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<document id="75846F4CC4307E9C5A276F321E638826" ID-DOI="10.11646/zootaxa.3994.2.4" ID-GBIF-Dataset="afcac252-ff9b-49f0-9c58-b35c9d4d80c2" ID-ISSN="1175-5326" ID-Zenodo-Dep="235688" ID-ZooBank="2E6312C0-FE02-4318-ADC9-1D37F714D0AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461274090604" checkinUser="plazi" docAuthor="Jäger, Peter &amp; Minn, Myin Zu" docDate="2015" docId="600523447F725522F4CCFD2AFCE8F9DA" docLanguage="en" docName="zt03994p252.pdf" docOrigin="Zootaxa 3994 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ctenus natmataung Jäger &amp; Minn, 2015, spec. nov." docType="treatment" docVersion="8" lastPageNumber="245" masterDocId="9C3C5B3C7F775528F45BFFEFFFD8FF94" masterDocTitle="New species in the family Ctenidae Keyserling, 1877 from high altitude habitats in Myanmar, with the first case of penetration of the females cuticle by a male in the RTA-clade (Arachnida: Araneae: Ctenidae)" masterLastPageNumber="252" masterPageNumber="235" pageNumber="240" updateTime="1698612306406" updateUser="plazi">
<mods:mods id="9B3C6D9C727E7D390FE6B1BA096D343C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="02791F4F9BD37CA67EE12B630B5B4D94">
<mods:title id="68B3A712BF88EA4F159B79256BD3A2F8">New species in the family Ctenidae Keyserling, 1877 from high altitude habitats in Myanmar, with the first case of penetration of the females cuticle by a male in the RTA-clade (Arachnida: Araneae: Ctenidae)</mods:title>
</mods:titleInfo>
<mods:name id="90FC28A44D129EA5A96C3561D4B57F52" type="personal">
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<mods:namePart id="54BA58C7A448B1E52C34D4AB1CD399D6">Jäger, Peter</mods:namePart>
</mods:name>
<mods:name id="F19DAA0FA932D909F7007D35BDD271D9" type="personal">
<mods:role id="7157F4B043CD717E6C1516E86C31E1BA">
<mods:roleTerm id="7635494F479A18C08D53677F1EAA4503">Author</mods:roleTerm>
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<mods:namePart id="CBA5F09943D53F4F56C109307738AE6D">Minn, Myin Zu</mods:namePart>
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<mods:typeOfResource id="1A9F224FA6AAA1A58D24F8AA1849420C">text</mods:typeOfResource>
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<mods:title id="F4986E8BD6E2DA7AAF4B314838A7DAD3">Zootaxa</mods:title>
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<mods:part id="ABE8FAF8963FA6370914EBF25567A766">
<mods:date id="24E92EBD42629F26549AFF969F8C5CA2">2015</mods:date>
<mods:detail id="4E1FAA91C938E98FCB06076B40EC2A76" type="volume">
<mods:number id="00F90EC0E684264CAF14F46CF11553E6">3994</mods:number>
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<mods:classification id="AB5CED38B658B337A3A1784BEB68A918">journal article</mods:classification>
<mods:identifier id="E09BC565F257926B1FD9493A3B74CF6B" type="DOI">10.11646/zootaxa.3994.2.4</mods:identifier>
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<treatment id="600523447F725522F4CCFD2AFCE8F9DA" ID-DOI="http://doi.org/10.5281/zenodo.6105742" ID-GBIF-Taxon="119639142" ID-Zenodo-Dep="6105742" LSID="urn:lsid:plazi:treatment:600523447F725522F4CCFD2AFCE8F9DA" httpUri="http://treatment.plazi.org/id/600523447F725522F4CCFD2AFCE8F9DA" lastPageId="10" lastPageNumber="245" pageId="5" pageNumber="240">
<subSubSection id="A0B6C1D97F72552DF4CCFD2AFE4AFC95" pageId="5" pageNumber="240" type="nomenclature">
<paragraph id="E81392527F72552DF4CCFD2AFDDCFD4B" blockId="5.[151,516,709,769]" box="[151,516,709,735]" pageId="5" pageNumber="240">
<heading id="B35B253E7F72552DF4CCFD2AFDDCFD4B" bold="true" box="[151,516,709,735]" fontSize="11" level="1" pageId="5" pageNumber="240" reason="1">
<emphasis id="DAD84E407F72552DF4CCFD2AFDDCFD4B" bold="true" box="[151,516,709,735]" pageId="5" pageNumber="240">
<taxonomicName id="2FACE9D17F72552DF4CCFD2AFE5DFD4B" ID-CoL="6BNMG" box="[151,389,709,735]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F72552DF4CCFD2AFE5DFD4B" bold="true" box="[151,389,709,735]" italics="true" pageId="5" pageNumber="240">Ctenus natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F72552DF5D7FD2AFDDCFD4B" box="[396,516,709,735]" pageId="5" pageNumber="240" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="E81392527F72552DF4CCFD07FE4AFC95" blockId="5.[151,516,709,769]" box="[151,402,744,769]" pageId="5" pageNumber="240">
<figureCitation id="70978ED77F72552DF4CCFD07FEB2FC95" box="[151,362,744,769]" captionStart-0="FIGURES 18 27" captionStart-1="FIGURES 28 30" captionStart-2="FIGURES 31 34" captionStart-3="FIGURES 35 41" captionStart-4="FIGURES 42 43" captionStartId-0="6.[151,264,1795,1818]" captionStartId-1="7.[151,264,1455,1478]" captionStartId-2="8.[151,264,1835,1858]" captionStartId-3="9.[151,264,1910,1933]" captionStartId-4="10.[151,264,807,830]" captionTargetBox-0="[173,1411,279,1748]" captionTargetBox-1="[231,1339,193,1433]" captionTargetBox-2="[150,1436,700,1807]" captionTargetBox-3="[151,1436,219,1889]" captionTargetBox-4="[153,1430,219,769]" captionTargetId-0="figure@6.[169,1415,265,1763]" captionTargetId-1="figure@7.[231,1355,193,1435]" captionTargetId-2="figure@8.[150,1436,697,1814]" captionTargetId-3="figure@9.[151,1436,219,1889]" captionTargetId-4="figure@10.[149,1437,214,776]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="8" captionTargetPageId-3="9" captionTargetPageId-4="10" captionText-0="FIGURES 18 27. Ctenus spp. from Mt Victoria, Myanmar, habitus (18, 20, 22, 24, 26 27 dorsal; 19, 21, 23, 25 ventral). 18 21, 26 27 Ctenus natmataung spec. nov., holotype male (18 19) and female paratype (20 21) 22 25 Ctenus sp. from Mt Victoria, Myanmar (22 23, male; 24 25 female)." captionText-1="FIGURES 28 30 Ctenus natmataung spec. nov., holotype male from Mt Victoria, Myanmar, male palp (28 prolateral; 29 ventral; 30 retrolateral)." captionText-2="FIGURES 31 34 Ctenus natmataung spec. nov., male paratype from Mt Victoria, Myanmar, male palp (31 prolateral; 32 ventral; 33 retrolateral; 34 embolus tip, ventral)." captionText-3="FIGURES 35 41 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, copulatory organ (35, 38, 39, 40 epigyne, ventral; 36 epigyne, posterior; 37 vulva, dorsal; 41 epigyne, lateral). I + II—two chambers of spermathecae, chamber II functionally between chamber I and fertilization duct; EF—epigynal field; EP—separate patches of epigynal field." captionText-4="FIGURES 42 43 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, epigyne with male RTA tip, ventral." httpUri-0="https://zenodo.org/record/235693/files/figure.png" httpUri-1="https://zenodo.org/record/235694/files/figure.png" httpUri-2="https://zenodo.org/record/235695/files/figure.png" httpUri-3="https://zenodo.org/record/235696/files/figure.png" httpUri-4="https://zenodo.org/record/235697/files/figure.png" pageId="5" pageNumber="240">Figs 1821, 2643</figureCitation>
,
<figureCitation id="70978ED77F72552DF52DFD07FE4AFC95" box="[374,402,744,769]" captionStart="FIGURE 63" captionStartId="14.[151,249,1795,1817]" captionTargetBox="[150,1435,872,1772]" captionTargetId="figure@14.[150,1436,872,1773]" captionTargetPageId="14" captionText="FIGURE 63. Distributional records of Ctenidae treated in this paper. Large Map: Records at Nat Ma Taung (= Mt Victoria). Red triangles— Ctenus cladarus, yellow circle— C. pingu spec. nov., green squares— C. natmataung spec. nov., white arrowsummit. White numbers indicate elevation [m]. Inset: Location of Mt Victoria (orange drop), and Mt Popa (blue cross; Anahita popa spec. nov., 1250 m)." httpUri="https://zenodo.org/record/235702/files/figure.png" pageId="5" pageNumber="240">63</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="A0B6C1D97F72552DF4CCFCC4FB88FB14" pageId="5" pageNumber="240" type="materials_examined">
<paragraph id="E81392527F72552DF4CCFCC4FB88FB14" blockId="5.[151,1437,810,2016]" pageId="5" pageNumber="240">
<emphasis id="DAD84E407F72552DF4CCFCC4FD75FCD0" bold="true" box="[151,685,810,836]" pageId="5" pageNumber="240">
<typeStatus id="37172CF07F72552DF4CCFCC4FF0AFCD0" box="[151,210,811,836]" pageId="5" pageNumber="240">Type</typeStatus>
material.
<collectingCountry id="90BBD2C27F72552DF53DFCC4FDDCFCD0" box="[358,516,811,836]" name="Myanmar" pageId="5" pageNumber="240">MYANMAR</collectingCountry>
:
<emphasis id="DAD84E407F72552DF645FCC5FD7CFCD7" bold="true" box="[542,676,810,835]" italics="true" pageId="5" pageNumber="240">
<collectingCountry id="90BBD2C27F72552DF645FCC5FD80FCD7" box="[542,600,810,835]" name="China" pageId="5" pageNumber="240">Chin</collectingCountry>
State
</emphasis>
:
</emphasis>
<typeStatus id="37172CF07F72552DF699FCC4FCF2FCD0" box="[706,810,811,836]" pageId="5" pageNumber="240" type="holotype">Holotype</typeStatus>
male, above Kampetlet, Mountain Oasis Resort,
<geoCoordinate id="8D98F4957F72552DF4CCFCA1FEE8FCF3" box="[151,304,846,871]" direction="north" orientation="latitude" pageId="5" pageNumber="240" precision="1" value="21.197083">21°11'49.5''N</geoCoordinate>
,
<geoCoordinate id="8D98F4957F72552DF51AFCA1FE00FCF3" box="[321,472,846,871]" direction="east" orientation="longitude" pageId="5" pageNumber="240" precision="1" value="94.040276">94°02'25.0''E</geoCoordinate>
,
<quantity id="2F543FB77F72552DF5B1FCA1FD9EFCF2" box="[490,582,846,871]" metricMagnitude="3" metricUnit="m" metricValue="1.716" pageId="5" pageNumber="240" unit="m" value="1716.0">1716 m</quantity>
, secondary forest, by hand, at night, P. Jäger leg.
<date id="9C12B4927F72552DF0F1FCA0FAE6FCF3" box="[1194,1342,847,871]" pageId="5" pageNumber="240" value="2014-05-08">8 May 2014</date>
(
<collectionCode id="8EBD0A977F72552DF108FCA0FA49FCF3" LSID="urn:lsid:biocol.org:col:34838" box="[1363,1425,847,871]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
).
<typeStatus id="37172CF07F72552DF4CCFC9DFEDCFC1E" box="[151,260,882,906]" pageId="5" pageNumber="240" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="FEAA59DB7F72552DF548FC9DFEA2FC1E" box="[275,378,881,906]" pageId="5" pageNumber="240" type="male">17 males</specimenCount>
,
<specimenCount id="FEAA59DB7F72552DF5DEFC9EFDDCFC1E" box="[389,516,881,906]" pageId="5" pageNumber="240" type="female">16 females</specimenCount>
):
<specimenCount id="FEAA59DB7F72552DF64DFC9DFDBBFC1E" box="[534,611,881,906]" pageId="5" pageNumber="240" type="male">1 male</specimenCount>
,
<specimenCount id="FEAA59DB7F72552DF635FC9DFD05FC1E" box="[622,733,881,906]" pageId="5" pageNumber="240" type="female">3 females</specimenCount>
, with same data as
<typeStatus id="37172CF07F72552DF7E8FC9EFBC0FC1E" box="[947,1048,881,906]" pageId="5" pageNumber="240" type="holotype">holotype</typeStatus>
, but
<date id="9C12B4927F72552DF017FC9DFB39FC1E" box="[1100,1249,881,906]" pageId="5" pageNumber="240" value="2015-05-17">17 May 2015</date>
(
<collectionCode id="8EBD0A977F72552DF0ABFC9DFAF6FC1E" LSID="urn:lsid:biocol.org:col:34838" box="[1264,1326,882,906]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
).
<specimenCount id="FEAA59DB7F72552DF164FC9DFA40FC1E" box="[1343,1432,881,906]" pageId="5" pageNumber="240" type="male">3 males</specimenCount>
,
<specimenCount id="FEAA59DB7F72552DF4CCFC7AFF24FC39" box="[151,252,916,941]" pageId="5" pageNumber="240" type="female">1 female</specimenCount>
, with same data as
<typeStatus id="37172CF07F72552DF581FC7BFDE7FC39" box="[474,575,916,941]" pageId="5" pageNumber="240" type="holotype">holotype</typeStatus>
, but
<date id="9C12B4927F72552DF62CFC7AFCD7FC39" box="[631,783,916,941]" pageId="5" pageNumber="240" value="2015-05-10">10 May 2015</date>
(
<collectionCode id="8EBD0A977F72552DF744FC7AFC85FC39" LSID="urn:lsid:biocol.org:col:34838" box="[799,861,917,941]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
).
<specimenCount id="FEAA59DB7F72552DF72AFC7AFC01FC39" box="[881,985,916,941]" pageId="5" pageNumber="240" type="male">12 males</specimenCount>
,
<specimenCount id="FEAA59DB7F72552DF7BEFC7AFB8DFC39" box="[997,1109,916,941]" pageId="5" pageNumber="240" type="female">8 females</specimenCount>
, with same data as
<typeStatus id="37172CF07F72552DF168FC7BFA40FC39" box="[1331,1432,916,941]" pageId="5" pageNumber="240" type="holotype">holotype</typeStatus>
, but
<date id="9C12B4927F72552DF49DFC58FEBCFC44" box="[198,356,951,976]" pageId="5" pageNumber="240" value="2015-05-15">15 May 2015</date>
(
<collectionCode id="8EBD0A977F72552DF52CFC57FE6CFC44" LSID="urn:lsid:biocol.org:col:34838" box="[375,436,952,976]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
;
<specimenCount id="FEAA59DB7F72552DF59FFC57FDF9FC44" box="[452,545,951,976]" pageId="5" pageNumber="240" type="male">2 males</specimenCount>
,
<specimenCount id="FEAA59DB7F72552DF66AFC57FD79FC44" box="[561,673,951,976]" pageId="5" pageNumber="240" type="female">2 females</specimenCount>
UYC;
<specimenCount id="FEAA59DB7F72552DF6A0FC57FC81FC44" box="[763,857,951,976]" pageId="5" pageNumber="240" type="male">2 males</specimenCount>
,
<specimenCount id="FEAA59DB7F72552DF733FC57FC01FC44" box="[872,985,951,976]" pageId="5" pageNumber="240" type="female">2 females</specimenCount>
<collectionCode id="8EBD0A977F72552DF7BFFC57FBE2FC44" box="[996,1082,952,976]" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="5" pageNumber="240">IZCAS</collectionCode>
).
<specimenCount id="FEAA59DB7F72552DF00AFC57FB61FC44" box="[1105,1209,951,976]" pageId="5" pageNumber="240" type="female">1 female</specimenCount>
, with same data as
<typeStatus id="37172CF07F72552DF4CCFC34FF24FC60" box="[151,252,987,1012]" pageId="5" pageNumber="240" type="holotype">holotype</typeStatus>
, but
<date id="9C12B4927F72552DF562FC34FE0EFC60" box="[313,470,987,1012]" pageId="5" pageNumber="240" value="2015-05-12">12 May 2015</date>
(
<collectionCode id="8EBD0A977F72552DF5B2FC34FDFFFC67" LSID="urn:lsid:biocol.org:col:34838" box="[489,551,987,1011]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
).
<specimenCount id="FEAA59DB7F72552DF666FC34FD7DFC60" box="[573,677,987,1012]" pageId="5" pageNumber="240" type="female">1 female</specimenCount>
, above Kampetlet, above Mountain Oasis Resort,
<geoCoordinate id="8D98F4957F72552DF0AAFC34FA4FFC60" box="[1265,1431,987,1012]" direction="north" orientation="latitude" pageId="5" pageNumber="240" precision="1" value="21.198372">21°11'54.14''N</geoCoordinate>
,
<geoCoordinate id="8D98F4957F72552DF4CCFC11FEE3FB83" box="[151,315,1022,1047]" direction="east" orientation="longitude" pageId="5" pageNumber="240" precision="1" value="94.037994">94°02'16.78''E</geoCoordinate>
,
<quantity id="2F543FB77F72552DF511FC11FE7CFB82" box="[330,420,1022,1047]" metricMagnitude="3" metricUnit="m" metricValue="1.752" pageId="5" pageNumber="240" unit="m" value="1752.0">1752 m</quantity>
, disturbed secondary forest, dry slopes,
<date id="9C12B4927F72552DF72EFC10FBC8FB83" box="[885,1040,1022,1047]" pageId="5" pageNumber="240" value="2015-05-12">12 May 2015</date>
(
<collectionCode id="8EBD0A977F72552DF079FC10FBB8FB83" LSID="urn:lsid:biocol.org:col:34838" box="[1058,1120,1023,1047]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
).
<specimenCount id="FEAA59DB7F72552DF02EFC10FB1EFB83" box="[1141,1222,1022,1047]" pageId="5" pageNumber="240" type="male">1 male</specimenCount>
, above Kampetlet, below Mountain Oasis Resort,
<geoCoordinate id="8D98F4957F72552DF5A1FBCEFD4BFBAE" box="[506,659,1057,1082]" direction="north" orientation="latitude" pageId="5" pageNumber="240" precision="1" value="21.190527">21°11'25.9''N</geoCoordinate>
,
<geoCoordinate id="8D98F4957F72552DF6FBFBCDFCE0FBAE" box="[672,824,1058,1082]" direction="east" orientation="longitude" pageId="5" pageNumber="240" precision="1" value="94.03001">94°01'48.0''E</geoCoordinate>
,
<quantity id="2F543FB77F72552DF71EFBCEFC45FBAD" box="[837,925,1057,1082]" metricMagnitude="3" metricUnit="m" metricValue="1.55" pageId="5" pageNumber="240" unit="m" value="1550.0">1550 m</quantity>
, secondary forest along stream, by hand and sieving leaf litter, at day, P. Jäger &amp; J. Martens leg.
<date id="9C12B4927F72552DF6BEFBABFCA4FBC9" box="[741,892,1092,1117]" pageId="5" pageNumber="240" value="2014-05-15">15 May 2014</date>
(
<collectionCode id="8EBD0A977F72552DF7D6FBAAFC13FBC9" LSID="urn:lsid:biocol.org:col:34838" box="[909,971,1093,1117]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
).
<specimenCount id="FEAA59DB7F72552DF784FBAAFB97FBC9" box="[991,1103,1092,1117]" pageId="5" pageNumber="240" type="female">2 females</specimenCount>
, with same data as preceding specimen, but
<geoCoordinate id="8D98F4957F72552DF562FB88FE0BFB14" box="[313,467,1127,1152]" direction="north" orientation="latitude" pageId="5" pageNumber="240" precision="1" value="21.195444">21°11'43.6''N</geoCoordinate>
,
<geoCoordinate id="8D98F4957F72552DF585FB87FDBFFB14" box="[478,615,1128,1152]" direction="east" orientation="longitude" pageId="5" pageNumber="240" precision="1" value="94.03364">94°02'1.1''E</geoCoordinate>
,
<quantity id="2F543FB77F72552DF629FB88FD11FBEB" box="[626,713,1127,1152]" metricMagnitude="3" metricUnit="m" metricValue="1.585" pageId="5" pageNumber="240" unit="m" value="1585.0">1585 m</quantity>
, P. Jäger leg.
<date id="9C12B4927F72552DF739FB87FC2FFB14" box="[866,1015,1128,1152]" pageId="5" pageNumber="240" value="2014-05-17">17 May 2014</date>
(
<collectionCode id="8EBD0A977F72552DF05DFB87FB9CFB14" LSID="urn:lsid:biocol.org:col:34838" box="[1030,1092,1128,1152]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="5" pageNumber="240">SMF</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="A0B6C1D97F72552DF49CFB64FECBFB5C" pageId="5" pageNumber="240" type="etymology">
<paragraph id="E81392527F72552DF49CFB64FECBFB5C" blockId="5.[151,1437,810,2016]" pageId="5" pageNumber="240">
<emphasis id="DAD84E407F72552DF49CFB64FE96FB30" bold="true" box="[199,334,1163,1188]" pageId="5" pageNumber="240">Etymology.</emphasis>
The species name is derived from the local name for Mt Victoria, the
<typeStatus id="37172CF07F72552DF0C7FB63FB14FB30" box="[1180,1228,1164,1188]" pageId="5" pageNumber="240">type</typeStatus>
locality; name in apposition.
</paragraph>
</subSubSection>
<subSubSection id="A0B6C1D97F72552DF49CFB3CFB6DF90F" pageId="5" pageNumber="240" type="diagnosis">
<paragraph id="E81392527F72552DF49CFB3CFB6DF90F" blockId="5.[151,1437,810,2016]" pageId="5" pageNumber="240">
<emphasis id="DAD84E407F72552DF49CFB3CFE99FB78" bold="true" box="[199,321,1235,1260]" pageId="5" pageNumber="240">Diagnosis.</emphasis>
Small to medium-sized
<taxonomicName id="2FACE9D17F72552DF631FB3CFD16FB78" box="[618,718,1235,1260]" class="Arachnida" family="Ctenidae" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="family">Ctenidae</taxonomicName>
(total length males 9.111.8, females 10.416.2). Males are similar to those of
<taxonomicName id="2FACE9D17F72552DF53DFB17FE3DFA9B" box="[358,485,1272,1295]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="ramosus">
<emphasis id="DAD84E407F72552DF53DFB17FE3DFA9B" box="[358,485,1272,1295]" italics="true" pageId="5" pageNumber="240">C. ramosus</emphasis>
</taxonomicName>
,
<taxonomicName id="2FACE9D17F72552DF5A8FB17FDADFA9B" box="[499,629,1271,1295]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="cladarus">
<emphasis id="DAD84E407F72552DF5A8FB17FDADFA9B" box="[499,629,1271,1295]" italics="true" pageId="5" pageNumber="240">C. cladarus</emphasis>
</taxonomicName>
and
<taxonomicName id="2FACE9D17F72552DF6F1FB17FCD2FA9B" box="[682,778,1272,1295]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="pingu" status="sp. nov.">
<emphasis id="DAD84E407F72552DF6F1FB17FCD2FA9B" box="[682,778,1272,1295]" italics="true" pageId="5" pageNumber="240">C. pingu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F72552DF74BFB18FC5AFA84" box="[784,898,1271,1296]" pageId="5" pageNumber="240" rank="species">
<emphasis id="DAD84E407F72552DF74BFB18FC5AFA84" bold="true" box="[784,898,1271,1296]" pageId="5" pageNumber="240">spec. nov.</emphasis>
</taxonomicNameLabel>
in having an easily breaking tip at distal RTA as well as a similar conformation of bulbal structures. Males of
<taxonomicName id="2FACE9D17F72552DF708FAF4FC22FAA7" box="[851,1018,1307,1331]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F72552DF708FAF4FC22FAA7" box="[851,1018,1307,1331]" italics="true" pageId="5" pageNumber="240">C. natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F72552DF05FFAF4FBA0FAA0" box="[1028,1144,1307,1332]" pageId="5" pageNumber="240" rank="species">
<emphasis id="DAD84E407F72552DF05FFAF4FBA0FAA0" bold="true" box="[1028,1144,1307,1332]" pageId="5" pageNumber="240">spec. nov.</emphasis>
</taxonomicNameLabel>
can be recognised by the following combination of characters (
<figureCitation id="70978ED77F72552DF61BFAD0FD1DFAC3" box="[576,709,1343,1368]" captionStart-0="FIGURES 28 30" captionStart-1="FIGURES 31 34" captionStartId-0="7.[151,264,1455,1478]" captionStartId-1="8.[151,264,1835,1858]" captionTargetBox-0="[231,1339,193,1433]" captionTargetBox-1="[150,1436,700,1807]" captionTargetId-0="figure@7.[231,1355,193,1435]" captionTargetId-1="figure@8.[150,1436,697,1814]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 28 30 Ctenus natmataung spec. nov., holotype male from Mt Victoria, Myanmar, male palp (28 prolateral; 29 ventral; 30 retrolateral)." captionText-1="FIGURES 31 34 Ctenus natmataung spec. nov., male paratype from Mt Victoria, Myanmar, male palp (31 prolateral; 32 ventral; 33 retrolateral; 34 embolus tip, ventral)." httpUri-0="https://zenodo.org/record/235694/files/figure.png" httpUri-1="https://zenodo.org/record/235695/files/figure.png" pageId="5" pageNumber="240">Figs 2834</figureCitation>
): 1. RTA tip with long and sharply pointed extension. 2. Tegular apophysis with distal part wide and proximally constricted and distal margin of proximal part transversal. 3. Membranous extension of embolus consisting of two conical parts, the median one distinctly longer than the prolateral one. For further details and a differential diagnosis see under
<taxonomicName id="2FACE9D17F72552DF79DFA44FBF0FA57" box="[966,1064,1451,1475]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="pingu" status="sp. nov.">
<emphasis id="DAD84E407F72552DF79DFA44FBF0FA57" box="[966,1064,1451,1475]" italics="true" pageId="5" pageNumber="240">C. pingu</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F72552DF06BFA44FB7CFA50" box="[1072,1188,1451,1476]" pageId="5" pageNumber="240" rank="species">
<emphasis id="DAD84E407F72552DF06BFA44FB7CFA50" bold="true" box="[1072,1188,1451,1476]" pageId="5" pageNumber="240">spec. nov.</emphasis>
</taxonomicNameLabel>
Copulatory organs of females similar to those of
<taxonomicName id="2FACE9D17F72552DF592FA3FFD91FA73" box="[457,585,1488,1511]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="ramosus">
<emphasis id="DAD84E407F72552DF592FA3FFD91FA73" box="[457,585,1488,1511]" italics="true" pageId="5" pageNumber="240">C. ramosus</emphasis>
</taxonomicName>
,
<taxonomicName id="2FACE9D17F72552DF603FA3FFCA4FA73" authority="Thorell, 1897" authorityName="Thorell" authorityYear="1897" box="[600,892,1487,1512]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="DAD84E407F72552DF603FA3FFD00FA73" box="[600,728,1487,1511]" italics="true" pageId="5" pageNumber="240">C. robustus</emphasis>
Thorell, 1897
</taxonomicName>
,
<taxonomicName id="2FACE9D17F72552DF7DCFA3FFB15FA73" authority="Jager, 2012" authorityName="Jager" authorityYear="2012" box="[903,1229,1487,1512]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="theodorianum">
<emphasis id="DAD84E407F72552DF7DCFA3FFB9DFA73" box="[903,1093,1487,1511]" italics="true" pageId="5" pageNumber="240">C. theodorianum</emphasis>
<bibRefCitation id="8C3DEFA37F72552DF016FA3FFB15FA73" author="Jager" box="[1101,1229,1487,1512]" pageId="5" pageNumber="240" refString="Jager, P. (2012) Asian species of the genera Anahita Karsch 1879, Ctenus Walckenaer 1805 and Amauropelma Raven, Stumkat &amp; Gray 2001 (Arachnida: Araneae: Ctenidae). Zootaxa, 3429, 1 - 63." type="journal article" year="2012">Jäger, 2012</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="2FACE9D17F72552DF15FFA3FFECFF99F" authority="Jager, 2012" authorityName="Jager" authorityYear="2012" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="lishuqiang">
<emphasis id="DAD84E407F72552DF15FFA3FFA44FA73" box="[1284,1436,1487,1511]" italics="true" pageId="5" pageNumber="240">C. lishuqiang</emphasis>
<bibRefCitation id="8C3DEFA37F72552DF4CCFA1BFECFF99F" author="Jager" box="[151,279,1523,1548]" pageId="5" pageNumber="240" refString="Jager, P. (2012) Asian species of the genera Anahita Karsch 1879, Ctenus Walckenaer 1805 and Amauropelma Raven, Stumkat &amp; Gray 2001 (Arachnida: Araneae: Ctenidae). Zootaxa, 3429, 1 - 63." type="journal article" year="2012">
Jäger,
<quantity id="2F543FB77F72552DF484FA1CFECFF99F" box="[223,279,1523,1547]" metricMagnitude="1" metricUnit="m" metricValue="5.11048" pageId="5" pageNumber="240" unit="in" value="2012.0">2012</quantity>
</bibRefCitation>
</taxonomicName>
in having the anterior part of the median plate distinctly narrower than the posterior part (
