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<document id="0923BD08824681396213F267B83F12E5" ID-DOI="10.5281/zenodo.5339158" ID-ISSN="0374-1036" ID-Zenodo-Dep="5339158" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630183512220" checkinUser="felipe" docAuthor="Švihla, Vladimír" docDate="2012" docId="03F87A534F23FF81F63A0C28A139942E" docLanguage="en" docName="ActaEntMusNatPra.2012.52.suppl.337-346.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 52" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Colobostomoides marshalli subsp. socotraensis Švihla 2012, ssp. nov." docType="treatment" docVersion="4" lastPageNumber="341" masterDocId="FFC1022B4F22FF85F7700E32A368953D" masterDocTitle="A review of the family Oedemeridae (Coleoptera) of the Socotra Archipelago" masterLastPageNumber="346" masterPageNumber="337" pageNumber="338" updateTime="1699178338177" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A7B9D77782219BE158ABEAFAFAAA7D80">A review of the family Oedemeridae (Coleoptera) of the Socotra Archipelago</mods:title>
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<mods:namePart id="AAF2AF576A82AB8C7814A4D25A614BCA">Švihla, Vladimír</mods:namePart>
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<mods:title id="AE2EE0FA36C9C51BA1993BB78EF4FE9F">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="AA8723DC2301A17FDCF7BB707C11C219">2012</mods:date>
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<mods:number id="296CB657D11508B488A163215272A457">2012-12-17</mods:number>
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<treatment id="03F87A534F23FF81F63A0C28A139942E" ID-DOI="http://doi.org/10.5281/zenodo.5415576" ID-Zenodo-Dep="5415576" LSID="urn:lsid:plazi:treatment:03F87A534F23FF81F63A0C28A139942E" httpUri="http://treatment.plazi.org/id/03F87A534F23FF81F63A0C28A139942E" lastPageId="4" lastPageNumber="341" pageId="1" pageNumber="338">
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<paragraph id="8BEECB454F23FF84F63A0C28A000970E" blockId="1.[330,872,538,563]" box="[330,872,538,563]" pageId="1" pageNumber="338">
<heading id="D0A67C294F23FF84F63A0C28A000970E" bold="true" box="[330,872,538,563]" centered="true" fontSize="8" level="2" pageId="1" pageNumber="338" reason="3">
<emphasis id="B92517574F23FF84F63A0C28A000970E" bold="true" box="[330,872,538,563]" pageId="1" pageNumber="338">
<taxonomicName id="4C51B0C64F23FF84F63A0C28A06A970E" authority="Švihla, 2012" authorityName="Švihla" authorityYear="2012" box="[330,770,538,563]" class="Insecta" family="Oedemeridae" genus="Colobostomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="338" phylum="Arthropoda" rank="subSpecies" species="marshalli" status="subsp. nov." subSpecies="socotraensis">
<emphasis id="B92517574F23FF84F63A0C28A06A970E" bold="true" box="[330,770,538,563]" italics="true" pageId="1" pageNumber="338">Colobostomoides marshalli socotraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216AA2C4F23FF84F4790C29A000970E" box="[777,872,539,563]" pageId="1" pageNumber="338" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C34B98CE4F23FF84F55A0C0BA1E09771" box="[554,648,569,588]" pageId="1" pageNumber="338" type="description">
<paragraph id="8BEECB454F23FF84F55A0C0BA1E09771" blockId="1.[554,648,569,588]" box="[554,648,569,588]" pageId="1" pageNumber="338">
(
<figureCitation id="136AD7C04F23FF84F5400C0BA1E99771" box="[560,641,569,588]" captionStart="Figs" captionStartId="2.[151,190,1570,1589]" captionTargetBox="[191,1086,219,1547]" captionTargetId="figure-12@2.[191,1087,219,1547]" captionTargetPageId="2" captionText="Figs. 1–4. General habitus of Socotran Oedemeridae. 1–2 – Colobostomoides marshalli socotraensis ssp. nov.; 3–4 – Dentostomus socotrensis (Švihla, 1986)." figureDoi="http://doi.org/10.5281/zenodo.5339160" httpUri="https://zenodo.org/record/5339160/files/figure.png" pageId="1" pageNumber="338">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34B98CE4F23FF84F7010C51A74597C6" pageId="1" pageNumber="338" type="materials_examined">
<paragraph id="8BEECB454F23FF84F7010C51A282974B" blockId="1.[113,1090,610,1227]" box="[113,490,610,630]" pageId="1" pageNumber="338">
<emphasis id="B92517574F23FF84F7010C51A382974B" bold="true" box="[113,234,610,630]" pageId="1" pageNumber="338">
<typeStatus id="54EA75E74F23FF84F7010C51A3F6974B" box="[113,158,611,630]" pageId="1" pageNumber="338">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3468BD54F23FF84F79F0C51A245974B" box="[239,301,611,630]" name="Yemen" pageId="1" pageNumber="338">Yemen</collectingCountry>
,
<collectingCountry id="F3468BD54F23FF84F6450C51A2DA974B" box="[309,434,611,630]" name="Socotra Island" pageId="1" pageNumber="338">Socotra Island</collectingCountry>
, Neet.
</paragraph>
<paragraph id="8BEECB454F23FF84F7010C4FA74597C6" blockId="1.[113,1090,610,1227]" pageId="1" pageNumber="338">
<emphasis id="B92517574F23FF84F7010C4FA39C97AC" bold="true" box="[113,244,637,657]" pageId="1" pageNumber="338">Type material.</emphasis>
<materialsCitation id="3B39C1184F23FF84F7890C4CA75397AD" box="[249,1083,632,657]" collectingDate="2000-10" collectionCode="NMPC, V" collectorName="Neet &amp; Bejcek, K" country="Yemen" location="Socotra" pageId="1" pageNumber="338" specimenCount="1" typeStatus="holotype">
<typeStatus id="54EA75E74F23FF84F7890C4CA22397B2" box="[249,331,638,657]" pageId="1" pageNumber="338" type="holotype">HOLOTYPE</typeStatus>
: J (
<collectionCode id="ED4053804F23FF84F6020C4FA2DA97AD" box="[370,434,637,656]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="338">NMPC</collectionCode>
), ‘
<collectingCountry id="F3468BD54F23FF84F6B30C4FA16F97AD" box="[451,519,637,656]" name="Yemen" pageId="1" pageNumber="338">Socotra</collectingCountry>
I.,
<date id="FFEFED854F23FF84F5530C4FA13597AD" box="[547,605,637,656]" pageId="1" pageNumber="338" value="2000-10">
<collectingDate id="EFAB146D4F23FF84F5530C4FA13597AD" box="[547,605,637,656]" pageId="1" pageNumber="338" value="2000-10">x.2000</collectingDate>
</date>
/
<collectorName id="26A4AE934F23FF84F51E0C4CA1FE97AC" box="[622,662,638,657]" pageId="1" pageNumber="338">Neet</collectorName>
/
<collectionCode id="ED4053804F23FF84F5D70C4CA1DE97AC" box="[679,694,638,657]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="338" type="Museum">V</collectionCode>
.
<collectorName id="26A4AE934F23FF84F5CE0C4FA07D97AC" box="[702,789,637,657]" pageId="1" pageNumber="338">Bejček, K</collectorName>
. Šťastný lgt. [white label, printed]
</materialsCitation>
’.
<materialsCitation id="3B39C1184F23FF84F7010CAAA2859791" box="[113,493,659,684]" collectionCode="NMPC" country="Yemen" pageId="1" pageNumber="338" specimenCount="1" typeStatus="paratype">
<typeStatus id="54EA75E74F23FF84F7010CAAA3A09797" box="[113,200,664,683]" pageId="1" pageNumber="338" type="paratype">PARATYPES</typeStatus>
(
<collectionCode id="ED4053804F23FF84F7A60CAAA27E9796" box="[214,278,664,683]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="338">NMPC</collectionCode>
), same label data, 2 JJ
</materialsCitation>
; ‘
<materialsCitation id="3B39C1184F23FF84F68D0CAAA04B97FA" collectorName="L. Purchart &amp; J. Vybiral" country="Yemen" latitude="12.445278" location="Wadi Dehlme" longLatPrecision="21" longitude="54.281666" pageId="1" pageNumber="338" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F3468BD54F23FF84F68D0CAAA1269796" box="[509,590,664,683]" name="Yemen" pageId="1" pageNumber="338">YEMEN</collectingCountry>
, SOCOTRA Island SE / sandy beach by
<location id="8E8E9D9E4F23FF84F4CC0CAAA75B9796" LSID="urn:lsid:plazi:treatment:03F87A534F23FF81F63A0C28A139942E:8E8E9D9E4F23FF84F4CC0CAAA75B9796" box="[956,1075,664,683]" country="Yemen" latitude="12.445278" longLatPrecision="21" longitude="54.281666" name="Wadi Dehlme" pageId="1" pageNumber="338">Wadi Dehlme</location>
/
<geoCoordinate id="EE65AD824F23FF84F7010C81A3B197FB" box="[113,217,690,710]" degrees="12" direction="north" minutes="26" orientation="latitude" pageId="1" pageNumber="338" precision="15" seconds="43" value="12.445278">N 12°26′43″</geoCoordinate>
,
<geoCoordinate id="EE65AD824F23FF84F7920C81A22F97FB" box="[226,327,690,710]" degrees="54" direction="east" minutes="16" orientation="longitude" pageId="1" pageNumber="338" precision="15" seconds="54" value="54.281666">E 54°16′54″</geoCoordinate>
/
<collectorName id="26A4AE934F23FF84F6250C81A2DA97FB" box="[341,434,691,710]" pageId="1" pageNumber="338">L. Purchart</collectorName>
&amp;
<collectorName id="26A4AE934F23FF84F6BA0C81A17097FB" box="[458,536,691,710]" pageId="1" pageNumber="338">J.Vybíral</collectorName>
lgt. [white label, printed]’, 3JJ
</materialsCitation>
; ‘
<materialsCitation id="3B39C1184F23FF84F4420C81A29E97DC" collectionCode="SHUAB" country="Yemen" location="YEMEN" pageId="1" pageNumber="338" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F3468BD54F23FF84F4420C81A0EB97FB" box="[818,899,691,710]" name="Yemen" pageId="1" pageNumber="338">YEMEN</collectingCountry>
,
<collectingCountry id="F3468BD54F23FF84F4F90C81A76597F9" box="[905,1037,691,710]" name="SOCOTRA ISLAND" pageId="1" pageNumber="338">SOCOTRA ISLAND</collectingCountry>
/ ca.
<quantity id="4CA966A04F23FF84F3460C81A3E497DD" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="338" unit="km" value="3.0">3 km</quantity>
NE of
<collectionCode id="ED4053804F23FF84F7B60CFFA39497E2" box="[198,252,717,736]" pageId="1" pageNumber="338">SHUAB</collectionCode>
/
<taxonomicName id="4C51B0C64F23FF84F6790CFCA2F497DD" box="[265,412,718,736]" class="Magnoliopsida" family="Acanthaceae" genus="Avicennia" kingdom="Plantae" order="Lamiales" pageId="1" pageNumber="338" phylum="Tracheophyta" rank="species" species="marina">
<emphasis id="B92517574F23FF84F6790CFCA2F497DD" box="[265,412,718,736]" italics="true" pageId="1" pageNumber="338">Avicennia marina</emphasis>
</taxonomicName>
mangrove
</materialsCitation>
; /
<materialsCitation id="3B39C1184F23FF84F5770CFFA74097C1" collectingDate="2012-06-20" collectingDateMax="2012-06-21" collectingDateMin="2012-06-20" collectionCode="V" collectorName="Expedition &amp; J. Bezdek &amp; J. Hajek &amp; Hula &amp; Kment &amp; Malenovsky &amp; J. Niedobova &amp; L. Purchart" country="Yemen" elevation="3" latitude="12.568334" location="SOCOTRA" longLatPrecision="129" longitude="53.398335" pageId="1" pageNumber="338" specimenCount="1" typeStatus="paratype">
sand dunes,
<date id="FFEFED854F23FF84F51C0CFFA18197DD" box="[620,745,717,736]" pageId="1" pageNumber="338" value="2012-06-20" valueMax="2012-06-21" valueMin="2012-06-20">
<collectingDate id="EFAB146D4F23FF84F51C0CFFA18197DD" box="[620,745,717,736]" pageId="1" pageNumber="338" value="2012-06-20" valueMax="2012-06-21" valueMin="2012-06-20">20.–21.vi.2012</collectingDate>
</date>
/
<geoCoordinate id="EE65AD824F23FF84F5870CFFA02497DC" box="[759,844,717,737]" degrees="12" direction="north" minutes="34.1" orientation="latitude" pageId="1" pageNumber="338" precision="92" value="12.568334">12°34.1′N</geoCoordinate>
<geoCoordinate id="EE65AD824F23FF84F4210CFFA0CE97DC" box="[849,934,717,737]" degrees="53" direction="east" minutes="23.9" orientation="longitude" pageId="1" pageNumber="338" precision="92" value="53.398335">53°23.9′E</geoCoordinate>
,
<quantity id="4CA966A04F23FF84F4DD0CFFA0A497DC" box="[941,972,717,737]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="338" unit="m" value="3.0">
<elevation id="007C2C764F23FF84F4DD0CFFA0A497DC" box="[941,972,717,737]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="1" pageNumber="338" unit="m" value="3.0">3 m</elevation>
</quantity>
//
<collectingCountry id="F3468BD54F23FF84F4AF0CFFA72897DD" box="[991,1088,717,736]" name="Yemen" pageId="1" pageNumber="338">SOCOTRA</collectingCountry>
<collectorName id="26A4AE934F23FF84F7010CDAA3A597C6" box="[113,205,744,763]" pageId="1" pageNumber="338">Expedition</collectorName>
2012 /
<collectorName id="26A4AE934F23FF84F67C0CDAA23697C6" box="[268,350,744,763]" pageId="1" pageNumber="338">J. Bezděk</collectorName>
,
<collectorName id="26A4AE934F23FF84F6170CDAA2C597C6" box="[359,429,744,763]" pageId="1" pageNumber="338">J. Hájek</collectorName>
,
<collectionCode id="ED4053804F23FF84F6C50CDAA2AC97C6" box="[437,452,744,763]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="338" type="Museum">V</collectionCode>
.
<collectorName id="26A4AE934F23FF84F6BC0CDAA29F97C6" box="[460,503,744,763]" pageId="1" pageNumber="338">Hula</collectorName>
, / P.
<collectorName id="26A4AE934F23FF84F56D0CDAA13097C6" box="[541,600,744,763]" pageId="1" pageNumber="338">Kment</collectorName>
, I.
<collectorName id="26A4AE934F23FF84F5010CDAA1B397C6" box="[625,731,744,763]" pageId="1" pageNumber="338">Malenovský</collectorName>
, /
<collectorName id="26A4AE934F23FF84F59E0CDAA03497C6" box="[750,860,744,763]" pageId="1" pageNumber="338">J. Niedobová</collectorName>
&amp;
<collectorName id="26A4AE934F23FF84F4060CDAA0BC97C6" box="[886,980,744,763]" pageId="1" pageNumber="338">L. Purchart</collectorName>
leg.’, 1 J
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C34B98CE4F23FF84F7010D21A01296B6" pageId="1" pageNumber="338" type="diagnosis">
<paragraph id="8BEECB454F23FF84F7010D21A1179676" blockId="1.[113,1090,610,1227]" pageId="1" pageNumber="338">
<emphasis id="B92517574F23FF84F7010D21A38E9616" bold="true" box="[113,230,787,811]" pageId="1" pageNumber="338">Diagnosis.</emphasis>
Length (J):
<quantity id="4CA966A04F23FF84F6000D26A1699616" box="[368,513,788,812]" metricMagnitude="-2" metricUnit="m" metricValue="1.18" metricValueMax="1.44" metricValueMin="0.9199999999999999" pageId="1" pageNumber="338" unit="mm" value="11.8" valueMax="14.4" valueMin="9.2">9.2–14.4 mm</quantity>
. Male terminalia similar to those of the nominotypical subspecies (cf.
<bibRefCitation id="EFC0B6B44F23FF84F6650D06A2DA9676" author="VAZQUEZ X. A." box="[277,434,819,843]" pageId="1" pageNumber="338" pagination="83 - 147" refId="ref4653" refString="VAZQUEZ X. A. 1996: Revision of the Southern African Oedemeridae (Coleoptera, Tenebrionoidea). Mitteilungen aus dem Zoologisches Museum in Berlin 72: 83 - 147." type="journal article" year="1996">VÁZQUEZ 1996</bibRefCitation>
), female unknown.
</paragraph>
<paragraph id="8BEECB454F23FF84F7010D61A01296B6" blockId="1.[113,1090,610,1227]" pageId="1" pageNumber="338">
<emphasis id="B92517574F23FF84F7010D61A2029656" bold="true" box="[113,362,851,875]" pageId="1" pageNumber="338">Differential diagnosis.</emphasis>
The newly described subspecies differs from the nominotypical form in the characters mentioned in
<tableCitation id="C6D3FEFE4F23FF84F6C90D41A16096B1" box="[441,520,883,908]" captionStart="Table 1" captionStartId="1.[113,160,1266,1285]" captionText="Table 1. Differential characters of Colobostomoides marshalli marshalli (Blair, 1926) and C. marshalii socotraensis ssp. nov." pageId="1" pageNumber="338">Table 1</tableCitation>
(cf.
<figureCitation id="136AD7C04F23FF84F5490D41A1F596B1" box="[569,669,883,908]" captionStart="Figs" captionStartId="2.[151,190,1570,1589]" captionTargetBox="[191,1086,219,1547]" captionTargetId="figure-12@2.[191,1087,219,1547]" captionTargetPageId="2" captionText="Figs. 1–4. General habitus of Socotran Oedemeridae. 1–2 – Colobostomoides marshalli socotraensis ssp. nov.; 3–4 – Dentostomus socotrensis (Švihla, 1986)." figureDoi="http://doi.org/10.5281/zenodo.5339160" httpUri="https://zenodo.org/record/5339160/files/figure.png" pageId="1" pageNumber="338">Figs. 1–2</figureCitation>
and
<bibRefCitation id="EFC0B6B44F23FF84F5A00D46A00796B6" author="VAZQUEZ X. A." box="[720,879,883,907]" pageId="1" pageNumber="338" pagination="83 - 147" refId="ref4653" refString="VAZQUEZ X. A. 1996: Revision of the Southern African Oedemeridae (Coleoptera, Tenebrionoidea). Mitteilungen aus dem Zoologisches Museum in Berlin 72: 83 - 147." type="journal article" year="1996">VÁZQUEZ 1996</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C34B98CE4F23FF84F7010DA1A078937A" pageId="1" pageNumber="338" type="discussion">
<paragraph id="8BEECB454F23FF84F7010DA1A1FB91F6" blockId="1.[113,1090,610,1227]" pageId="1" pageNumber="338">
<emphasis id="B92517574F23FF84F7010DA1A39B9696" bold="true" box="[113,243,915,939]" pageId="1" pageNumber="338">Comments.</emphasis>
<taxonomicName id="4C51B0C64F23FF84F7880DA7A28E9691" authority="marshalli" authorityName="marshalli (Blair" authorityYear="1926" box="[248,486,917,940]" class="Insecta" family="Oedemeridae" genus="Colobostomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="338" phylum="Arthropoda" rank="subSpecies" species="marshalli" subSpecies="marshalli">
<emphasis id="B92517574F23FF84F7880DA7A28E9691" box="[248,486,917,940]" italics="true" pageId="1" pageNumber="338">C. marshalli marshalli</emphasis>
</taxonomicName>
is hitherto known only from the
<collectingCountry id="F3468BD54F23FF84F44A0DA1A7299696" box="[826,1089,915,939]" name="South Africa" pageId="1" pageNumber="338">Republic of South Africa</collectingCountry>
(
<collectingRegion id="499505A74F23FF84F7080D86A26096F1" box="[120,264,948,972]" country="South Africa" name="Eastern Cape" pageId="1" pageNumber="338">Eastern Cape</collectingRegion>
and
<collectingRegion id="499505A74F23FF84F64B0D81A28A96F6" box="[315,482,947,971]" country="South Africa" name="KwaZulu-Natal" pageId="1" pageNumber="338">KwaZulu-Natal</collectingRegion>
provinces). It is probable, that
<taxonomicName id="4C51B0C64F23FF84F45F0D87A72896F1" authorityName="Švihla" authorityYear="2012" box="[815,1088,949,972]" class="Insecta" family="Oedemeridae" genus="Colobostomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="338" phylum="Arthropoda" rank="subSpecies" species="marshalli" subSpecies="socotraensis">
<emphasis id="B92517574F23FF84F45F0D87A72896F1" box="[815,1088,949,972]" italics="true" pageId="1" pageNumber="338">C. marshalli socotraensis</emphasis>
</taxonomicName>
ssp. nov. was introduced to
<collectingCountry id="F3468BD54F23FF84F6ED0DE6A28696D1" box="[413,494,980,1004]" name="Yemen" pageId="1" pageNumber="338">Socotra</collectingCountry>
by ocean streams and here it differentiated on the subspecific level. However, there is also the possibility, that
<taxonomicName id="4C51B0C64F23FF84F5A10DC7A03E9131" baseAuthorityName="Blair" baseAuthorityYear="1926" box="[721,854,1013,1036]" class="Insecta" family="Oedemeridae" genus="Colobostomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="338" phylum="Arthropoda" rank="species" sensu="lato" species="marshalli">
<emphasis id="B92517574F23FF84F5A10DC7A03E9131" box="[721,854,1013,1036]" italics="true" pageId="1" pageNumber="338">C. marshalli</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216AA2C4F23FF84F42D0DC6A0FF9136" box="[861,919,1011,1035]" pageId="1" pageNumber="338" sensu="lato">s. lat.</taxonomicNameLabel>
originates from
<collectingCountry id="F3468BD54F23FF84F7010A26A3AA9111" box="[113,194,1044,1068]" name="Yemen" pageId="1" pageNumber="338">Socotra</collectingCountry>
(and/or Asian mainland) and it was introduced to southern Africa by the commercial ships commuting, very frequent during the Antiquity and the Middle Ages. Another possibility is that this species is widely distributed from
<collectingCountry id="F3468BD54F23FF84F5E90A61A0499156" box="[665,801,1107,1131]" name="South Africa" pageId="1" pageNumber="338">South Africa</collectingCountry>
to
<collectingCountry id="F3468BD54F23FF84F4330A66A0F89156" box="[835,912,1108,1131]" name="Yemen" pageId="1" pageNumber="338">Yemen</collectingCountry>
, but still has not been discovered in the intermediate areas. This pattern of distribution is known for example in
<taxonomicName id="4C51B0C64F23FF84F7FE0AA7A12C9196" authority="Kaszab, 1956" authorityName="Kaszab" authorityYear="1956" box="[142,580,1171,1196]" class="Insecta" family="Meloidae" genus="Stenoria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="338" phylum="Arthropoda" rank="species" species="muiri">
<emphasis id="B92517574F23FF84F7FE0AA7A2439191" box="[142,299,1173,1196]" italics="true" pageId="1" pageNumber="338">Stenoria muiri</emphasis>
Kaszab, 1956 (Meloidae)
</taxonomicName>
–
<collectingCountry id="F3468BD54F23FF84F5100AA6A1C29196" box="[608,682,1172,1195]" name="Yemen" pageId="1" pageNumber="338">Yemen</collectingCountry>
and southern
<collectingCountry id="F3468BD54F23FF84F4360AA1A0BE9196" box="[838,982,1171,1195]" name="Mozambique" pageId="1" pageNumber="338">Mozambique</collectingCountry>
, with two subspecies scarcely distinct by phenetic characters.
</paragraph>
<caption id="DF2E9BCD4F23FF84F7010AC0A3D5901D" pageId="1" pageNumber="338" startId="1.[113,160,1266,1285]">
<paragraph id="8BEECB454F23FF84F7010AC0A3D5901D" blockId="1.[113,1089,1266,1607]" pageId="1" pageNumber="338">
Table 1. Differential characters of
<taxonomicName id="4C51B0C64F23FF84F6E10AC1A0299038" authority="marshalli (Blair, 1926)" authorityName="marshalli (Blair" authorityYear="1926" baseAuthorityName="Blair" baseAuthorityYear="1926" box="[401,833,1266,1285]" class="Insecta" family="Oedemeridae" genus="Colobostomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="338" phylum="Arthropoda" rank="subSpecies" species="marshalli" subSpecies="marshalli">
<emphasis id="B92517574F23FF84F6E10AC1A1B99038" box="[401,721,1267,1285]" italics="true" pageId="1" pageNumber="338">Colobostomoides marshalli marshalli</emphasis>
(Blair, 1926)
</taxonomicName>
and
<taxonomicName id="4C51B0C64F23FF84F4180AC1A7299038" box="[872,1089,1267,1285]" class="Insecta" family="Oedemeridae" genus="Colobostomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="338" phylum="Arthropoda" rank="subSpecies" species="marshalii" subSpecies="socotraensis">
<emphasis id="B92517574F23FF84F4180AC1A7299038" box="[872,1089,1267,1285]" italics="true" pageId="1" pageNumber="338">C. marshalii socotraensis</emphasis>
</taxonomicName>
ssp. nov.
</paragraph>
</caption>
<paragraph id="8BEECB454F23FF84F7090B67A09B90A0" blockId="1.[113,1089,1266,1607]" pageId="1" pageNumber="338">Pronotum pale lemon yellow with narrow medio-longi- Pronotum either entirely pale lemon yellow or (in one tudinal terra-cotta to sienna stripe, not reaching anterior specimen) almost entirely sienna with narrow anterior, and posterior margin. ventro-lateral and posterior paler margins only.</paragraph>
<paragraph id="8BEECB454F23FF84F7090BCCA078937A" blockId="1.[113,1089,1266,1607]" pageId="1" pageNumber="338">
Elytra more or less darkened submarginally. Elytra either entirely pale lemon yellow or (in
<specimenCount id="9D5700CC4F23FF84F4810BCDA1FF9311" pageId="1" pageNumber="338" type="generic">one specimen</specimenCount>
) almost entirely sienna with narrow sutural and lateral margins only.
</paragraph>
</subSubSection>
<caption id="DF2E9BCD4F20FF87F7E70810A14E936D" ID-DOI="http://doi.org/10.5281/zenodo.5339160" ID-Zenodo-Dep="5339160" httpUri="https://zenodo.org/record/5339160/files/figure.png" pageId="2" pageNumber="339" startId="2.[151,190,1570,1589]" targetBox="[191,1086,219,1547]" targetPageId="2">
<paragraph id="8BEECB454F20FF87F7E70810A14E936D" blockId="2.[151,1129,1570,1616]" pageId="2" pageNumber="339">
Figs. 1–4. General habitus of Socotran
<taxonomicName id="4C51B0C64F20FF87F6840810A1019308" authorityName="Latreille" authorityYear="1810" box="[500,617,1570,1589]" class="Insecta" family="Oedemeridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="339" phylum="Arthropoda" rank="family">Oedemeridae</taxonomicName>
. 1–2 –
<taxonomicName id="4C51B0C64F20FF87F5DE0811A7669308" authority="Švihla, 2012" authorityName="Švihla" authorityYear="2012" box="[686,1038,1571,1589]" class="Insecta" family="Oedemeridae" genus="Colobostomoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="339" phylum="Arthropoda" rank="subSpecies" species="marshalli" status="subsp. nov." subSpecies="socotraensis">
<emphasis id="B92517574F20FF87F5DE0811A7669308" box="[686,1038,1571,1589]" italics="true" pageId="2" pageNumber="339">Colobostomoides marshalli socotraensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216AA2C4F20FF87F3660810A70E9308" box="[1046,1126,1570,1589]" pageId="2" pageNumber="339" rank="subSpecies">ssp. nov.</taxonomicNameLabel>
; 3–4 –
<taxonomicName id="4C51B0C64F20FF87F7BC080CA14A936D" authority="(Svihla, 1986)" baseAuthorityName="Svihla" baseAuthorityYear="1986" box="[204,546,1596,1616]" class="Insecta" family="Oedemeridae" genus="Dentostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="339" phylum="Arthropoda" rank="species" species="socotrensis">
<emphasis id="B92517574F20FF87F7BC080CA2C8936D" box="[204,416,1598,1616]" italics="true" pageId="2" pageNumber="339">Dentostomus socotrensis</emphasis>
(
<bibRefCitation id="EFC0B6B44F20FF87F6DC080EA174936D" author="SVIHLA V." box="[428,540,1596,1616]" pageId="2" pageNumber="339" pagination="141 - 238" refId="ref4395" refString="SVIHLA V. 1986: Revision of the generic classification of the Old World Oedemeridae (Coleoptera). Sbornik Narodniho Muzea v Praze 41 B (1985): 141 - 238." type="journal article" year="1986">Švihla, 1986</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF2E9BCD4F21FF86F7010836A267930E" ID-DOI="http://doi.org/10.5281/zenodo.5339162" ID-Zenodo-Dep="5339162" httpUri="https://zenodo.org/record/5339162/files/figure.png" pageId="3" pageNumber="340" startId="3.[113,152,1540,1559]" targetBox="[160,1041,217,1499]" targetPageId="3">
<paragraph id="8BEECB454F21FF86F7010836A267930E" blockId="3.[113,1088,1540,1587]" pageId="3" pageNumber="340">
Figs.5–8. General habitus of Socotran
<taxonomicName id="4C51B0C64F21FF86F6C70836A143932A" authorityName="Latreille" authorityYear="1810" box="[439,555,1540,1559]" class="Insecta" family="Oedemeridae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="340" phylum="Arthropoda" rank="family">Oedemeridae</taxonomicName>
. 5 –
<taxonomicName id="4C51B0C64F21FF86F53F0837A0FB9325" authority="(Svihla, 1987)" baseAuthorityName="Svihla" baseAuthorityYear="1987" box="[591,915,1540,1560]" class="Insecta" family="Oedemeridae" genus="Dentostomus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="340" phylum="Arthropoda" rank="species" species="guichardi">
<emphasis id="B92517574F21FF86F53F0837A07C932A" box="[591,788,1541,1559]" italics="true" pageId="3" pageNumber="340">Dentostomus guichardi</emphasis>
(
<bibRefCitation id="EFC0B6B44F21FF86F46E0836A0E59325" author="SVIHLA V." box="[798,909,1540,1560]" pageId="3" pageNumber="340" pagination="1 - 27" refId="ref4429" refString="SVIHLA V. 1987: Contribution to the knowledge of the Old World Oedemeridae (Coleoptera). Annotationes Zoologicae et Botanicae (Bratislava) 181: 1 - 27." type="journal article" year="1987">Švihla, 1987</bibRefCitation>
)
</taxonomicName>
; 6–8 –
<taxonomicName id="4C51B0C64F21FF86F4BD0837A3AE930F" authority="Švihla, 2012" authorityName="Švihla" authorityYear="2012" class="Insecta" family="Oedemeridae" genus="Anisochrodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="340" phylum="Arthropoda" rank="species" species="janavlada" status="sp. nov.">
<emphasis id="B92517574F21FF86F4BD0837A3AE930F" italics="true" pageId="3" pageNumber="340">Anisochrodes janavlada</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A216AA2C4F21FF86F7BC0812A267930E" box="[204,271,1568,1587]" pageId="3" pageNumber="340" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<subSubSection id="C34B98CE4F26FF81F7E70EEEA1F795CE" box="[151,671,219,244]" pageId="4" pageNumber="341" type="etymology">
<paragraph id="8BEECB454F26FF81F7E70EEEA1F795CE" blockId="4.[151,671,219,276]" box="[151,671,219,244]" pageId="4" pageNumber="341">
<emphasis id="B92517574F26FF81F7E70EEEA27095C9" bold="true" box="[151,280,220,244]" pageId="4" pageNumber="341">Etymology.</emphasis>
Named according to its distribution.
</paragraph>
</subSubSection>
<subSubSection id="C34B98CE4F26FF81F7E70EC9A139942E" box="[151,593,251,276]" pageId="4" pageNumber="341" type="distribution">
<paragraph id="8BEECB454F26FF81F7E70EC9A139942E" blockId="4.[151,671,219,276]" box="[151,593,251,276]" pageId="4" pageNumber="341">
<emphasis id="B92517574F26FF81F7E70EC9A242942E" bold="true" box="[151,298,251,275]" pageId="4" pageNumber="341">Distribution.</emphasis>
Endemic to
<collectingCountry id="F3468BD54F26FF81F6C10ECEA126942E" box="[433,590,251,276]" name="Socotra Island" pageId="4" pageNumber="341">Socotra Island</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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