treatments-xml/data/05/8F/87/058F87894B18573E8931563DEA3F645E.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1030.62544" ID-GBIF-Dataset="91fec855-7fa9-4ad6-84d7-9b9d35e23e8b" ID-PMC="PMC8060247" ID-Pensoft-Pub="1313-2970-1030-1" ID-Pensoft-UUID="875199397EEE5F7898EA1DB25DA62D25" ID-PubMed="33958904" ModsDocID="1313-2970-1030-1" checkinTime="1618371561543" checkinUser="pensoft" docAuthor="Means, Jackson C., Hennen, Derek A. &amp; Marek, Paul E." docDate="2021" docId="058F87894B18573E8931563DEA3F645E" docLanguage="en" docName="ZooKeys 1030: 1-180" docOrigin="ZooKeys 1030" docPubDate="2021-04-13" docSource="http://dx.doi.org/10.3897/zookeys.1030.62544" docTitle="Nannaria blackmountainensis Means &amp; Hennen &amp; Marek 2021, sp. nov." docType="treatment" docVersion="5" id="875199397EEE5F7898EA1DB25DA62D25" lastPageNumber="1" masterDocId="875199397EEE5F7898EA1DB25DA62D25" masterDocTitle="A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)" masterLastPageNumber="180" masterPageNumber="1" pageNumber="1" updateTime="1673961214978" updateUser="julia">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A revision of the minor species group in the millipede genus Nannaria Chamberlin, 1918 (Diplopoda, Polydesmida, Xystodesmidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Means, Jackson C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7377-0696</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
<mods:nameIdentifier type="email">mjacks4@vt.edu</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hennen, Derek A.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-7005-1151</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Marek, Paul E.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7048-2514</mods:nameIdentifier>
<mods:affiliation>Virginia Tech, Department of Entomology, Blacksburg, Virginia 24061, USA</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-04-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1030</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>180</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.1030.62544</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1030.62544</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1030-1</mods:identifier>
<mods:identifier type="Pensoft-UUID">875199397EEE5F7898EA1DB25DA62D25</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="181004765" LSID="urn:lsid:zoobank.org:act:2ACF8697-BA97-4E11-BFBD-233FE0452FAB" httpUri="http://treatment.plazi.org/id/058F87894B18573E8931563DEA3F645E" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="8RVTX" LSID="http://zoobank.org/2ACF8697-BA97-4E11-BFBD-233FE0452FAB" authority="Means &amp; Hennen &amp; Marek, 2021" authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Nannaria blackmountainensis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="blackmountainensis" status="sp. nov.">Nannaria blackmountainensis</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Nannaria blackmountainensis sp. nov. holotype ♂ left gonopod (VTEC, SPC 001090) A anterior view; red arrow indicates hooked lateral flange B medial view; red triangle indicates reduced prefemoral spine C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure73" httpUri="https://binary.pensoft.net/fig/531176" pageId="0" pageNumber="1">Figs 73</figureCitation>
<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Nannaria blackmountainensis sp. nov. paratype ♂ (VMNH, SPC 001002) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure74" httpUri="https://binary.pensoft.net/fig/531177" pageId="0" pageNumber="1">, 74 Vernacular name: &quot;The Black Mountain Twisted-Claw Millipede&quot;</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355611" collectingDate="2011-05-10" collectorName="P. Marek, C. Hall, D. &amp; M. Beamer" country="United States" county="Harlan Co." elevation="1250" latitude="36.9156" location="Black Mountain Ridge Rd." longLatPrecision="6" longitude="-82.893" municipality="Black Mountain" specimenCount="♂" stateProvince="Kentucky" typeStatus="Holotype">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Holotype</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry name="United States of America">United States</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Kentucky">Kentucky</collectingRegion>
</emphasis>
• 
<specimenCount>♂</specimenCount>
; 
<collectingCounty>Harlan Co.</collectingCounty>
, 
<collectingMunicipality>Black Mountain</collectingMunicipality>
summit, radar station access road off 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:AB54246FB81832D164970C0DF77A2C2C" country="United States" county="Harlan Co." latitude="36.9156" longLatPrecision="6" longitude="-82.893" municipality="Black Mountain" name="Black Mountain Ridge Rd." stateProvince="Kentucky">Black Mountain Ridge Rd.</location>
, about 2.4 rd. km E jct w/ KY-160; 
<geoCoordinate degrees="36.9156" direction="north" orientation="latitude" precision="5" value="36.9156">36.9156°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.8930" direction="west" orientation="longitude" precision="5" value="-82.893">82.8930°W</geoCoordinate>
; elev. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">
<elevation metricMagnitude="3" metricUnit="m" metricValue="1.25" unit="m" value="1250.0">1250 m</elevation>
</quantity>
; 
<collectingDate value="2011-05-10">10 May 2011</collectingDate>
; hand collected; 
<collectorName>P. Marek</collectorName>
, 
<collectorName>C. Hall</collectorName>
and 
<collectorName>D. &amp; M. Beamer</collectorName>
leg.; VTEC SPC001090.
</materialsCitation>
</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355749" collectingDate="2006-05-31" collectorName="P. Marek" country="United States" county="Harlan Co." elevation="570" latitude="36.7607" location="Stone Mountain State Natural Area" longLatPrecision="6" longitude="-83.14" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="Paratypes">
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">
<typeStatus>Paratypes</typeStatus>
</emphasis>
:
</emphasis>
<collectingCountry name="United States of America">United States</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Kentucky">Kentucky</collectingRegion>
</emphasis>
• 
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingCounty>Harlan Co.</collectingCounty>
, 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:81D2202A205AFA06164E3E37F7726E00" country="United States" county="Harlan Co." latitude="36.7607" longLatPrecision="6" longitude="-83.14" name="Stone Mountain State Natural Area" stateProvince="Kentucky">Stone Mountain State Natural Area</location>
, wooded slope above entrance ex US-421; 
<geoCoordinate degrees="36.7607" direction="north" orientation="latitude" precision="5" value="36.7607">36.7607°N</geoCoordinate>
, - 
<geoCoordinate degrees="83.1400" direction="west" orientation="longitude" precision="5" value="-83.14">83.1400°W</geoCoordinate>
; elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.7" unit="m" value="570.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.7" unit="m" value="570.0">570 m</elevation>
</quantity>
; 
<collectingDate value="2006-05-31">31 May 2006</collectingDate>
; hand collected; 
<collectorName>P. Marek</collectorName>
leg.; VMNH SPC001002
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="3071355510" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="P. Marek" country="United States" county="Harlan Co." elevation="570" latitude="36.7607" location="Stone Mountain State Natural Area" longLatPrecision="6" longitude="-83.14" specimenCount="1" specimenCount-female="1" stateProvince="Kentucky" typeStatus="paratype">
<specimenCount type="female">1 ♀</specimenCount>
same collection data as for preceding; VTEC SPC001009
</materialsCitation>
• 
<materialsCitation ID-GBIF-Occurrence="3071355390" collectingDate="2005-07-25" collectorName="N Slope, P. Marek, C. Spruill" country="United States" county="Harlan Co." elevation="691" latitude="36.7328" location="Cumberland Mountain" longLatPrecision="6" longitude="-83.2216" municipality="Kentucky" specimenCount="1" specimenCount-male="1" stateProvince="Kentucky" typeStatus="paratype">
<specimenCount type="male">1 ♂</specimenCount>
<collectingMunicipality>Kentucky</collectingMunicipality>
, 
<collectingCounty>Harlan Co.</collectingCounty>
, 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:78CDD30286AAA100E2DAB042AEE1446F" country="United States" county="Harlan Co." latitude="36.7328" longLatPrecision="6" longitude="-83.2216" municipality="Kentucky" name="Cumberland Mountain" stateProvince="Kentucky">Cumberland Mountain</location>
(
<collectorName>N Slope</collectorName>
), 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:004871720D292183FD5FBD451F54BC10" country="United States" county="Harlan Co." latitude="36.7328" longLatPrecision="6" longitude="-83.2216" municipality="Kentucky" name="Wagonroad Tunnel Trail" stateProvince="Kentucky">Wagonroad Tunnel Trail</location>
; 
<geoCoordinate degrees="36.7328" direction="north" orientation="latitude" precision="5" value="36.7328">36.7328°N</geoCoordinate>
, - 
<geoCoordinate degrees="83.2216" direction="west" orientation="longitude" precision="5" value="-83.2216">83.2216°W</geoCoordinate>
; elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" unit="m" value="691.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="6.909999999999999" unit="m" value="691.0">691 m</elevation>
</quantity>
; 
<collectingDate value="2005-07-25">25 July 2005</collectingDate>
; hand collected; 
<collectorName>P. Marek</collectorName>
and 
<collectorName>C. Spruill</collectorName>
leg.; FMNH SPC000652
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<materialsCitation ID-GBIF-Occurrence="3071355326" collectingDate="1941-08-09" collectingDateMax="2005-07-26" collectingDateMin="1941-08-09" collectorName="R. Hoffman, J. Ettman, Coll, P. Marek, C. Spruill, Plumetree, L. Hubricht, Banner Elk, NC, Grandfather Mtn., R. Shelley, Newland, D. Massee, Valle Crucis, NC, J. Clamp, Roan Mtn., Brooks, Carter County, N, Roane Mtn, Dr., Mrs. Brooks, Cumberland Mtns., E. Cope, Pennington Gap, VA" country="United States" county="Bell County" elevation="528" latitude="36.1079" location="Honeymoon Falls" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" specimenCount="58" specimenCount-female="2" specimenCount-male="56" stateProvince="Kentucky">
<collectingCountry name="United States of America">United States</collectingCountry>
- 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Kentucky">Kentucky</collectingRegion>
</emphasis>
• 
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingCounty>Bell County</collectingCounty>
, 
<collectingMunicipality>Pine Mountain State Park</collectingMunicipality>
, trail to 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:9D08ED08298CF32F8BC0E5C2B0C83634" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Honeymoon Falls" stateProvince="Kentucky">Honeymoon Falls</location>
; 
<geoCoordinate degrees="36.7434" direction="north" orientation="latitude" precision="5" value="36.7434">36.7434°N</geoCoordinate>
, - 
<geoCoordinate degrees="83.7121" direction="west" orientation="longitude" precision="5" value="-83.7121">83.7121°W</geoCoordinate>
; 
<collectingDate value="1976-09-26">26 Sept.1976</collectingDate>
; 
<collectorName>R. Hoffman</collectorName>
leg.; VMNH NAN0184 • 
<specimenCount type="male">1 ♂</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:B156333021A8C353383C5B9776C17D51" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Bell County" stateProvince="Kentucky">Bell County</location>
, 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:6DF68D55A8C45F44945968F1B327916C" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Pine Mtn. State Park" stateProvince="Kentucky">Pine Mtn. State Park</location>
; 
<geoCoordinate degrees="36.6844" direction="north" orientation="latitude" precision="5" value="36.6844">36.6844°N</geoCoordinate>
, - 
<geoCoordinate degrees="83.8351" direction="west" orientation="longitude" precision="5" value="-83.8351">83.8351°W</geoCoordinate>
; 
<collectingDate value="1975-05-10">10 May 1975</collectingDate>
; 
<collectorName>J. Ettman</collectorName>
leg.; VMNH NAN0493 • 
<specimenCount type="male">2 ♂♂</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:AB3823788009D064BAEDACF379DC56D2" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Bell County" stateProvince="Kentucky">Bell County</location>
, 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:5F8C4C7889D76258FB8760EE87ADC4A4" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Pine Mtn. State Park" stateProvince="Kentucky">Pine Mtn. State Park</location>
, 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:96396F08CF9E560B205ED0602F185E2E" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Wildflower Garden Area" stateProvince="Kentucky">Wildflower Garden Area</location>
; 
<geoCoordinate degrees="36.7359" direction="north" orientation="latitude" precision="5" value="36.7359">36.7359°N</geoCoordinate>
, - 
<geoCoordinate degrees="83.7379" direction="west" orientation="longitude" precision="5" value="-83.7379">83.7379°W</geoCoordinate>
; 
<collectingDate value="1976-02-01">1 Feb. 1976</collectingDate>
; 
<collectorName>J. Ettman</collectorName>
leg.; NCSM NAN0545 • 
<specimenCount type="female">1 ♀</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:B54F15DAAE10F9AFCA316FFF80B1EAC1" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Harlan County" stateProvince="Kentucky">Harlan County</location>
, 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:964599E0B538FF417D516150C7A4A6DD" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Cumberland Mtn" stateProvince="Kentucky">Cumberland Mtn</location>
(N slope), 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:1ECE430D9E3FCA36894C415A4AF0BCD0" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Wagonroad Tunnel Trail" stateProvince="Kentucky">Wagonroad Tunnel Trail</location>
(
<geoCoordinate degrees="36.7389" direction="north" orientation="latitude" precision="5" value="36.7389">36.7389°N</geoCoordinate>
, - 
<geoCoordinate degrees="83.2197" direction="west" orientation="longitude" precision="5" value="-83.2197">83.2197°W</geoCoordinate>
, elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" unit="m" value="528.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="5.279999999999999" unit="m" value="528.0">528 m</elevation>
</quantity>
), 
<collectingDate value="2005-07-26">26 July 2005</collectingDate>
(
<collectorName>Coll</collectorName>
: 
<collectorName>P. Marek</collectorName>
, 
<collectorName>C. Spruill</collectorName>
; VTEC SPC000794. SCAU - 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="North Carolina">North Carolina</collectingRegion>
</emphasis>
• 
<specimenCount type="male">1 ♂</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:29D3D50544F3A451FB0C4A69545C3E1A" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Avery County" stateProvince="Kentucky">Avery County</location>
, wooded hillside at 
<collectorName>Plumetree</collectorName>
; 
<geoCoordinate degrees="36.0269" direction="north" orientation="latitude" precision="5" value="36.0269">36.0269°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.0080" direction="west" orientation="longitude" precision="5" value="-82.008">82.0080°W</geoCoordinate>
; 
<collectingDate value="1964-06-04">4 June 1964</collectingDate>
; 
<collectorName>L. Hubricht</collectorName>
leg.; VMNH NAN0183 • 
<specimenCount type="male">11 ♂♂</specimenCount>
<specimenCount>♀♀</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:AAA02A8BD75F58D9474E1201C6451A23" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Avery County" stateProvince="Kentucky">Avery County</location>
, 3.7 SSE 
<collectorName>Banner Elk, NC</collectorName>
105 &amp; 184, 
<collectorName>Grandfather Mtn.</collectorName>
; 
<geoCoordinate degrees="36.1170" direction="north" orientation="latitude" precision="5" value="36.117">36.1170°N</geoCoordinate>
, - 
<geoCoordinate degrees="81.8386" direction="west" orientation="longitude" precision="5" value="-81.8386">81.8386°W</geoCoordinate>
; 
<collectingDate value="1973-09-08">8 Sept. 1973</collectingDate>
; 
<collectorName>R. Shelley</collectorName>
; NCSM NAN0498 • 
<specimenCount type="male">6 ♂♂</specimenCount>
; same collection data as for preceding; 1978; 
<collectorName>R. Shelley</collectorName>
leg.; NCSM NAN0518 • 
<specimenCount type="male">20 ♂♂</specimenCount>
<specimenCount>♀♀</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:06B1C7B269FB9C9F51C2856709C1AA28" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Avery County" stateProvince="Kentucky">Avery County</location>
, 
<collectorName>Newland</collectorName>
; 
<geoCoordinate degrees="36.0873" direction="north" orientation="latitude" precision="5" value="36.0873">36.0873°N</geoCoordinate>
, - 
<geoCoordinate degrees="81.9273" direction="west" orientation="longitude" precision="5" value="-81.9273">81.9273°W</geoCoordinate>
; 
<collectingDate value="1984-08-20">20 Aug. 1984</collectingDate>
; 
<collectorName>D. Massee</collectorName>
leg.; NCSM NAN0522 • 
<specimenCount type="male">1 ♂</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:ACB72814D348E88FE34A386BD6B15D6A" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Watauga County" stateProvince="Kentucky">Watauga County</location>
, 
<collectorName>Valle Crucis, NC</collectorName>
194, ca. 6S 1113; 
<geoCoordinate degrees="36.2092" direction="north" orientation="latitude" precision="5" value="36.2092">36.2092°N</geoCoordinate>
, - 
<geoCoordinate degrees="81.7783" direction="west" orientation="longitude" precision="5" value="-81.7783">81.7783°W</geoCoordinate>
; 
<collectingDate value="1975-10-11">11 Oct. 1975</collectingDate>
; 
<collectorName>J. Clamp</collectorName>
leg.; NCSM NAN0515; SCAU - 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Tennessee">Tennessee</collectingRegion>
</emphasis>
• 
<specimenCount type="male">1 ♂</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:F1AC03991DE103B9881AA34F47C7DBFB" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Carter County" stateProvince="Kentucky">Carter County</location>
, 
<locationDeviation location="Burbank">
<quantity metricMagnitude="3" metricUnit="m" metricValue="4.02336" unit="mi" value="2.5">2.5 miles</quantity>
S of Burbank
</locationDeviation>
; 
<geoCoordinate degrees="36.1123" direction="north" orientation="latitude" precision="5" value="36.1123">36.1123°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.1019" direction="west" orientation="longitude" precision="5" value="-82.1019">82.1019°W</geoCoordinate>
; 
<collectingDate value="1951-05-02">2 May 1951</collectingDate>
; 
<collectorName>L. Hubricht</collectorName>
leg.; VMNH, NAN0185 • 
<specimenCount type="male">6 ♂</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:3724D194045A69192CAE8E01D49F99BC" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Carter County" stateProvince="Kentucky">Carter County</location>
, 
<collectorName>Roan Mtn.</collectorName>
; 
<geoCoordinate degrees="36.1943" direction="north" orientation="latitude" precision="5" value="36.1943">36.1943°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.0710" direction="west" orientation="longitude" precision="5" value="-82.071">82.0710°W</geoCoordinate>
; 
<collectingDate value="1941-08-09">9 Aug. 1941</collectingDate>
; 
<collectorName>Brooks</collectorName>
leg.; VMNH, NAN0186 • 
<specimenCount type="male">5 ♂♂</specimenCount>
; 
<collectorName>
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:50247736C667FC86A6E49FF92C5A4E61" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Carter County" stateProvince="Kentucky">Carter County</location>
, N
</collectorName>
slope, 
<collectorName>Roane Mtn</collectorName>
; 
<geoCoordinate degrees="36.1079" direction="north" orientation="latitude" precision="5" value="36.1079">36.1079°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.1280" direction="west" orientation="longitude" precision="5" value="-82.128">82.1280°W</geoCoordinate>
; 
<collectingDate value="1941-08-09">9 Aug. 1941</collectingDate>
; 
<collectorName>Dr.</collectorName>
&amp; 
<collectorName>Mrs. Brooks</collectorName>
leg.; VMNH, NAN0294 • 
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectorName>Cumberland Mtns.</collectorName>
; 
<collectorName>E. Cope</collectorName>
leg.; VMNH, NAN0181; SCAU - 
<emphasis bold="true" pageId="0" pageNumber="1">
<collectingRegion country="United States of America" name="Virginia">Virginia</collectingRegion>
</emphasis>
• 
<specimenCount type="female">1 ♀</specimenCount>
; 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:04374BA83B342179A6A4DD7583731A96" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Lee County" stateProvince="Kentucky">Lee County</location>
, 
<location LSID="urn:lsid:plazi:treatment:058F87894B18573E8931563DEA3F645E:7281714050DD3DBC4FE9C3EEB7AD4D98" country="United States" county="Bell County" latitude="36.1079" longLatPrecision="6" longitude="-82.128" municipality="Pine Mountain State Park" name="Poor Valley" stateProvince="Kentucky">Poor Valley</location>
, 
<collectorName>Pennington Gap, VA</collectorName>
. 621, ca. 
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.6" unit="km" value="1.6">1.6 km</quantity>
E
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3071355313" collectingDate="1994-09-13" collectingDateMax="2005-06-10" collectingDateMin="1994-09-13" collectorName="P. Marek, Powell Mtn., C. Hobson, D. Stevenson, For, Suppl." country="US" county="Scott County" elevation="460" latitude="36.7311" location="Scott County" longLatPrecision="6" longitude="-82.8071" specimenCount="1" specimenCount-male="1">
<collectingCountry name="United States of America">US</collectingCountry>
. 421, on N facing slope; 
<geoCoordinate degrees="36.7768" direction="north" orientation="latitude" precision="5" value="36.7768">36.7768°N</geoCoordinate>
, - 
<geoCoordinate degrees="83.0157" direction="west" orientation="longitude" precision="5" value="-83.0157">83.0157°W</geoCoordinate>
, elev. 
<quantity metricMagnitude="2" metricUnit="m" metricValue="4.6" unit="m" value="460.0">
<elevation metricMagnitude="2" metricUnit="m" metricValue="4.6" unit="m" value="460.0">460 m</elevation>
</quantity>
; 
<collectingDate value="2005-06-10">10 June 2005</collectingDate>
; hand collected; 
<collectorName>P. Marek</collectorName>
leg.; VTEC, SPC000540 • 
<specimenCount type="male">1 ♂</specimenCount>
; 
<collectingCounty>Scott County</collectingCounty>
, 
<collectorName>Powell Mtn.</collectorName>
off FS 642, ca. 
<locationDeviation location="Duffield">
<quantity metricMagnitude="3" metricUnit="m" metricValue="1.609344" unit="mi" value="1.0">1 mi</quantity>
NW of Duffield
</locationDeviation>
, JNF; 
<geoCoordinate degrees="36.7311" direction="north" orientation="latitude" precision="5" value="36.7311">36.7311°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.8071" direction="west" orientation="longitude" precision="5" value="-82.8071">82.8071°W</geoCoordinate>
; 
<collectingDate value="1994-09-13">13 Sept. 1994</collectingDate>
; 
<collectorName>C. Hobson</collectorName>
, 
<collectorName>D. Stevenson</collectorName>
leg.; VMNH NAN0182. 
<collectorName>For</collectorName>
detailed collection data see 
<collectorName>Suppl.</collectorName>
material 7.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
Adult males of 
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria blackmountainensis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="blackmountainensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria blackmountainensis</emphasis>
</taxonomicName>
sp. nov. are distinct from other 
<taxonomicName authorityName="Chamberlin" authorityYear="1918" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria</emphasis>
</taxonomicName>
and the nearby 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. domestica" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="domestica">
<emphasis italics="true" pageId="0" pageNumber="1">N. domestica</emphasis>
</taxonomicName>
based on the following combination of characters: 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Gonopods</emphasis>
.
</emphasis>
Gonopodal acropodite gently curving dorsomedially with pronounced medial swelling (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Nannaria blackmountainensis sp. nov. holotype ♂ left gonopod (VTEC, SPC 001090) A anterior view; red arrow indicates hooked lateral flange B medial view; red triangle indicates reduced prefemoral spine C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure73" httpUri="https://binary.pensoft.net/fig/531176" pageId="0" pageNumber="1">73A</figureCitation>
). Acropodite tip with large, hooked lateral flange (Fig. 
<figureCitation captionStart="Figure 73" captionStartId="F73" captionText="Figure 73. Nannaria blackmountainensis sp. nov. holotype ♂ left gonopod (VTEC, SPC 001090) A anterior view; red arrow indicates hooked lateral flange B medial view; red triangle indicates reduced prefemoral spine C posterior view. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure73" httpUri="https://binary.pensoft.net/fig/531176" pageId="0" pageNumber="1">73A</figureCitation>
, red arrow), not small, triangular lateral flange as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. domestica" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="domestica">
<emphasis italics="true" pageId="0" pageNumber="1">N. domestica</emphasis>
</taxonomicName>
. Tip terminating in small, dorsally directed rectangular point, not sharp, caudally directed as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. domestica" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="domestica">
<emphasis italics="true" pageId="0" pageNumber="1">N. domestica</emphasis>
</taxonomicName>
. Height of telopodite basal zone ca. 1/2 length of prefemoral process, not &lt;1/3 length as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. domestica" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="domestica">
<emphasis italics="true" pageId="0" pageNumber="1">N. domestica</emphasis>
</taxonomicName>
. Prefemoral process straight, acuminate, not laminate and serpentine as in 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. domestica" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="domestica">
<emphasis italics="true" pageId="0" pageNumber="1">N. domestica</emphasis>
</taxonomicName>
. Prefemoral spine reduced and fused to prefemoral process, forming small ridge (Fig. 
<figureCitation captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Nannaria fowleri Chamberlin, 1947, ♂ (VTEC, MPE 03017) left gonopod A anterior view B medial view C posterior view; red triangle indicates stout, tooth-like prefemoral spine. Scale bar: 0.5 mm." figureDoi="10.3897/zookeys.1030.62544.figure15" httpUri="https://binary.pensoft.net/fig/531118" pageId="0" pageNumber="1">15B</figureCitation>
, red triangle). 
<emphasis bold="true" pageId="0" pageNumber="1">
<emphasis italics="true" pageId="0" pageNumber="1">Color</emphasis>
.
</emphasis>
Tergites with hot orange/red paranotal spots (Fig. 
<figureCitation captionStart="Figure 74" captionStartId="F74" captionText="Figure 74. Nannaria blackmountainensis sp. nov. paratype ♂ (VMNH, SPC 001002) coloration. Scale bar: 4.0 mm." figureDoi="10.3897/zookeys.1030.62544.figure74" httpUri="https://binary.pensoft.net/fig/531177" pageId="0" pageNumber="1">74</figureCitation>
). Jet black background. Dorsum of collum smooth with orange margin.
</paragraph>
<caption doi="10.3897/zookeys.1030.62544.figure73" httpUri="https://binary.pensoft.net/fig/531176" pageId="0" pageNumber="1" start="Figure 73" startId="F73">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 73.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria blackmountainensis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="blackmountainensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria blackmountainensis</emphasis>
</taxonomicName>
sp. nov. holotype ♂ left gonopod (VTEC, SPC001090) 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
anterior view; red arrow indicates hooked lateral flange 
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
medial view; red triangle indicates reduced prefemoral spine 
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
posterior view. Scale bar: 0.5 mm.
</paragraph>
</caption>
<caption doi="10.3897/zookeys.1030.62544.figure74" httpUri="https://binary.pensoft.net/fig/531177" pageId="0" pageNumber="1" start="Figure 74" startId="F74">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 74.</emphasis>
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria blackmountainensis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="blackmountainensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria blackmountainensis</emphasis>
</taxonomicName>
sp. nov. paratype ♂ (VMNH, SPC001002) coloration. Scale bar: 4.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements.</paragraph>
<paragraph pageId="0" pageNumber="1">♂ holotype (VTEC, SPC001090): BL = 31.2, CW = 3.6, IW = 2.3, ISW = 0.7, B11W = 4.7, B11H = 3.2. ♀ paratype (VTEC, SPC001009): BL = 35.4, CW = 4.1, IW = 2.8, ISW = 1.0, B11W = 5.5, B11H = 3.9.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="variation">
<paragraph pageId="0" pageNumber="1">Variation.</paragraph>
<paragraph pageId="0" pageNumber="1">
There exists a fair amount of variation amongst individuals of 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
</taxonomicName>
sp. nov. from throughout the 
<normalizedToken originalValue="species’">species'</normalizedToken>
range. Individuals from the northern border of Tennessee and North Carolina have medially curving prefemoral processes and sharp, pronounced prefemoral spines, while the only known specimen from near Crummies, Kentucky (SPC000652) has a small, triangular prefemoral spine. The southern populations of 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
</taxonomicName>
may be discovered to be a separate species; however, such a discovery is contingent on the further collection of genetic material.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Means &amp; Hennen &amp; Marek" authorityYear="2021" class="Diplopoda" family="Xystodesmidae" genus="Nannaria" kingdom="Animalia" lsidName="Nannaria blackmountainensis" order="Polydesmida" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="blackmountainensis">
<emphasis italics="true" pageId="0" pageNumber="1">Nannaria blackmountainensis</emphasis>
</taxonomicName>
sp. nov. has a disjunct distribution in the confluence of eastern Kentucky and Tennessee and western Virginia and North Carolina (Kentucky: Harlan and Bell counties; Virginia: Scott County; Tennessee: Cumberland and Carter counties; North Carolina: Avery and Watauga counties; Suppl. material 7; Fig. 
<figureCitation captionStart="Figure 126" captionStartId="F126" captionText="Figure 126. Distribution map of Nannaria minor species group collection localities in the central Appalachians." figureDoi="10.3897/zookeys.1030.62544.figure126" httpUri="https://binary.pensoft.net/fig/531229" pageId="0" pageNumber="1">126</figureCitation>
). Distribution area: 5,582 km2; status: SRE.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="ecology">
<paragraph pageId="0" pageNumber="1">Ecology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Individuals of 
<taxonomicName class="Dicotyledoneae" family="Ericaceae" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="N. blackmountainensis" order="Ericales" pageId="0" pageNumber="1" phylum="Angiospermae" rank="species" species="blackmountainensis">
<emphasis italics="true" pageId="0" pageNumber="1">N. blackmountainensis</emphasis>
</taxonomicName>
sp. nov. were collected from mesic forests of hemlock, tuliptree, maple, and oak, with an understory of jewelweed and stinging nettle.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">This species is named for its type locality. The specific name is an adjective.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
United States, Kentucky, Harlan Co., Black Mountain summit, radar station access road off Black Mountain Ridge Rd., about 2.4 rd. km E jct w/ KY-160, 
<geoCoordinate degrees="36.9156" direction="north" orientation="latitude" precision="5" value="36.9156">36.9156°N</geoCoordinate>
, - 
<geoCoordinate degrees="82.8930" direction="west" orientation="longitude" precision="5" value="-82.893">82.8930°W</geoCoordinate>
.
</paragraph>
</subSubSection>
</treatment>
</document>