<figureCitation id="70978ED77F72552DF156FA1CFA49F998" box="[1293,1425,1523,1548]" captionStart-0="FIGURES 35 41" captionStart-1="FIGURES 42 43" captionStartId-0="9.[151,264,1910,1933]" captionStartId-1="10.[151,264,807,830]" captionTargetBox-0="[151,1436,219,1889]" captionTargetBox-1="[153,1430,219,769]" captionTargetId-0="figure@9.[151,1436,219,1889]" captionTargetId-1="figure@10.[149,1437,214,776]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 35 41 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, copulatory organ (35, 38, 39, 40 epigyne, ventral; 36 epigyne, posterior; 37 vulva, dorsal; 41 epigyne, lateral). I + II—two chambers of spermathecae, chamber II functionally between chamber I and fertilization duct; EF—epigynal field; EP—separate patches of epigynal field." captionText-1="FIGURES 42 43 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, epigyne with male RTA tip, ventral." httpUri-0="https://zenodo.org/record/235696/files/figure.png" httpUri-1="https://zenodo.org/record/235697/files/figure.png" pageId="5" pageNumber="240">Figs 3543</figureCitation>
). Epigynes of
<taxonomicName id="2FACE9D17F72552DF572F9F7FE0AF9BB" box="[297,466,1560,1583]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F72552DF572F9F7FE0AF9BB" box="[297,466,1560,1583]" italics="true" pageId="5" pageNumber="240">C. natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F72552DF587F9F8FD89F9A4" box="[476,593,1559,1584]" pageId="5" pageNumber="240" rank="species">
<emphasis id="DAD84E407F72552DF587F9F8FD89F9A4" bold="true" box="[476,593,1559,1584]" pageId="5" pageNumber="240">spec. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished from those of
<taxonomicName id="2FACE9D17F72552DF7A8F9F7FBADF9BB" box="[1011,1141,1559,1583]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="DAD84E407F72552DF7A8F9F7FBADF9BB" box="[1011,1141,1559,1583]" italics="true" pageId="5" pageNumber="240">C. robustus</emphasis>
</taxonomicName>
,
<taxonomicName id="2FACE9D17F72552DF0DDF9F7FA9FF9BB" box="[1158,1351,1559,1583]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="theodorianum">
<emphasis id="DAD84E407F72552DF0DDF9F7FA9FF9BB" box="[1158,1351,1559,1583]" italics="true" pageId="5" pageNumber="240">C. theodorianum</emphasis>
</taxonomicName>
and
<taxonomicName id="2FACE9D17F72552DF1D9F9F7FED5F9C7" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="lishuqiang">
<emphasis id="DAD84E407F72552DF1D9F9F7FED5F9C7" italics="true" pageId="5" pageNumber="240">C. lishuqiang</emphasis>
</taxonomicName>
by the presence of two furrows within the median plate, and from those of
<taxonomicName id="2FACE9D17F72552DF038F9D4FB3BF9C7" box="[1123,1251,1595,1619]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="ramosus">
<emphasis id="DAD84E407F72552DF038F9D4FB3BF9C7" box="[1123,1251,1595,1619]" italics="true" pageId="5" pageNumber="240">C. ramosus</emphasis>
</taxonomicName>
by having these not as converging as in
<taxonomicName id="2FACE9D17F72552DF5F4F98FFDEAF9E3" box="[431,562,1632,1655]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="ramosus">
<emphasis id="DAD84E407F72552DF5F4F98FFDEAF9E3" box="[431,562,1632,1655]" italics="true" pageId="5" pageNumber="240">C. ramosus</emphasis>
</taxonomicName>
(i.e. posterior ends of the additional furrows posteriad, not laterad as in
<taxonomicName id="2FACE9D17F72552DF1D9F98FFF2EF90F" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="ramosus">
<emphasis id="DAD84E407F72552DF1D9F98FFF2EF90F" italics="true" pageId="5" pageNumber="240">C. ramosus</emphasis>
</taxonomicName>
). Moreover, epigynal teeth not bending ventrally over median plate as in
<taxonomicName id="2FACE9D17F72552DF075F96CFB76F90F" box="[1070,1198,1667,1691]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="5" pageNumber="240" phylum="Arthropoda" rank="species" species="ramosus">
<emphasis id="DAD84E407F72552DF075F96CFB76F90F" box="[1070,1198,1667,1691]" italics="true" pageId="5" pageNumber="240">C. ramosus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="A0B6C1D97F725522F49CF948FC1FFA0D" lastPageId="10" lastPageNumber="245" pageId="5" pageNumber="240" type="description">
<paragraph id="E81392527F72552EF49CF948FBD0FF40" blockId="5.[151,1437,810,2016]" lastBlockId="6.[151,1436,151,213]" lastPageId="6" lastPageNumber="241" pageId="5" pageNumber="240">
<emphasis id="DAD84E407F72552DF49CF948FE81F954" bold="true" box="[199,345,1703,1728]" pageId="5" pageNumber="240">Description.</emphasis>
Male (
<typeStatus id="37172CF07F72552DF5F0F948FDCCF954" box="[427,532,1703,1728]" pageId="5" pageNumber="240" type="holotype">holotype</typeStatus>
). PL 5.8, PW 4.5, AW 2.2, OL 4.2, OW 2.8. Eye diameters and interdistances: AME 0.28, ALE 0.20, PME 0.41, PLE 0.40, AMEAME 0.14, AMEALE 0.32, PMEPME 0.12, PMEPLE 0.30, AMEPME 0.06, ALEPLE 0.11, clypeus height at AME 0.15, clypeus height at ALE 0.45. Palp and leg measurements: palp 8.2 (3.0, 1.1, 1.3, -, 1.8), I 19.2 (4.9, 2.3, 5.0, 5.0, 2.0), II 16.9 (4.6, 2.1, 4.3, 4.3, 1.6), III 14.7 (4.1, 1.8, 3.6, 3.8, 1.4), IV 20.7 (5.3, 2.0, 5.2, 6.4, 1.8). Leg formula 4123. Spination of palp and legs: palp 151, 100, 1111; femora I p003, d111, r112, IIIII p112, d111, r112, IV p112, d111, r012; patellae 101; tibiae III p010, d111, r110, v22222, IIIIV p11, d111, r11, v222; metatarsi I p111, r111, v222, II p111, d00(1)0, r111, v222, III p111, d012, r111, v222, IV p111, d012, r111,
<date id="9C12B4927F72552DF6CBF84CFD02F82F" box="[656,730,1955,1979]" pageId="5" pageNumber="240" value="2122-05">v2122</date>
. Chelicerae with 3 promarginal, 4 retromarginal teeth, and with c. 20 denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 1 strong bristle and 6 thinner bristles. Coxae ventrally with stout setae, increasingly distinct from I to IV, leaving on each coxa proxiomedian patch free of setae. Sparse scopula on all tarsi and distal metatarsi III.
</paragraph>
<caption id="BCD3C2DA7F71552EF4CCF8ECFD52F8C3" httpUri="https://zenodo.org/record/235693/files/figure.png" pageId="6" pageNumber="241" targetBox="[173,1411,279,1748]" targetPageId="6">
<paragraph id="E81392527F71552EF4CCF8ECFD52F8C3" blockId="6.[151,1437,1795,1879]" pageId="6" pageNumber="241">
<emphasis id="DAD84E407F71552EF4CCF8ECFE0EF88D" bold="true" box="[151,470,1795,1818]" pageId="6" pageNumber="241">
FIGURES 1827.
<taxonomicName id="2FACE9D17F71552EF501F8ECFE7BF88D" box="[346,419,1795,1817]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="DAD84E407F71552EF501F8ECFE7BF88D" bold="true" box="[346,419,1795,1817]" italics="true" pageId="6" pageNumber="241">Ctenus</emphasis>
</taxonomicName>
spp.
</emphasis>
from Mt Victoria, Myanmar, habitus (18, 20, 22, 24, 2627 dorsal; 19, 21, 23, 25 ventral).
<emphasis id="DAD84E407F71552EF12DF8ECFD92F8A3" bold="true" pageId="6" pageNumber="241">
18 21, 2627
<taxonomicName id="2FACE9D17F71552EF55CF8CDFE00F8AC" box="[263,472,1826,1848]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F71552EF55CF8CDFE00F8AC" bold="true" box="[263,472,1826,1848]" italics="true" pageId="6" pageNumber="241">Ctenus natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F71552EF5BAF8CDFD92F8A3" box="[481,586,1826,1847]" pageId="6" pageNumber="241" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (1819) and female paratype (2021)
<emphasis id="DAD84E407F71552EF0D8F8CDFAE3F8A3" bold="true" box="[1155,1339,1826,1848]" pageId="6" pageNumber="241">
2225
<taxonomicName id="2FACE9D17F71552EF090F8CDFACCF8AC" box="[1227,1300,1826,1848]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="6" pageNumber="241" phylum="Arthropoda" rank="genus">
<emphasis id="DAD84E407F71552EF090F8CDFACCF8AC" bold="true" box="[1227,1300,1826,1848]" italics="true" pageId="6" pageNumber="241">Ctenus</emphasis>
</taxonomicName>
sp.
</emphasis>
from Mt Victoria, Myanmar (2223, male; 2425 female).
</paragraph>
</caption>
<caption id="BCD3C2DA7F70552FF4CCFA40FE51FA71" httpUri="https://zenodo.org/record/235694/files/figure.png" pageId="7" pageNumber="242" targetBox="[231,1339,193,1433]" targetPageId="7">
<paragraph id="E81392527F70552FF4CCFA40FE51FA71" blockId="7.[151,1435,1455,1509]" pageId="7" pageNumber="242">
<emphasis id="DAD84E407F70552FF4CCFA40FD43FA51" bold="true" box="[151,667,1455,1478]" pageId="7" pageNumber="242">
FIGURES 2830
<taxonomicName id="2FACE9D17F70552FF503FA5FFDF1FA52" box="[344,553,1456,1478]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="7" pageNumber="242" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F70552FF503FA5FFDF1FA52" bold="true" box="[344,553,1456,1478]" italics="true" pageId="7" pageNumber="242">Ctenus natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F70552FF669FA5FFD43FA51" box="[562,667,1456,1477]" pageId="7" pageNumber="242" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, holotype male from Mt Victoria, Myanmar, male palp (28 prolateral; 29 ventral; 30 retrolateral).
</paragraph>
</caption>
<paragraph id="E81392527F70552FF49CF9C4FD56F940" blockId="7.[151,1437,1579,2038]" pageId="7" pageNumber="242">
Palp as in diagnosis (
<figureCitation id="70978ED77F70552FF5E1F9C4FDE6F9D7" box="[442,574,1579,1604]" captionStart-0="FIGURES 28 30" captionStart-1="FIGURES 31 34" captionStartId-0="7.[151,264,1455,1478]" captionStartId-1="8.[151,264,1835,1858]" captionTargetBox-0="[231,1339,193,1433]" captionTargetBox-1="[150,1436,700,1807]" captionTargetId-0="figure@7.[231,1355,193,1435]" captionTargetId-1="figure@8.[150,1436,697,1814]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="FIGURES 28 30 Ctenus natmataung spec. nov., holotype male from Mt Victoria, Myanmar, male palp (28 prolateral; 29 ventral; 30 retrolateral)." captionText-1="FIGURES 31 34 Ctenus natmataung spec. nov., male paratype from Mt Victoria, Myanmar, male palp (31 prolateral; 32 ventral; 33 retrolateral; 34 embolus tip, ventral)." httpUri-0="https://zenodo.org/record/235694/files/figure.png" httpUri-1="https://zenodo.org/record/235695/files/figure.png" pageId="7" pageNumber="242">Figs 2834</figureCitation>
). RTA of left palp with, of right palp without distal part. Cymbium with tip slightly conical, retro-proximally with shallow pit. Embolus arising at 7:30- to 8-oclock-position from tegulum, semicircular in ventral view, with broad base, its tip slightly indented and situated in distal half of tegulum. Conductor arising at 12- to 12:30oclock-position subdistally. Tegular apophysis arising at 6-oclock-position from tegulum, shallowly excavated on dorsal side.
</paragraph>
<paragraph id="E81392527F705520F49CF931FBF1FF6C" blockId="7.[151,1437,1579,2038]" lastBlockId="8.[151,1436,151,645]" lastPageId="8" lastPageNumber="243" pageId="7" pageNumber="242">
Colouration (
<figureCitation id="70978ED77F70552FF504F931FDCCF963" box="[351,532,1758,1783]" captionStart="FIGURES 18 27" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[173,1411,279,1748]" captionTargetId="figure@6.[169,1415,265,1763]" captionTargetPageId="6" captionText="FIGURES 18 27. Ctenus spp. from Mt Victoria, Myanmar, habitus (18, 20, 22, 24, 26 27 dorsal; 19, 21, 23, 25 ventral). 18 21, 26 27 Ctenus natmataung spec. nov., holotype male (18 19) and female paratype (20 21) 22 25 Ctenus sp. from Mt Victoria, Myanmar (22 23, male; 24 25 female)." httpUri="https://zenodo.org/record/235693/files/figure.png" pageId="7" pageNumber="242">Figs 1819, 26</figureCitation>
). Yellowish- to reddish-brown with slightly darker markings. Dorsal prosoma with light median band consisting of white dense hairs, laterally with radial marble-like pattern in dark lateral bands; distinctly marked fovea, in 2nd and 3rd third of prosoma. Sternum and ventral coxae yellowish-brown with faint dark marking; labium and gnathocoxae greyish-brown, the latter with light distal parts. Chelicerae brown. Legs yellowish-brown with femora dorsally darker, femora ventrally with indistinct banded pattern. Opisthosoma dorsally with light median area and lateral dark patches in anterior half; posterior half with pairs of dark spots in light median band; laterally irregularly dotted; ventrally dark with four posteriorly converging rows of light muscle sigilla; epiandrum light. Spinnerets generally light yellowish-brown, anterior lateral spinnerets ventrally dark, posterior lateral spinnerets dorsally slightly darker. For colouration pattern of a living specimen see
<figureCitation id="70978ED77F7F5520F15DFF78FA81FF24" box="[1286,1369,151,176]" captionStart="FIGURES 18 27" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[173,1411,279,1748]" captionTargetId="figure@6.[169,1415,265,1763]" captionTargetPageId="6" captionText="FIGURES 18 27. Ctenus spp. from Mt Victoria, Myanmar, habitus (18, 20, 22, 24, 26 27 dorsal; 19, 21, 23, 25 ventral). 18 21, 26 27 Ctenus natmataung spec. nov., holotype male (18 19) and female paratype (20 21) 22 25 Ctenus sp. from Mt Victoria, Myanmar (22 23, male; 24 25 female)." httpUri="https://zenodo.org/record/235693/files/figure.png" pageId="8" pageNumber="243">Fig. 26</figureCitation>
. Note the continuous wide light band at the margin of the dark lateral band of the prosoma dorsally. This character might prove as diagnostic when larger series with this character being stable are known.
</paragraph>
<paragraph id="E81392527F7F5520F49CFEEBFC08FD11" blockId="8.[151,1436,151,645]" pageId="8" pageNumber="243">
<emphasis id="DAD84E407F7F5520F49CFEEBFEC6FE89" bold="true" box="[199,286,260,285]" pageId="8" pageNumber="243">Female</emphasis>
(
<typeStatus id="37172CF07F7F5520F574FEEBFE4CFE88" box="[303,404,260,284]" pageId="8" pageNumber="243" type="paratype">paratype</typeStatus>
, collected with
<typeStatus id="37172CF07F7F5520F616FEEBFD77FE89" box="[589,687,260,285]" pageId="8" pageNumber="243" type="holotype">holotype</typeStatus>
<date id="9C12B4927F7F5520F6E2FEEBFC82FE89" box="[697,858,260,285]" pageId="8" pageNumber="243" value="2015-05-12">12 May 2015</date>
). PL 6.3, PW 4.9, AW 3.1, OL 6.3, OW 4.0. Eye diameters and interdistances: AME 0.29, ALE 0.24, PME 0.41, PLE 0.38, AMEAME 0.18, AMEALE 0.40, PMEPME 0.18, PMEPLE 0.36, AMEPME 0.13, ALEPLE 0.14, clypeus height at AME 0.21, clypeus height at ALE 0.58. Palp and leg measurements: palp 6.2 (2.2, 1.1, 1.4, -, 1.5), I 15.0 (4.2, 2.4, 3.9, 3.3, 1.2), II 14.2 (4.1, 2.2, 3.9, 3.3, 1.2), III 12.8 (3.6, 1.9, 2.9, 3.1, 1.3), IV 17.9 (4.7, 2.1, 4.3, 5.2, 1.6). Leg formula 4123. Spination of palp and legs: palp 141, 100, 1101, 1013; femora I p003, d111, r110, II p112, d111, r110, III p112, d111, r112, IV p112, d111, r001; patellae III 0 0 0, IIIIV 101; tibiae III v22222, IIIIV p11, d111, r11, v222; metatarsi III v222, III p111, d012, r111, v222, IV p111, d012, r111,
<date id="9C12B4927F7F5520F6E5FDEFFCD0FD8C" box="[702,776,512,536]" pageId="8" pageNumber="243" value="2122-05">v2122</date>
. Chelicerae with 3 promarginal, 4 retromarginal teeth, and with elongated patch of 3033 denticles along entire cheliceral furrow. Retromargin of chelicerae close to fang base with 1 slightly thicker bristle and 7 thin bristles. Coxae ventrally with thin setae. Tarsi and metatarsi IIII distally with sparse scopula ventrally. Palpal claw with 5 secondary teeth.
</paragraph>
<caption id="BCD3C2DA7F7F5520F4CCF8C4FD58F8F4" httpUri="https://zenodo.org/record/235695/files/figure.png" pageId="8" pageNumber="243" targetBox="[150,1436,700,1807]" targetPageId="8">
<paragraph id="E81392527F7F5520F4CCF8C4FD58F8F4" blockId="8.[151,1436,1835,1888]" pageId="8" pageNumber="243">
<emphasis id="DAD84E407F7F5520F4CCF8C4FD43F8D5" bold="true" box="[151,667,1835,1858]" pageId="8" pageNumber="243">
FIGURES 3134
<taxonomicName id="2FACE9D17F7F5520F502F8C3FDF1F8D6" box="[345,553,1836,1858]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="243" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F7F5520F502F8C3FDF1F8D6" bold="true" box="[345,553,1836,1858]" italics="true" pageId="8" pageNumber="243">Ctenus natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F7F5520F669F8C3FD43F8D5" box="[562,667,1836,1857]" pageId="8" pageNumber="243" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, male paratype from Mt Victoria, Myanmar, male palp (31 prolateral; 32 ventral; 33 retrolateral; 34 embolus tip, ventral).
</paragraph>
</caption>
<caption id="BCD3C2DA7F7E5521F4CCF899FA4CF85E" httpUri="https://zenodo.org/record/235696/files/figure.png" pageId="9" pageNumber="244" targetBox="[151,1436,219,1889]" targetPageId="9">
<paragraph id="E81392527F7E5521F4CCF899FA4CF85E" blockId="9.[151,1436,1910,1994]" pageId="9" pageNumber="244">
<emphasis id="DAD84E407F7E5521F4CCF899FD56F818" bold="true" box="[151,654,1910,1933]" pageId="9" pageNumber="244">
FIGURES 3541
<taxonomicName id="2FACE9D17F7E5521F508F899FDF9F818" box="[339,545,1910,1932]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="9" pageNumber="244" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F7E5521F508F899FDF9F818" bold="true" box="[339,545,1910,1932]" italics="true" pageId="9" pageNumber="244">Ctenus natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F7E5521F67CF898FD56F818" box="[551,654,1911,1932]" pageId="9" pageNumber="244" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, female paratypes from Mt Victoria, Myanmar, copulatory organ (35, 38, 39, 40 epigyne, ventral; 36 epigyne, posterior; 37 vulva, dorsal; 41 epigyne, lateral). I + II—two chambers of spermathecae, chamber II functionally between chamber I and fertilization duct; EF—epigynal field; EP—separate patches of epigynal field.
</paragraph>
</caption>
<caption id="BCD3C2DA7F7D5522F4CCFCC8FF3DFCC8" httpUri="https://zenodo.org/record/235697/files/figure.png" pageId="10" pageNumber="245" targetBox="[153,1430,219,769]" targetPageId="10">
<paragraph id="E81392527F7D5522F4CCFCC8FF3DFCC8" blockId="10.[151,1436,807,860]" pageId="10" pageNumber="245">
<emphasis id="DAD84E407F7D5522F4CCFCC8FD4BFCA9" bold="true" box="[151,659,807,830]" pageId="10" pageNumber="245">
FIGURES 4243
<taxonomicName id="2FACE9D17F7D5522F50EFCC8FDFBFCA9" box="[341,547,807,829]" class="Arachnida" family="Ctenidae" genus="Ctenus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="245" phylum="Arthropoda" rank="species" species="natmataung" status="sp. nov.">
<emphasis id="DAD84E407F7D5522F50EFCC8FDFBFCA9" bold="true" box="[341,547,807,829]" italics="true" pageId="10" pageNumber="245">Ctenus natmataung</emphasis>
</taxonomicName>
<taxonomicNameLabel id="C1EBF33B7F7D5522F670FCC7FD4BFCA9" box="[555,659,808,829]" pageId="10" pageNumber="245" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
, female paratypes from Mt Victoria, Myanmar, epigyne with male RTA tip, ventral.
</paragraph>
</caption>
<paragraph id="E81392527F7D5522F49CFC67FAE2FBA5" blockId="10.[151,1436,904,1614]" pageId="10" pageNumber="245">
Copulatory organ as in diagnosis (
<figureCitation id="70978ED77F7D5522F614FC67FD0DFC35" box="[591,725,904,929]" captionStart-0="FIGURES 35 41" captionStart-1="FIGURES 42 43" captionStartId-0="9.[151,264,1910,1933]" captionStartId-1="10.[151,264,807,830]" captionTargetBox-0="[151,1436,219,1889]" captionTargetBox-1="[153,1430,219,769]" captionTargetId-0="figure@9.[151,1436,219,1889]" captionTargetId-1="figure@10.[149,1437,214,776]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 35 41 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, copulatory organ (35, 38, 39, 40 epigyne, ventral; 36 epigyne, posterior; 37 vulva, dorsal; 41 epigyne, lateral). I + II—two chambers of spermathecae, chamber II functionally between chamber I and fertilization duct; EF—epigynal field; EP—separate patches of epigynal field." captionText-1="FIGURES 42 43 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, epigyne with male RTA tip, ventral." httpUri-0="https://zenodo.org/record/235696/files/figure.png" httpUri-1="https://zenodo.org/record/235697/files/figure.png" pageId="10" pageNumber="245">Figs 3843</figureCitation>
). Epigyne wider than long, antero-laterally with two indistinct separate patches, laterally with three slit sensilla (2 left, 1 right). Median plate roughly trapezoid, epigynal teeth arising posterio-laterally. Longitudinal furrows dividing median plate in vase-shaped median part and two slightly bent lateral parts. Internal duct system with lateral folds diverging posteriorly. Longitudinal ovoid spermathecae, internally with a large anterior and small posterior chamber, fertilisation ducts arising posteriorly, mediad.
</paragraph>
<paragraph id="E81392527F7D5522F49CFBD2FF32FA9D" blockId="10.[151,1436,904,1614]" pageId="10" pageNumber="245">
Colouration (
<figureCitation id="70978ED77F7D5522F506FBD2FDD4FBC1" box="[349,524,1085,1110]" captionStart="FIGURES 18 27" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[173,1411,279,1748]" captionTargetId="figure@6.[169,1415,265,1763]" captionTargetPageId="6" captionText="FIGURES 18 27. Ctenus spp. from Mt Victoria, Myanmar, habitus (18, 20, 22, 24, 26 27 dorsal; 19, 21, 23, 25 ventral). 18 21, 26 27 Ctenus natmataung spec. nov., holotype male (18 19) and female paratype (20 21) 22 25 Ctenus sp. from Mt Victoria, Myanmar (22 23, male; 24 25 female)." httpUri="https://zenodo.org/record/235693/files/figure.png" pageId="10" pageNumber="245">Figs 2021, 27</figureCitation>
). As in male, but generally darker and with more contrasting pattern at prosoma dorsally. Femora ventrally without banded pattern, similarly dark as dorsally. Chelicerae dark reddish-brown with yellowish-brown proximo-lateral humps. Anterior lateral spinnerets ventro-laterally dark. For colouration pattern of a living specimen see
<figureCitation id="70978ED77F7D5522F5EBFB47FDDBFB55" box="[432,515,1192,1217]" captionStart="FIGURES 18 27" captionStartId="6.[151,264,1795,1818]" captionTargetBox="[173,1411,279,1748]" captionTargetId="figure@6.[169,1415,265,1763]" captionTargetPageId="6" captionText="FIGURES 18 27. Ctenus spp. from Mt Victoria, Myanmar, habitus (18, 20, 22, 24, 26 27 dorsal; 19, 21, 23, 25 ventral). 18 21, 26 27 Ctenus natmataung spec. nov., holotype male (18 19) and female paratype (20 21) 22 25 Ctenus sp. from Mt Victoria, Myanmar (22 23, male; 24 25 female)." httpUri="https://zenodo.org/record/235693/files/figure.png" pageId="10" pageNumber="245">Fig. 27</figureCitation>
. Note the continuous wide light band at the margin of the dark lateral band of the prosoma dorsally. This character might prove as diagnostic when larger series with this character being stable are known.
</paragraph>
<paragraph id="E81392527F7D5522F49CFAFAFC1FFA0D" blockId="10.[151,1436,904,1614]" pageId="10" pageNumber="245">
<emphasis id="DAD84E407F7D5522F49CFAFAFEE3FABA" bold="true" box="[199,315,1301,1326]" pageId="10" pageNumber="245">Variation.</emphasis>
Male PL 5.36.6, OL 3.86.3. Spination: palpal patella 171, tibia I d001. RTA tip present in 12 out of
<specimenCount id="FEAA59DB7F7D5522F4EEFAD6FEC2FAC5" box="[181,282,1336,1361]" pageId="10" pageNumber="245" type="male">18 males</specimenCount>
(with one tip present: left 4 [
<figureCitation id="70978ED77F7D5522F63AFAD7FD6FFAC5" box="[609,695,1336,1361]" captionStart="FIGURES 28 30" captionStartId="7.[151,264,1455,1478]" captionTargetBox="[231,1339,193,1433]" captionTargetId="figure@7.[231,1355,193,1435]" captionTargetPageId="7" captionText="FIGURES 28 30 Ctenus natmataung spec. nov., holotype male from Mt Victoria, Myanmar, male palp (28 prolateral; 29 ventral; 30 retrolateral)." httpUri="https://zenodo.org/record/235694/files/figure.png" pageId="10" pageNumber="245">Fig. 29</figureCitation>
], right 6; both tips present 2; both tips absent 6 [
<figureCitation id="70978ED77F7D5522F08DFAD7FAF5FAC5" box="[1238,1325,1336,1361]" captionStart="FIGURES 31 34" captionStartId="8.[151,264,1835,1858]" captionTargetBox="[150,1436,700,1807]" captionTargetId="figure@8.[150,1436,697,1814]" captionTargetPageId="8" captionText="FIGURES 31 34 Ctenus natmataung spec. nov., male paratype from Mt Victoria, Myanmar, male palp (31 prolateral; 32 ventral; 33 retrolateral; 34 embolus tip, ventral)." httpUri="https://zenodo.org/record/235695/files/figure.png" pageId="10" pageNumber="245">Fig. 32</figureCitation>
]); Female PL 5.18.1, OL 5.39.1. Epigynal field with one slit sensillum on each side and shape of anterior field and median plate with its median part as well as epigynal teeth varying (
<figureCitation id="70978ED77F7D5522F763FA6FFC63FA0D" box="[824,955,1408,1433]" captionStart="FIGURES 35 41" captionStartId="9.[151,264,1910,1933]" captionTargetBox="[151,1436,219,1889]" captionTargetId="figure@9.[151,1436,219,1889]" captionTargetPageId="9" captionText="FIGURES 35 41 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, copulatory organ (35, 38, 39, 40 epigyne, ventral; 36 epigyne, posterior; 37 vulva, dorsal; 41 epigyne, lateral). I + II—two chambers of spermathecae, chamber II functionally between chamber I and fertilization duct; EF—epigynal field; EP—separate patches of epigynal field." httpUri="https://zenodo.org/record/235696/files/figure.png" pageId="10" pageNumber="245">Figs 3537</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="A0B6C1D97F7D5522F49CFA4AFCE8F9DA" pageId="10" pageNumber="245" type="distribution">
<paragraph id="E81392527F7D5522F49CFA4AFF24FA75" blockId="10.[151,1436,904,1614]" pageId="10" pageNumber="245">
<emphasis id="DAD84E407F7D5522F49CFA4AFEB8FA2A" bold="true" box="[199,352,1445,1470]" pageId="10" pageNumber="245">Distribution.</emphasis>
Known only from the
<typeStatus id="37172CF07F7D5522F63BFA49FD48FA2A" box="[608,656,1446,1470]" pageId="10" pageNumber="245">type</typeStatus>
locality, Mt Victoria from elevations of
<quantity id="2F543FB77F7D5522F00EFA4AFB28FA29" box="[1109,1264,1445,1470]" metricMagnitude="3" metricUnit="m" metricValue="1.6684999999999999" metricValueMax="1.752" metricValueMin="1.585" pageId="10" pageNumber="245" unit="m" value="1668.5" valueMax="1752.0" valueMin="1585.0">15851752 m</quantity>
(
<figureCitation id="70978ED77F7D5522F0A4FA4AFA8BFA2A" box="[1279,1363,1445,1470]" captionStart="FIGURE 63" captionStartId="14.[151,249,1795,1817]" captionTargetBox="[150,1435,872,1772]" captionTargetId="figure@14.[150,1436,872,1773]" captionTargetPageId="14" captionText="FIGURE 63. Distributional records of Ctenidae treated in this paper. Large Map: Records at Nat Ma Taung (= Mt Victoria). Red triangles— Ctenus cladarus, yellow circle— C. pingu spec. nov., green squares— C. natmataung spec. nov., white arrowsummit. White numbers indicate elevation [m]. Inset: Location of Mt Victoria (orange drop), and Mt Popa (blue cross; Anahita popa spec. nov., 1250 m)." httpUri="https://zenodo.org/record/235702/files/figure.png" pageId="10" pageNumber="245">Fig. 63</figureCitation>
: green squares).
</paragraph>
<paragraph id="E81392527F7D5522F49CFA02FCE8F9DA" blockId="10.[151,1436,904,1614]" pageId="10" pageNumber="245">
<emphasis id="DAD84E407F7D5522F49CFA02FEC9F992" bold="true" box="[199,273,1517,1542]" pageId="10" pageNumber="245">Notes.</emphasis>
Two females were observed with the distal part of the males RTA clinging to their epigyne. The long and pointed appendix of the RTA penetrated the females cuticle in the right epigynal tooth (
<figureCitation id="70978ED77F7D5522F0C4F9FFFAC7F9BD" box="[1183,1311,1552,1577]" captionStart-0="FIGURES 35 41" captionStart-1="FIGURES 42 43" captionStartId-0="9.[151,264,1910,1933]" captionStartId-1="10.[151,264,807,830]" captionTargetBox-0="[151,1436,219,1889]" captionTargetBox-1="[153,1430,219,769]" captionTargetId-0="figure@9.[151,1436,219,1889]" captionTargetId-1="figure@10.[149,1437,214,776]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="FIGURES 35 41 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, copulatory organ (35, 38, 39, 40 epigyne, ventral; 36 epigyne, posterior; 37 vulva, dorsal; 41 epigyne, lateral). I + II—two chambers of spermathecae, chamber II functionally between chamber I and fertilization duct; EF—epigynal field; EP—separate patches of epigynal field." captionText-1="FIGURES 42 43 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, epigyne with male RTA tip, ventral." httpUri-0="https://zenodo.org/record/235696/files/figure.png" httpUri-1="https://zenodo.org/record/235697/files/figure.png" pageId="10" pageNumber="245">Figs 4042</figureCitation>
) and in the cuticle right beside the median plate (
<figureCitation id="70978ED77F7D5522F660F9DAFD57F9D9" box="[571,655,1589,1614]" captionStart="FIGURES 42 43" captionStartId="10.[151,264,807,830]" captionTargetBox="[153,1430,219,769]" captionTargetId="figure@10.[149,1437,214,776]" captionTargetPageId="10" captionText="FIGURES 42 43 Ctenus natmataung spec. nov., female paratypes from Mt Victoria, Myanmar, epigyne with male RTA tip, ventral." httpUri="https://zenodo.org/record/235697/files/figure.png" pageId="10" pageNumber="245">Fig. 43</figureCitation>
), respectively.
</paragraph>
</subSubSection>
</treatment>
</document>