201 lines
21 KiB
XML
201 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.295.4905" ID-GBIF-Dataset="59cbf2a9-41ca-4f5a-a350-ef034991798b" ID-PMC="PMC3677376" ID-Pensoft-Pub="1313-2970-295-1" ID-PubMed="23794874" ModsDocAuthor="" ModsDocDate="2013" ModsDocID="1313-2970-295-1" ModsDocOrigin="ZooKeys 295" ModsDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" checkinTime="1451247354359" checkinUser="pensoft" docAuthor="Branstetter, Michael G." docDate="2013" docId="13555CF39F778660495AD6BD5AE6AED0" docLanguage="en" docName="ZooKeys 295: 1-277" docOrigin="ZooKeys 295" docSource="http://dx.doi.org/10.3897/zookeys.295.4905" docTitle="Stenamma catracho Branstetter, 2013, sp. n." docType="treatment" docVersion="4" lastPageNumber="53" masterDocId="FFB3D3328201C177FF99FFC5FFA8A72C" masterDocTitle="Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)" masterLastPageNumber="277" masterPageNumber="1" pageNumber="50" updateTime="1668155749461" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision of the Middle American clade of the ant genus Stenamma Westwood (Hymenoptera, Formicidae, Myrmicinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Branstetter, Michael G.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2013</mods:date>
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<mods:detail type="volume">
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<mods:number>295</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>277</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.295.4905</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.295.4905</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-295-1</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="152043649" LSID="urn:lsid:zoobank.org:act:6D8C5D59-F8F2-4C8A-8D0B-1A543DF13706" httpUri="http://treatment.plazi.org/id/13555CF39F778660495AD6BD5AE6AED0" lastPageId="52" lastPageNumber="53" pageId="49" pageNumber="50">
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<subSubSection pageId="49" pageNumber="50" type="nomenclature">
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<paragraph pageId="49" pageNumber="50">
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<taxonomicName LSID="urn:lsid:zoobank.org:act:6D8C5D59-F8F2-4C8A-8D0B-1A543DF13706" class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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<taxonomicNameLabel pageId="49" pageNumber="50">sp. n.</taxonomicNameLabel>
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Worker: Figures 61, 62; Queen: Figure 63; Map: Figure 60
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</paragraph>
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</subSubSection>
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<subSubSection pageId="49" pageNumber="50" type="type material">
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<paragraph pageId="49" pageNumber="50">Type material.</paragraph>
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<paragraph pageId="49" pageNumber="50">
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Holotype worker. HONDURAS: Olancho, Parque Nacional La Muralla,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.09965">15.09965°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.74072">86.74072°W</geoCoordinate>
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<normalizedToken originalValue="±">+/-</normalizedToken>
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20m, 1530m, 2 May 2010, cloud forest, ex sifted leaf litter (LLAMA, collection Wa-C-01-1-36) [USNM, specimen CASENT0621306]. Paratypes: same data as holotype [1dq, 1w, USNM, CASENT0621305, CASENT0621307], [1w, CAS, CASENT0621308]; same data as holotype but
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.09925">15.09925°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.74063">86.74063°W</geoCoordinate>
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<normalizedToken originalValue="±">+/-</normalizedToken>
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20m, 1530m, 2 May 2010 (LLAMA, Wa-C-01-1-27) [1w, EAPZ, CASENT0621298], [1w, ECOSCE, CASENT0621299], [1w, FMNH, CASENT0621297], [1w, ICN, CASENT0621300];
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.09859">15.09859°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.74216">86.74216°W</geoCoordinate>
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<normalizedToken originalValue="±">+/-</normalizedToken>
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10m, 1510m, 2 May 2010 (LLAMA, Wm-C-01-1-03) [1w, INBio, CASENT0622022], [1w, JTLC, CASENT0622023], [1w, LACM, CASENT0622024], [1w, MGBPC, CASENT0623530], [1w, MCZ, CASENT0623531], [1w, MZSP, CASENT0623532], [1w, UCD, CASENT0623533];
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.09694">15.09694°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.74533">86.74533°W</geoCoordinate>
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<normalizedToken originalValue="±">+/-</normalizedToken>
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10m, 1460m, 5 May 2010 (LLAMA, Wm-C-01-1-09) [1w, UNAM, CASENT0622048], [1w, UVGC, CASENT0623534].
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50" type="worker diagnosis">
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<paragraph pageId="49" pageNumber="50">Worker diagnosis.</paragraph>
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<paragraph lastPageId="50" lastPageNumber="51" pageId="49" pageNumber="50">
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Integument mostly dark red-brown to orange-brown; small-sized species (see HL, ML, PrW below); anterior clypeal margin viewed from anterodorsal angle undulating (straight in full-face view), appearing as 2-4 blunt teeth; basal margin of mandible sinuous, with a shallow, but distinct basal depression; head and mesosoma densely sculptured, mostly rugoreticulae; gastral pilosity short, relatively dense, and usually appearing uniformly suberect to subdecumbent, but sometimes
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<pageBreakToken pageId="50" pageNumber="51" start="start">more</pageBreakToken>
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clearly bilayered; petiole somewhat long and gracile, with node relatively small; petiolar and postpetiolar nodes smooth only on anterior faces; eye relatively small (EL 0.08-0.09, REL 14-16), subcircular to oval-shaped, with 4-5 ommatidia in greatest diameter; frontal lobes moderately expanded (FLD 0.15-0.18, FLI 28-33), mostly to completely covering torular lobes in full-face view; propodeal spines short to medium length (PSL 0.11-0.18, PSI 1.6-2.1). Similar species:
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma crypticum" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="crypticum">Stenamma crypticum</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma cusuco" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="cusuco">Stenamma cusuco</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma hojarasca" order="Hymenoptera" pageId="50" pageNumber="51" phylum="Arthropoda" rank="species" species="hojarasca">Stenamma hojarasca</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="50" pageNumber="51" type="geographic range">
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<paragraph pageId="50" pageNumber="51">Geographic range.</paragraph>
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<paragraph pageId="50" pageNumber="51">Guatemala to Honduras.</paragraph>
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</subSubSection>
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<subSubSection lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51" type="worker description">
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<paragraph pageId="50" pageNumber="51">Worker description.</paragraph>
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<paragraph pageId="50" pageNumber="51">(9 measured) HL 0.61-0.68 (0.68), HW 0.51-0.59 (0.58), FLD 0.15-0.18 (0.18), PCW 0.01-0.03 (0.03), SL 0.52-0.55 (0.55), EL 0.08-0.09 (0.08), ACL 0.51-0.54 (0.53), ML 0.75-0.82 (0.82), PrW 0.36-0.41 (0.41), PSL 0.11-0.12 (0.12), SDL 0.05-0.07 (0.07), PL 0.28-0.32 (0.32), PH 0.15-0.17 (0.17), PW 0.12-0.13 (0.13) PPL 0.13-0.15 (0.15), PPH 0.13-0.15 (0.15), PPW 0.15-0.16 (0.16), MFL 0.57-0.63 (0.62), MTL 0.47-0.51 (0.51), CI 82-88 (86), SI 92-104 (94), REL 14-16 (14), FLI 28-33 (32), PSI 1.6-2.1 (1.8); MFI 84-95 (93), ACI1 68-71 (68), ACI2 96-102 (96).</paragraph>
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<paragraph lastPageId="52" lastPageNumber="53" pageId="50" pageNumber="51">
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Small species; general body color dark red-brown to orange-brown, with patches of dark brown to brown on gaster; appendages lighter, brown to yellow-brown; setae golden brown; mandible with 6 teeth, consisting of 3 distinct apical teeth, a basal tooth, and 2 middle teeth, which are often worn and indistinct; basal margin of mandible sinuous, with a shallow, but distinct basal depression, accompanied by attenuation of cuticle where mandible fits underneath clypeus; mandible mostly smooth and shining, with scattered piligerous punctae and basal striae; anterior clypeal margin viewed at anterodorsal angle undulating (straight in full-face view), appearing as 2-4 blunt teeth; median lobe of clypeus somewhat flattened, longitudinal carinulae absent or very faint, apex with a short transverse carinula; remainder of clypeus mostly smooth and shiny; posterior extension of clypeus between antennal insertions narrow (PCW 0.01-0.03), sides diverging posteriad; frontal lobes moderately expanded (FLD 0.15-0.18, FLI 28-33), with lateral apices shifted slightly posteriad of torular lobes, which are mostly to completely covered in full-face view; head subrectangular to oval-shaped (CI 82-88), posterior margin slightly depressed medially; eye relatively small (EL 0.08-0.09, REL 14-16), subcircular to oval-shaped, with 4-5 ommatidia in greatest diameter; head mostly rugoreticulate, with a few longitudinal rugae along midline, interstices faintly punctate; scape relatively long (SI 94-104), but variable, either reaching or not quite reaching posterior margin when laid back; scape surface shiny, but somewhat rough, with punctae and faint striae; flagellum with distinct 4-segmented antennal club; mesosoma densely sculptured, except for propodeal declivity, which only has a few faint transverse carinulae; promesonotal dorsum rugose to rugoreticulae, interstices faintly punctate; mesosomal side mostly punctate, with a few rugulae; propodeal dorsum with a few transverse carinae; promesonotum in profile, low-domed, roughly symmetrical; metantoal groove distinct, but somewhat shallow; anterodorsal margin of propodeum in profile with a small welt; propodeal spines present, short to medium length
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<pageBreakToken pageId="51" pageNumber="52" start="start">(</pageBreakToken>
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PSL 0.11-0.18, PSI 1.6-2.1); petiole somewhat long and gracile (PL/HW 0.53-0.59); petiolar node in profile relatively small (PH/PL 0.49-0.59), asymmetrical, with anterior face long and sloping and posterior face short and nearly vertical, dorsum of node rounded, apex pointing posteriad; postpetiole relatively small (PPH/PH 0.85-0.91), somewhat dorsoventrally compressed; anterior faces of petiolar and postpetiolar nodes smooth and shiny, remainder of waist mostly punctate, with a few small rugulae around postpetiolar node; gaster mostly smooth and shiny, with
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<pageBreakToken pageId="52" pageNumber="53" start="start">scattered</pageBreakToken>
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piligerous punctae; most of body dorsum with short to moderately long standing pilosity; scape with suberect to decumbent setae; gastral pilosity variable, usually short, relatively dense, and uniformly suberect to subdecumbent, but sometimes clearly bilayered, with a longer suberect layer, and a shorter decumbent layer; setae on legs mostly decumbent to appressed, with some suberect setae on coxae and venter of profemur.
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</paragraph>
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<caption pageId="52" pageNumber="53">
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<paragraph pageId="52" pageNumber="53">
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Figure 61.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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holotype worker (CASENT0621306) A Profile B Face C Dorsum D Anterior clypeal margin in anterodorsal view E Gaster.
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</paragraph>
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</caption>
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<caption pageId="52" pageNumber="53">
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<paragraph pageId="52" pageNumber="53">
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Figure 62.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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worker variant (CASENT0621563) A Face B Profile C Dorsum.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="52" pageNumber="53" type="queen description">
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<paragraph pageId="52" pageNumber="53">Queen description.</paragraph>
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<paragraph pageId="52" pageNumber="53">(3 measured) HL 0.67-0.69 (0.69), HW 0.59-0.62 (0.61), FLD 0.16-0.21 (0.21), PCW 0.02-0.04 (0.02), SL 0.56-0.58 (0.56), EL 0.16-0.18 (0.17), ACL 0.54-0.57 (054), ML 0.92-1.00 (0.95), PrW 0.50-0.55 (0.53), PSL 0.15-0.16 (0.15), SDL 0.08-0.09 (0.09), PL 0.36-0.38 (0.38), PH 0.18-0.20 (0.19), PW 0.15-0.16 (0.15), PPL 0.15-0.17 (0.15), PPH 0.18-0.19 (0.18), PPW 0.19-0.20 (0.19), MFL 0.63-0.69 (0.63), MTL 0.53-0.58 (0.53), CI 88-90 (88), SI 92-96 (92), REL 27-30 (27), FLI 25-34 (34), PSI 1.6-2.1 (1.6), MFI 88-97 (97), ACI1 67-69 (69), ACI2 96-98 (69).</paragraph>
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<paragraph pageId="52" pageNumber="53">Same as worker except for standard queen modifications and as follows: mesoscutum mostly longitudinally rugose, but with a central longitudinal carina that is thick at anterior margin; most of katepisternum and part of anepisternum smooth and shiny; gastral pilosity denser, and more distinctly bilayered.</paragraph>
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<caption pageId="52" pageNumber="53">
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<paragraph pageId="52" pageNumber="53">
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Figure 63.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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paratype queen (CASENT0621305) A Profile B Face C Dorsum.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="52" pageNumber="53" type="male">
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<paragraph pageId="52" pageNumber="53">Male.</paragraph>
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<paragraph pageId="52" pageNumber="53">Unknown.</paragraph>
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</subSubSection>
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<subSubSection pageId="52" pageNumber="53" type="biology">
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<paragraph pageId="52" pageNumber="53">Biology.</paragraph>
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<paragraph pageId="52" pageNumber="53">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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is known only from sifted leaf litter collected from the forest floor. It occurs from approximately 1100-1900 m elevation in montane wet forest habitats.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="52" pageNumber="53" type="comments">
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<paragraph pageId="52" pageNumber="53">Comments.</paragraph>
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<paragraph pageId="52" pageNumber="53">
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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is most likely to be confused with
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma cusuco" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="cusuco">Stenamma cusuco</taxonomicName>
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or
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma hojarasca" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="hojarasca">Stenamma hojarasca</taxonomicName>
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, both of which have expanded frontal lobes and somewhat elongate petioles.
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma cusuco" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="cusuco">Stenamma cusuco</taxonomicName>
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can be distinguished easily by its distinctive tridentate clypeal margin, and
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma hojarasca" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="hojarasca">Stenamma hojarasca</taxonomicName>
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can be separated by its longer petiole and completely shiny petiolar and postpetiolar nodes.
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</paragraph>
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<paragraph pageId="52" pageNumber="53">
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There are some morphological differences unique to the Meambar population (Figure 62) of
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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. Meambar specimens have most pilosity longer, with the gastral pilosity clearly bilayered; they have lighter integument color; and they have narrower frontal lobes. The other populations have the pilosity shorter overall and the gastral pilosity denser and not as clearly bilayered. I treat these differences as intraspecific variation until more material is collected.
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</paragraph>
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<paragraph pageId="52" pageNumber="53">
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Molecular phylogenetic data show that
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma catracho" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="catracho">Stenamma catracho</taxonomicName>
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most likely belongs to a clade that includes
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma crypticum" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="crypticum">Stenamma crypticum</taxonomicName>
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,
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma monstrosum" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="monstrosum">Stenamma monstrosum</taxonomicName>
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, and
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma saenzae" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="saenzae">Stenamma saenzae</taxonomicName>
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, with
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<taxonomicName class="Insecta" family="Formicidae" genus="Stenamma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Stenamma monstrosum" order="Hymenoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="monstrosum">Stenamma monstrosum</taxonomicName>
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being its sister species (Branstetter unpublished data).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="52" pageNumber="53" type="material examined">
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<paragraph pageId="52" pageNumber="53">Material examined.</paragraph>
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<paragraph pageId="52" pageNumber="53">
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GUATEMALA:Zacapa: 2km SE La
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<normalizedToken originalValue="Unión">Union</normalizedToken>
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,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.95344">14.95344°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-89.27587">89.27587°W</geoCoordinate>
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, 1430m, 12 May 2009 (LLAMA); HONDURAS: Comayagua: PN Cerro Azul Meambar,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="14.8696">14.86960°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-87.89843">87.89843°W</geoCoordinate>
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, 1140m, 20 May 2010 (LLAMA); Olancho: PN La Muralla,
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<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.09511">15.09511°N</geoCoordinate>
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,
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<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.73925">86.73925°W</geoCoordinate>
|
|
, 1420m, 2 May 2010 (LLAMA); PN La Muralla,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.09965">15.09965°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.74072">86.74072°W</geoCoordinate>
|
|
, 1530m, 2 May 2010 (LLAMA); PN La Muralla,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.10257">15.10257°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.73642">86.73642°W</geoCoordinate>
|
|
, 1650m, 2 May 2010 (R. S. Anderson); PN La Muralla,
|
|
<geoCoordinate direction="north" orientation="latitude" precision="1" value="15.09807">15.09807°N</geoCoordinate>
|
|
,
|
|
<geoCoordinate direction="west" orientation="longitude" precision="1" value="-86.72047">86.72047°W</geoCoordinate>
|
|
, 1860m, 3 May 2010 (LLAMA).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |