - 39." type="journal article" year="1997">1997</bibRefCitation>
|
||
). It differs from these species in typically having 20–24 preclocal pores in males (
|
||
<emphasis id="CE9EEAD1B115A135EBD1FF48E158FF35" box="[235,306,153,176]" italics="true" pageId="25" pageNumber="476">versus</emphasis>
|
||
<quantity id="3B129B26B115A135EA06FF46E1CBFF35" box="[316,417,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="4.191" metricValueMax="4.318" metricValueMin="4.064" pageId="25" pageNumber="476" unit="in" value="16.5" valueMax="17.0" valueMin="16.0">16–17 in</quantity>
|
||
<taxonomicName id="3BEA4D40B115A135EA91FF49E255FF35" box="[427,575,151,176]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="multiporis">
|
||
<emphasis id="CE9EEAD1B115A135EA91FF49E255FF35" box="[427,575,151,176]" italics="true" pageId="25" pageNumber="476">A. multiporis</emphasis>
|
||
</taxonomicName>
|
||
, generally
|
||
<quantity id="3B129B26B115A135E9FBFF46E34DFF35" box="[705,807,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="6.731" metricValueMax="7.112" metricValueMin="6.35" pageId="25" pageNumber="476" unit="in" value="26.5" valueMax="28.0" valueMin="25.0">25–28 in</quantity>
|
||
<taxonomicName id="3BEA4D40B115A135E80AFF49E3C2FF35" box="[816,936,151,176]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="haackei">
|
||
<emphasis id="CE9EEAD1B115A135E80AFF49E3C2FF35" box="[816,936,151,176]" italics="true" pageId="25" pageNumber="476">A. haackei</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity id="3B129B26B115A135E8D1FF49E43AFF35" box="[1003,1104,151,176]" metricMagnitude="-1" metricUnit="m" metricValue="4.699" metricValueMax="4.826" metricValueMin="4.572" pageId="25" pageNumber="476" unit="in" value="18.5" valueMax="19.0" valueMin="18.0">18–19 in</quantity>
|
||
<taxonomicName id="3BEA4D40B115A135EF63FF49E4D4FF35" box="[1113,1214,152,176]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="CE9EEAD1B115A135EF63FF49E4D4FF35" box="[1113,1214,152,176]" italics="true" pageId="25" pageNumber="476">A. major</emphasis>
|
||
</taxonomicName>
|
||
). It may further be differentiated from
|
||
<taxonomicName id="3BEA4D40B115A135EA4BFF6DE269FF50" box="[369,515,188,213]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="multiporis">
|
||
<emphasis id="CE9EEAD1B115A135EA4BFF6DE269FF50" box="[369,515,188,213]" italics="true" pageId="25" pageNumber="476">A. multiporis</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="3BEA4D40B115A135E900FF6DE2F6FF51" box="[570,668,188,212]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="CE9EEAD1B115A135E900FF6DE2F6FF51" box="[570,668,188,212]" italics="true" pageId="25" pageNumber="476">A. major</emphasis>
|
||
</taxonomicName>
|
||
by its lower number of supracaudal scale rows per vertical (6
|
||
<emphasis id="CE9EEAD1B115A135EE6EFF6CE5F1FF51" box="[1364,1435,189,212]" italics="true" pageId="25" pageNumber="476">versus</emphasis>
|
||
7 and 8, respectively). It is significantly larger (max SVL
|
||
<quantity id="3B129B26B115A135E818FF0EE319FF7D" box="[802,883,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="6.2" pageId="25" pageNumber="476" unit="mm" value="62.0">62 mm</quantity>
|
||
) than
|
||
<taxonomicName id="3BEA4D40B115A135E883FF31E445FF7D" box="[953,1071,223,248]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="haackei">
|
||
<emphasis id="CE9EEAD1B115A135E883FF31E445FF7D" box="[953,1071,223,248]" italics="true" pageId="25" pageNumber="476">A. haackei</emphasis>
|
||
</taxonomicName>
|
||
(max SVL
|
||
<quantity id="3B129B26B115A135EF89FF0EE56DFF7D" box="[1203,1287,223,248]" metricMagnitude="-2" metricUnit="m" metricValue="5.2" pageId="25" pageNumber="476" unit="mm" value="52.0">52 mm</quantity>
|
||
), from which it also has a significantly higher number of supralabials (p <0.05, t28 d.f.) and scales from eye to ear (p <0.001, t25 d.f.), and smaller than
|
||
<taxonomicName id="3BEA4D40B115A135EA84FEF9E24AFEC5" box="[446,544,296,320]" class="Reptilia" family="Gekkonidae" genus="Afroedura" kingdom="Animalia" order="Squamata" pageId="25" pageNumber="476" phylum="Chordata" rank="species" species="major">
|
||
<emphasis id="CE9EEAD1B115A135EA84FEF9E24AFEC5" box="[446,544,296,320]" italics="true" pageId="25" pageNumber="476">A. major</emphasis>
|
||
</taxonomicName>
|
||
(max SVL
|
||
<quantity id="3B129B26B115A135E99EFEF6E29DFEC5" box="[676,759,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="7.6" pageId="25" pageNumber="476" unit="mm" value="76.0">76 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |