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<document id="586521787704934BDA5B8F8828E90452" ID-DOI="10.26107/RBZ-2020-0001" ID-ISSN="2345-7600" ID-Zenodo-Dep="4577165" ID-ZooBank="4A76F694-DD52-48CC-A0D4-C1AD9D33B0A1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614784698889" checkinUser="carolina" docAuthor="Páll-Gergely, Barna, Reischütz, Alexander, Maassen, Wim J. M., Grego, Jozef &amp; Hunyadi, András" docDate="2020" docId="03EE1927C50C1402FC98F95DFE021A9D" docLanguage="en" docName="RafflesBZool.68.1-13.pdf" docOrigin="Raffles Bulletin of Zoology 68" docStyle="DocumentStyle:AF09A0581917EACD12E7F0991C3E7C26.5:RafflesBZool.2019-.journal_article" docStyleId="AF09A0581917EACD12E7F0991C3E7C26" docStyleName="RafflesBZool.2019-.journal_article" docStyleVersion="5" docTitle="Sinoennea variabilis Pall-Gergely &amp; Grego 2020, new species" docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="FFD7615FC5041409FFDAFFC0FF961E28" masterDocTitle="New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)" masterLastPageNumber="13" masterPageNumber="1" pageNumber="9" updateTime="1698878575407" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
<mods:mods id="A584B9AAFFA1C41834F1DEE5EEDE3EA0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E39B65CD165A330AE5841D96A4BE909E">
<mods:title id="DD89B89BD2E3555D4F43C0E072225395">New taxa of Diapheridae Panha &amp; Naggs in Sutcharit et al., 2010 from Laos and Thailand (Gastropoda: Eupulmonata: Stylommatophora)</mods:title>
</mods:titleInfo>
<mods:name id="B74796FAD0A58DE5EA23E236AAF94496" type="personal">
<mods:role id="F2087C2BEB332BFEA833BC60D19EA648">
<mods:roleTerm id="D87AE88921FE5BBBAE9AE56C38758756">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44F369D1B314A33A669E62833680747B">Páll-Gergely, Barna</mods:namePart>
</mods:name>
<mods:name id="629FCA3DDE1671134E5D750717A1095D" type="personal">
<mods:role id="61E699F009ADF0527C33CAA38E9047A2">
<mods:roleTerm id="17AA747B3726CE7D7AD68C2073B057F5">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="26121A75C908E072FFBAA12E094A5CF0">Reischütz, Alexander</mods:namePart>
</mods:name>
<mods:name id="9CFCB56AA2B89B4600406181D2DD4C16" type="personal">
<mods:role id="F75962A7909C265DD5C0D5DB4A875557">
<mods:roleTerm id="F9FB6FA5B938646473749DF364BB3DCC">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3FA4890B9136B0E62CC68A4BDFBF3697">Maassen, Wim J. M.</mods:namePart>
</mods:name>
<mods:name id="2D5F7B9D6D6938AC2DA8FB6BD1A7252A" type="personal">
<mods:role id="A1670C7B5729C31E91D32209C4DA064E">
<mods:roleTerm id="EE33DC9B389F7B4BFB284E1DF83E11B8">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9A5D4155CFDDC758A28D94EC059E8808">Grego, Jozef</mods:namePart>
</mods:name>
<mods:name id="ED877A6618D9858121FF387545F4A16C" type="personal">
<mods:role id="8971B3D507AC15DFA1B664E6C8A1E75C">
<mods:roleTerm id="2DDAD08B7DA670A4A5900AF097B7601E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4CEA10935D17287F7323D82B50E45703">Hunyadi, András</mods:namePart>
</mods:name>
<mods:typeOfResource id="4538444E80E5A161EC2B90DBCFAA388C">text</mods:typeOfResource>
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<mods:title id="0CFFC2C505FA5F17658C1A8DBA834B9F">Raffles Bulletin of Zoology</mods:title>
</mods:titleInfo>
<mods:part id="12870457ACF473994176D901B29D6487">
<mods:date id="08E1A97E6C90146B1BA8687AFDFECBEA">2020</mods:date>
<mods:detail id="631B620585438250DCD7931BAA8B4A1E" type="pubDate">
<mods:number id="0D60A8F002AF20F26A673A66DC1F5484">2020-01-15</mods:number>
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<mods:detail id="7F24CA7C2F4B207C1D75941B9186DA1B" type="volume">
<mods:number id="86902BA5FA081F612162F42F01890401">68</mods:number>
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<mods:start id="A8EB685D8189E50DAD986F37D2C54C13">1</mods:start>
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<mods:classification id="04E3D187D9D7EAD07592AF5397E43FC0">journal article</mods:classification>
<mods:identifier id="A6175C815755A6EE04BD4AF42AF68610" type="DOI">10.26107/RBZ-2020-0001</mods:identifier>
<mods:identifier id="5AA292DCC726EAA1C4F4E41406A71E9F" type="ISSN">2345-7600</mods:identifier>
<mods:identifier id="D1C44C0AE574838AA1EE18C9AD835473" type="Zenodo-Dep">4577165</mods:identifier>
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</mods:mods>
<treatment id="03EE1927C50C1402FC98F95DFE021A9D" LSID="urn:lsid:plazi:treatment:03EE1927C50C1402FC98F95DFE021A9D" httpUri="http://treatment.plazi.org/id/03EE1927C50C1402FC98F95DFE021A9D" lastPageId="11" lastPageNumber="12" pageId="8" pageNumber="9">
<subSubSection id="C35DFBBAC50C1401FC98F95DFA54189D" box="[834,1474,1692,1717]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BF8A831C50C1401FC98F95DFA54189D" blockId="8.[834,1474,1692,1749]" box="[834,1474,1692,1717]" pageId="8" pageNumber="9">
<heading id="D0B01F5DC50C1401FC98F95DFA54189D" bold="true" box="[834,1474,1692,1717]" centered="true" fontSize="10" level="2" pageId="8" pageNumber="9" reason="2">
<emphasis id="B9337423C50C1401FC98F95DFA54189D" bold="true" box="[834,1474,1692,1717]" pageId="8" pageNumber="9">
<taxonomicName id="4C47D3B2C50C1401FC98F95DFAB9189D" authority="Pall-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego" authorityYear="2020" box="[834,1327,1692,1717]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="8" pageNumber="9" phylum="Mollusca" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B9337423C50C1401FC98F95DFBB0189D" bold="true" box="[834,1062,1693,1717]" italics="true" pageId="8" pageNumber="9">Sinoennea variabilis</emphasis>
Páll-Gergely &amp; Grego
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50C1401FAE6F95DFA54189D" box="[1340,1474,1693,1717]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC50C1401FBF4F97DFB4018FD" box="[1070,1238,1725,1749]" pageId="8" pageNumber="9" type="description">
<paragraph id="8BF8A831C50C1401FBF4F97DFB4018FD" blockId="8.[834,1474,1692,1749]" box="[1070,1238,1725,1749]" pageId="8" pageNumber="9">
(
<figureCitation id="137CB4B4C50C1401FBEFF97DFB1518FD" box="[1077,1155,1725,1749]" captionStart="Fig" captionStartId="9.[113,148,1915,1937]" captionTargetBox="[113,1451,338,1900]" captionTargetId="figure-11@9.[113,1451,338,1900]" captionTargetPageId="9" captionText="Fig. 5. Sinoennea variabilis Páll-Gergely &amp; Grego, new species. A, holotype (3L07); B, 4L07 (not paratype); C, JG2B (not paratype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577177" httpUri="https://zenodo.org/record/4577177/files/figure.png" pageId="8" pageNumber="9">Figs. 5</figureCitation>
,
<figureCitation id="137CB4B4C50C1401FB4AF97DFB0A18FD" box="[1168,1180,1725,1749]" captionStart="Fig" captionStartId="10.[135,170,1193,1215]" captionTargetBox="[135,1473,190,1177]" captionTargetId="figure-112@10.[135,1473,190,1177]" captionTargetPageId="10" captionText="Fig. 6. Synoptic view of Sinoennea and Laoennea, new genus, species of Laos (all species shown) and northern Thailand. A, Laoennea carychioides Páll-Gergely, A. Reischütz &amp; Maassen, new species; B, Sinoennea infantilis Páll-Gergely &amp; Grego, new species; C–E, Sinoennea variabilis Páll-Gergely &amp; Grego, new species; F, Sinoennea sutchariti Páll-Gergely &amp; Hunyadi, new species; G, Sinoennea panhai Páll-Gergely &amp; Hunyadi, new species; H, Sinoennea euryomphala Inkhavilay &amp; Panha, 2016; I, Sinoennea ljudmilena Páll-Gergely, new species; J, Sinoennea lizae Maassen, 2008; K, Sinoennea otostoma Páll-Gergely, A. Reischütz &amp; Maassen, new species; L, Sinoennea angustistoma Páll-Gergely, A. Reischütz &amp; Maassen, new species. Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577179" httpUri="https://zenodo.org/record/4577179/files/figure.png" pageId="8" pageNumber="9">6</figureCitation>
C–E)
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC50C1403FC98F93DFE2A18F6" lastPageId="10" lastPageNumber="11" pageId="8" pageNumber="9" type="materials_examined">
<paragraph id="8BF8A831C50C1401FC98F93DFBAD163D" blockId="8.[834,1474,1789,2069]" pageId="8" pageNumber="9">
<emphasis id="B9337423C50C1401FC98F93DFC62193D" bold="true" box="[834,1012,1789,1813]" pageId="8" pageNumber="9">Type material.</emphasis>
<materialsCitation id="3B2FA26CC50C1401FC25F93DFBAD163D" collectingDate="2007-11-25" collectionCode="H, D" collectorName="A. Abdou &amp; I. V. Muratov" country="Laos" county="South-Central" elevation="150" latitude="17.4326" location="Muang" longLatPrecision="1" longitude="105.16115" municipality="Thakhek" pageId="8" pageNumber="9" specimenCount="3" stateProvince="Khammouan" typeStatus="holotype">
<typeStatus id="54FC1693C50C1401FC25F93DFBF0193D" box="[1023,1126,1789,1813]" pageId="8" pageNumber="9" type="holotype">Holotype</typeStatus>
(1 shell,
<collectionCode id="ED5630F4C50C1401FB00F93EFB67193D" box="[1242,1265,1790,1813]" pageId="8" pageNumber="9">H</collectionCode>
:
<quantity id="4CBF05D4C50C1401FADBF93DFAFF193D" box="[1281,1385,1789,1813]" metricMagnitude="-3" metricUnit="m" metricValue="1.96" pageId="8" pageNumber="9" unit="mm" value="1.96">1.96 mm</quantity>
,
<collectionCode id="ED5630F4C50C1401FAA3F93EFA06193D" box="[1401,1424,1790,1813]" pageId="8" pageNumber="9">D</collectionCode>
:
<quantity id="4CBF05D4C50C1401FA45F93DFCE6191D" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="8" pageNumber="9" unit="mm" value="1.1">1.1 mm</quantity>
;
<figureCitation id="137CB4B4C50C1401FC5AF8DDFC4C191D" box="[896,986,1821,1845]" captionStart="Fig" captionStartId="9.[113,148,1915,1937]" captionTargetBox="[113,1451,338,1900]" captionTargetId="figure-11@9.[113,1451,338,1900]" captionTargetPageId="9" captionText="Fig. 5. Sinoennea variabilis Páll-Gergely &amp; Grego, new species. A, holotype (3L07); B, 4L07 (not paratype); C, JG2B (not paratype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577177" httpUri="https://zenodo.org/record/4577177/files/figure.png" pageId="8" pageNumber="9">Fig. 5A</figureCitation>
) (MNHN-IM-2012-27279),
<collectingCounty id="6299D0BDC50C1401FAF9F8DDFA56191D" box="[1315,1472,1821,1845]" pageId="8" pageNumber="9">South-Central</collectingCounty>
<collectingCountry id="F350E8A1C50C1401FC99F8FEFCEC197D" box="[835,890,1854,1877]" name="Laos" pageId="8" pageNumber="9">Laos</collectingCountry>
,
<collectingRegion id="498366D3C50C1401FC52F8FDFB17197D" box="[904,1153,1853,1877]" country="Laos" name="Khammouan" pageId="8" pageNumber="9">Khammouan Province</collectingRegion>
, ca.
<quantity id="4CBF05D4C50C1401FB62F8FDFB69197D" box="[1208,1279,1853,1877]" metricMagnitude="4" metricUnit="m" metricValue="3.7" pageId="8" pageNumber="9" unit="km" value="37.0">37 km</quantity>
east-northeast of
<collectingMunicipality id="6B9C324BC50C1401FC99F89DFC36195D" box="[835,928,1885,1909]" pageId="8" pageNumber="9">Thakhek</collectingMunicipality>
(
<location id="8E98FEEAC50C1401FC74F89DFC6D195D" LSID="urn:lsid:plazi:treatment:03EE1927C50C1402FC98F95DFE021A9D:8E98FEEAC50C1401FC74F89DFC6D195D" box="[942,1019,1885,1909]" country="Laos" county="South-Central" latitude="17.4326" longLatPrecision="1" longitude="105.16115" municipality="Thakhek" name="Muang" pageId="8" pageNumber="9" stateProvince="Khammouan">Muang</location>
<collectingRegion id="498366D3C50C1401FBD9F89DFB07195D" box="[1027,1169,1885,1909]" country="Laos" name="Khammouan" pageId="8" pageNumber="9">Khammouan</collectingRegion>
), ca.
<quantity id="4CBF05D4C50C1401FB10F89DFA83195D" box="[1226,1301,1885,1909]" metricMagnitude="3" metricUnit="m" metricValue="4.5" pageId="8" pageNumber="9" unit="km" value="4.5">4.5 km</quantity>
west-northwest of
<location id="8E98FEEAC50C1401FCBEF8BDFC5019BD" LSID="urn:lsid:plazi:treatment:03EE1927C50C1402FC98F95DFE021A9D:8E98FEEAC50C1401FCBEF8BDFC5019BD" box="[868,966,1917,1941]" country="Laos" county="South-Central" latitude="17.4326" longLatPrecision="1" longitude="105.16115" municipality="Thakhek" name="Mahaxai" pageId="8" pageNumber="9" stateProvince="Khammouan">Mahaxai</location>
, on and under rocks in dry secondary forest under E exposed cliff,
<geoCoordinate id="EE73CEF6C50C1401FBE3F85CFB55199D" box="[1081,1219,1948,1973]" degrees="17" direction="north" minutes="25.956" orientation="latitude" pageId="8" pageNumber="9" precision="1" value="17.4326">17°25.956′N</geoCoordinate>
,
<geoCoordinate id="EE73CEF6C50C1401FB0AF85CFAF2199D" box="[1232,1380,1948,1973]" degrees="105" direction="east" minutes="09.669" orientation="longitude" pageId="8" pageNumber="9" precision="1" value="105.16115">105°09.669′E</geoCoordinate>
, alt.
<quantity id="4CBF05D4C50C1401FA43F85DFCCF19FD" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="m" value="150.0">
<elevation id="006A4F02C50C1401FA43F85DFCCF19FD" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="8" pageNumber="9" unit="m" value="150.0">150 m</elevation>
</quantity>
, locality code: 3L07, coll.
<collectorName id="26B2CDE7C50C1401FBA6F87EFB7119FD" box="[1148,1255,1981,2006]" pageId="8" pageNumber="9">A. Abdou</collectorName>
&amp;
<collectorName id="26B2CDE7C50C1401FAD1F87EFA0D19FD" box="[1291,1435,1982,2005]" pageId="8" pageNumber="9">I.V. Muratov</collectorName>
,
<date id="FFF98EF1C50C1401FA7CF87DFC6B19DD" pageId="8" pageNumber="9" value="2007-11-25">
<collectingDate id="EFBD7719C50C1401FA7CF87DFC6B19DD" pageId="8" pageNumber="9" value="2007-11-25">25 November 2007</collectingDate>
</date>
;
<specimenCount id="9D4163B8C50C1401FBD6F81DFB1819DE" box="[1036,1166,2013,2038]" pageId="8" pageNumber="9" type="generic" typeStatus="paratypes">3 paratypes</specimenCount>
(MNHN-IM-2012-27280), same data as holotype.
</materialsCitation>
</paragraph>
<caption id="DF38F8B9C50D1400FFABF8BBFEEB1986" ID-DOI="http://doi.org/10.5281/zenodo.4577177" ID-Zenodo-Dep="4577177" httpUri="https://zenodo.org/record/4577177/files/figure.png" pageId="9" pageNumber="10" startId="9.[113,148,1915,1937]" targetBox="[113,1451,338,1900]" targetPageId="9">
<paragraph id="8BF8A831C50D1400FFABF8BBFEEB1986" blockId="9.[113,1453,1915,1966]" pageId="9" pageNumber="10">
Fig. 5.
<taxonomicName id="4C47D3B2C50D1400FF61F8BCFDFF19B9" authority="Pall-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego" authorityYear="2020" box="[187,617,1915,1937]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="9" pageNumber="10" phylum="Mollusca" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B9337423C50D1400FF61F8BCFE1519B8" box="[187,387,1915,1937]" italics="true" pageId="9" pageNumber="10">Sinoennea variabilis</emphasis>
Páll-Gergely &amp; Grego
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50D1400FDACF8BCFD7819B9" box="[630,750,1915,1937]" pageId="9" pageNumber="10" rank="species">new species</taxonomicNameLabel>
. A, holotype (3L07); B, 4L07 (not paratype); C, JG2B (not paratype). Scale bar represents 1 mm.
</paragraph>
</caption>
<caption id="DF38F8B9C50E1403FF5DFB69FBBE1B66" ID-DOI="http://doi.org/10.5281/zenodo.4577179" ID-Zenodo-Dep="4577179" httpUri="https://zenodo.org/record/4577179/files/figure.png" pageId="10" pageNumber="11" startId="10.[135,170,1193,1215]" targetBox="[135,1473,190,1177]" targetPageId="10">
<paragraph id="8BF8A831C50E1403FF5DFB69FBBE1B66" blockId="10.[135,1474,1193,1359]" pageId="10" pageNumber="11">
Fig. 6. Synoptic view of
<taxonomicName id="4C47D3B2C50E1403FEA5FB69FE751A96" authorityName="Kobelt" authorityYear="1904" box="[383,483,1193,1214]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus">
<emphasis id="B9337423C50E1403FEA5FB69FE751A96" box="[383,483,1193,1214]" italics="true" pageId="10" pageNumber="11">Sinoennea</emphasis>
</taxonomicName>
and
<taxonomicName id="4C47D3B2C50E1403FDCCFB69FDE31A96" authority="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi, 2020" authorityName="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi" authorityYear="2020" box="[534,629,1193,1214]" class="Gastropoda" family="Diapheridae" genus="Laoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9337423C50E1403FDCCFB69FDE31A96" box="[534,629,1193,1214]" italics="true" pageId="10" pageNumber="11">Laoennea</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FD58FB69FD7A1A96" box="[642,748,1193,1214]" pageId="10" pageNumber="11" rank="genus">new genus</taxonomicNameLabel>
, species of Laos (all species shown) and northern Thailand. A,
<taxonomicName id="4C47D3B2C50E1403FAB9FB69FD011AF3" authority="Pall-Gergely, A. Reischutz &amp; Maassen" authorityName="Pall-Gergely, A. Reischutz &amp; Maassen" authorityYear="2020" class="Gastropoda" family="Diapheridae" genus="Laoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="carychioides" status="sp. nov.">
<emphasis id="B9337423C50E1403FAB9FB69FE951AF2" italics="true" pageId="10" pageNumber="11">Laoennea carychioides</emphasis>
Páll-Gergely, A. Reischütz &amp; Maassen
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FD7EFB06FC881AF3" box="[676,798,1221,1243]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
; B,
<taxonomicName id="4C47D3B2C50E1403FC91FB06FB6D1AF3" authority="Pall-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" box="[843,1275,1221,1243]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="infantilis" status="sp. nov.">
<emphasis id="B9337423C50E1403FC91FB06FB871AF2" box="[843,1041,1221,1243]" italics="true" pageId="10" pageNumber="11">Sinoennea infantilis</emphasis>
Páll-Gergely &amp; Grego
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FAD2FB06FA151AF3" box="[1288,1411,1221,1243]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
; C–E,
<taxonomicName id="4C47D3B2C50E1403FF5DFB23FDA01ADF" authority="Pall-Gergely &amp; Grego" authorityName="Pall-Gergely &amp; Grego" authorityYear="2020" box="[135,566,1250,1272]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B9337423C50E1403FF5DFB23FEC61ADF" box="[135,336,1250,1272]" italics="true" pageId="10" pageNumber="11">Sinoennea variabilis</emphasis>
Páll-Gergely &amp; Grego
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FD99FB23FD2A1AD0" box="[579,700,1250,1272]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
; F,
<taxonomicName id="4C47D3B2C50E1403FD3FFB23FB3D1AD0" authority="Pall-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" box="[741,1195,1250,1272]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="sutchariti" status="sp. nov.">
<emphasis id="B9337423C50E1403FD3FFB23FC381ADF" box="[741,942,1250,1272]" italics="true" pageId="10" pageNumber="11">Sinoennea sutchariti</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FB62FB23FAA71AD0" box="[1208,1329,1250,1272]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
; G,
<taxonomicName id="4C47D3B2C50E1403FA84FB23FE2B1B3D" authority="Pall-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="panhai" status="sp. nov.">
<emphasis id="B9337423C50E1403FA84FB23FF5C1B3C" italics="true" pageId="10" pageNumber="11">Sinoennea panhai</emphasis>
Páll-Gergely &amp; Hunyadi
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FE1CFAC0FDAA1B3D" box="[454,572,1279,1301]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
; H,
<taxonomicName id="4C47D3B2C50E1403FDB8FB3FFBD81B3D" authority="Inkhavilay &amp; Panha, 2016" authorityName="Inkhavilay &amp; Panha in Inkhavilay, Sutcharit, Tongkerd &amp; Panha" authorityYear="2016" box="[610,1102,1279,1301]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="euryomphala">
<emphasis id="B9337423C50E1403FDB8FB3FFCDD1B3C" box="[610,843,1279,1300]" italics="true" pageId="10" pageNumber="11">Sinoennea euryomphala</emphasis>
<bibRefCitation id="EFD6D5C0C50E1403FC8AFB3FFBD81B3D" author="Inkhavilay K &amp; Sutcharit C &amp; Tongkerd P &amp; Panha S" box="[848,1102,1279,1301]" pageId="10" pageNumber="11" pagination="213 - 232" refId="ref7929" refString="Inkhavilay K, Sutcharit C, Tongkerd P &amp; Panha S (2016) New species of micro snails from Laos (Pulmonata: Vertiginidae and Diapheridae). Journal of Conchology, 42 (4): 213 - 232." type="journal article" year="2016">Inkhavilay &amp; Panha, 2016</bibRefCitation>
</taxonomicName>
; I,
<taxonomicName id="4C47D3B2C50E1403FBB1FB3FFA281B3D" authority="Pall-Gergely" authorityName="Pall-Gergely in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" box="[1131,1470,1279,1301]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="ljudmilena" status="sp. nov.">
<emphasis id="B9337423C50E1403FBB1FB3FFAAD1B3C" box="[1131,1339,1279,1300]" italics="true" pageId="10" pageNumber="11">Sinoennea ljudmilena</emphasis>
Páll-Gergely
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FF5DFADCFF6B1B1A" box="[135,253,1308,1330]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
; J,
<taxonomicName id="4C47D3B2C50E1403FEC4FADCFDD91B19" authority="Maassen, 2008" authorityName="Maassen" authorityYear="2008" box="[286,591,1308,1329]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="lizae">
<emphasis id="B9337423C50E1403FEC4FADCFE231B19" box="[286,437,1308,1329]" italics="true" pageId="10" pageNumber="11">Sinoennea lizae</emphasis>
<bibRefCitation id="EFD6D5C0C50E1403FE61FADCFDD91B19" author="Maassen WJM" box="[443,591,1308,1329]" pageId="10" pageNumber="11" pagination="233 - 240" refId="ref8222" refString="Maassen WJM (2008) Remarks on a small collection of terrestrial molluscs from north-west Laos, with descriptions of three new species (Mollusca: Pulmonata: Streptaxidae, Vertiginidae). Basteria, 72: 233 - 240." type="journal article" year="2008">Maassen, 2008</bibRefCitation>
</taxonomicName>
; K,
<taxonomicName id="4C47D3B2C50E1403FDADFADCFB2E1B19" authority="Pall-Gergely, A. Reischutz &amp; Maassen" authorityName="Pall-Gergely, A. Reischutz &amp; Maassen in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" box="[631,1208,1308,1330]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="otostoma" status="sp. nov.">
<emphasis id="B9337423C50E1403FDADFADCFCAF1B1A" box="[631,825,1308,1330]" italics="true" pageId="10" pageNumber="11">Sinoennea otostoma</emphasis>
Páll-Gergely, A. Reischütz &amp; Maassen
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FB18FADCFAAE1B1A" box="[1218,1336,1308,1330]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
; L,
<taxonomicName id="4C47D3B2C50E1403FA84FADCFD1B1B66" authority="Pall-Gergely, A. Reischutz &amp; Maassen" authorityName="Pall-Gergely, A. Reischutz &amp; Maassen in Pall-Gergely, A. Reischutz, Maassen, Grego &amp; Hunyadi" authorityYear="2020" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="10" pageNumber="11" phylum="Mollusca" rank="species" species="angustistoma" status="sp. nov.">
<emphasis id="B9337423C50E1403FA84FADCFE911B66" italics="true" pageId="10" pageNumber="11">Sinoennea angustistoma</emphasis>
Páll-Gergely, A. Reischütz &amp; Maassen
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50E1403FD43FAF9FC871B67" box="[665,785,1337,1359]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
. Scale bar represents 1 mm.
</paragraph>
</caption>
<paragraph id="8BF8A831C50E1403FF5DFA46FE2A18F6" blockId="10.[135,774,1414,1758]" pageId="10" pageNumber="11">
<emphasis id="B9337423C50E1403FF5DFA46FEEA1BB6" bold="true" box="[135,380,1414,1438]" pageId="10" pageNumber="11">Additional material.</emphasis>
<materialsCitation id="3B2FA26CC50E1403FE5DFA46FE561876" collectingDate="2007-11-25" collectorName="A. Abdou &amp; I. V. Muratov" country="Laos" county="Thakhek" elevation="169" latitude="17.4456" location="Mahaxai" longLatPrecision="1" longitude="105.139336" municipality="Muang" pageId="10" pageNumber="11" specimenCount="1" stateProvince="Khammouan">
1 shell (MNHN-IM-2012-27281), south-central
<collectingCountry id="F350E8A1C50E1403FEC6FA66FEC41B95" box="[284,338,1446,1469]" name="Laos" pageId="10" pageNumber="11">Laos</collectingCountry>
,
<collectingRegion id="498366D3C50E1403FE84FA66FDC51B96" box="[350,595,1446,1470]" country="Laos" name="Khammouan" pageId="10" pageNumber="11">Khammouan Province</collectingRegion>
, ca.
<quantity id="4CBF05D4C50E1403FD5CFA66FD5D1B96" box="[646,715,1446,1470]" metricMagnitude="4" metricUnit="m" metricValue="3.5" pageId="10" pageNumber="11" unit="km" value="35.0">35 km</quantity>
eastnortheast of
<collectingCounty id="6299D0BDC50E1403FED6FA06FEFF1BF6" box="[268,361,1478,1502]" pageId="10" pageNumber="11">Thakhek</collectingCounty>
(
<collectingMunicipality id="6B9C324BC50E1403FEADFA06FE521BF6" box="[375,452,1478,1502]" pageId="10" pageNumber="11">Muang</collectingMunicipality>
<collectingRegion id="498366D3C50E1403FE11FA06FDCF1BF6" box="[459,601,1478,1502]" country="Laos" name="Khammouan" pageId="10" pageNumber="11">Khammouan</collectingRegion>
), ca.
<quantity id="4CBF05D4C50E1403FD4AFA06FD501BF6" box="[656,710,1478,1502]" metricMagnitude="3" metricUnit="m" metricValue="7.0" pageId="10" pageNumber="11" unit="km" value="7.0">7 km</quantity>
westnorthwest of
<location id="8E98FEEAC50E1403FECFFA26FEE31BD6" LSID="urn:lsid:plazi:treatment:03EE1927C50C1402FC98F95DFE021A9D:8E98FEEAC50E1403FECFFA26FEE31BD6" box="[277,373,1510,1534]" country="Laos" county="Thakhek" latitude="17.4456" longLatPrecision="1" longitude="105.139336" municipality="Muang" name="Mahaxai" pageId="10" pageNumber="11" stateProvince="Khammouan">Mahaxai</location>
, on and under rocks in dry secondary forest under south exposed cliff,
<geoCoordinate id="EE73CEF6C50E1403FE06F9C4FDF31835" box="[476,613,1540,1565]" degrees="17" direction="north" minutes="26.736" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="17.4456">17°26.736′N</geoCoordinate>
,
<geoCoordinate id="EE73CEF6C50E1403FDB5F9C4FC941835" box="[623,770,1540,1565]" degrees="105" direction="east" minutes="08.360" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="105.139336">105°08.360′E</geoCoordinate>
, alt.
<elevation id="006A4F02C50E1403FF6AF9E6FEB81816" box="[176,302,1574,1598]" metricMagnitude="2" metricUnit="m" metricValue="1.69" pageId="10" pageNumber="11" unit="m" value="169.0">
<quantity id="4CBF05D4C50E1403FF6AF9E6FF621815" box="[176,244,1574,1598]" metricMagnitude="2" metricUnit="m" metricValue="1.69" pageId="10" pageNumber="11" unit="m" value="169.0">169 m</quantity>
a.s.l.
</elevation>
, locality code: 4L07, coll.
<collectorName id="26B2CDE7C50E1403FD8AF9E6FD2F1816" box="[592,697,1574,1598]" pageId="10" pageNumber="11">A. Abdou</collectorName>
&amp;
<collectorName id="26B2CDE7C50E1403FD06F9E6FF701875" pageId="10" pageNumber="11">I.V. Muratov</collectorName>
,
<date id="FFF98EF1C50E1403FF2AF986FE561876" box="[240,448,1606,1630]" pageId="10" pageNumber="11" value="2007-11-25">
<collectingDate id="EFBD7719C50E1403FF2AF986FE561876" box="[240,448,1606,1630]" pageId="10" pageNumber="11" value="2007-11-25">25 November 2007</collectingDate>
</date>
</materialsCitation>
;
<materialsCitation id="3B2FA26CC50E1403FE10F986FE2E18F6" collectingDate="2017-02-11" collectionCode="JG" collectorName="J. Grego" country="Laos" county="South-Central" elevation="160" latitude="17.563583" location="Dome" longLatPrecision="1" longitude="104.871635" municipality="Tham Don Cave" pageId="10" pageNumber="11" specimenCode="JG/2, JG2" specimenCount="1" stateProvince="Khammouan">
2 shells (
<specimenCode id="DBE1004AC50E1403FDFDF986FDC81876" box="[551,606,1606,1630]" collectionCode="JG" pageId="10" pageNumber="11">JG/2</specimenCode>
),
<collectingCounty id="6299D0BDC50E1403FDABF986FC901876" box="[625,774,1606,1630]" pageId="10" pageNumber="11">South-Central</collectingCounty>
<collectingCountry id="F350E8A1C50E1403FF5DF9A6FF2B1855" box="[135,189,1638,1661]" name="Laos" pageId="10" pageNumber="11">Laos</collectingCountry>
,
<collectingRegion id="498366D3C50E1403FF10F9A6FE291856" box="[202,447,1638,1662]" country="Laos" name="Khammouan" pageId="10" pageNumber="11">Khammouan Province</collectingRegion>
,
<collectingMunicipality id="6B9C324BC50E1403FE16F9A6FD151856" box="[460,643,1638,1662]" pageId="10" pageNumber="11">Tham Don Cave</collectingMunicipality>
, earthquake
<location id="8E98FEEAC50E1403FF5DF946FF5D18B5" LSID="urn:lsid:plazi:treatment:03EE1927C50C1402FC98F95DFE021A9D:8E98FEEAC50E1403FF5DF946FF5D18B5" box="[135,203,1670,1693]" country="Laos" county="South-Central" latitude="17.563583" longLatPrecision="1" longitude="104.871635" municipality="Tham Don Cave" name="Dome" pageId="10" pageNumber="11" stateProvince="Khammouan">Dome</location>
, sand sediments at bank of cave river,
<geoCoordinate id="EE73CEF6C50E1403FDADF944FC9418B5" box="[631,770,1668,1693]" degrees="17" direction="north" minutes="33.815" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="17.563583">17°33.815′N</geoCoordinate>
,
<geoCoordinate id="EE73CEF6C50E1403FF5DF964FE8B1895" box="[135,285,1700,1725]" degrees="104" direction="east" minutes="52.298" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="104.871635">104°52.298′E</geoCoordinate>
, alt.
<elevation id="006A4F02C50E1403FE8CF966FE4C1896" box="[342,474,1702,1726]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="11" unit="m" value="160.0">
<quantity id="4CBF05D4C50E1403FE8CF966FE081895" box="[342,414,1702,1726]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="10" pageNumber="11" unit="m" value="160.0">160 m</quantity>
a.s.l.
</elevation>
, locality code:
<specimenCode id="DBE1004AC50E1403FD53F966FD211896" box="[649,695,1702,1726]" collectionCode="JG" pageId="10" pageNumber="11">JG2</specimenCode>
B, coll.
<collectorName id="26B2CDE7C50E1403FF5DF906FF7018F6" box="[135,230,1734,1758]" pageId="10" pageNumber="11">J. Grego</collectorName>
,
<date id="FFF98EF1C50E1403FF29F906FE2E18F6" box="[243,440,1734,1758]" pageId="10" pageNumber="11" value="2017-02-11">
<collectingDate id="EFBD7719C50E1403FF29F906FE2E18F6" box="[243,440,1734,1758]" pageId="10" pageNumber="11" value="2017-02-11">11 February 2017</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC50E1403FF5DF8C6FD921956" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BF8A831C50E1403FF5DF8C6FD921956" blockId="10.[134,775,1798,1918]" pageId="10" pageNumber="11">
<emphasis id="B9337423C50E1403FF5DF8C6FF6D1936" bold="true" box="[135,251,1798,1822]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
A minute species with conical, strongly, regularly ribbed shell, closed umbilicus with a slight inner groove, weak apertural barriers (short parietal lamella, two palatal denticles, and columellar denticle).
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC50E1403FF5DF866FA8F1636" pageId="10" pageNumber="11" type="description">
<paragraph id="8BF8A831C50E1403FF5DF866FBCA1996" blockId="10.[135,775,1958,2078]" lastBlockId="10.[834,1476,1414,1982]" pageId="10" pageNumber="11">
<emphasis id="B9337423C50E1403FF5DF866FE821996" bold="true" box="[135,276,1958,1982]" pageId="10" pageNumber="11">Description.</emphasis>
Shell whitish, gradually growing to become conical to slender conical or pupiform. Body whorl widest from standard apertural view. Entire shell with 4.75–5 whorls. Due to the obscure boundary between protoconch and teleoconch, size of protoconch could not be determined. Protoconch not glossy, very finely pitted. Teleoconch strongly and regularly ribbed. Whorls strongly bulging, suture deep. Aperture suboval/deformed heart-shaped. Parietal callus rounded, reaches ca. half of preceding whorl, relatively thick, not “smeared” onto preceding whorl. Apertural dentition four or five-fold. Parietal lamella strong, elevated, relatively short. Outer portion slightly bent in palatal direction, inner part bent in columellar direction. Sinulus relatively wide, opens slightly laterally. Palatal denticles pointed, somewhat elongate and situated on a ridge, upper one situated closer to peristome. Columellar tooth moderately deeply situated, small, pointed, not elongated. An additional basal denticle might be present on lip. Peristome thickened, expanded, and slightly reflected. Basal swelling slight, situated in some distance from peristome. Impression anterior to basal swelling not visible. Umbilicus closed, not keeled, inner side with shallow groove.
</paragraph>
<paragraph id="8BF8A831C50E1403FC99F826FA8F1636" blockId="10.[835,1472,2022,2078]" pageId="10" pageNumber="11">
<emphasis id="B9337423C50E1403FC99F826FC6A19D6" bold="true" box="[835,1020,2022,2046]" pageId="10" pageNumber="11">Measurements.</emphasis>
H =
<quantity id="4CBF05D4C50E1403FB9BF826FB4919D5" box="[1089,1247,2022,2046]" metricMagnitude="-3" metricUnit="m" metricValue="1.77" metricValueMax="2.1" metricValueMin="1.44" pageId="10" pageNumber="11" unit="mm" value="1.77" valueMax="2.1" valueMin="1.44">1.44–2.1 mm</quantity>
, D =
<quantity id="4CBF05D4C50E1403FAF2F826FA5619D5" box="[1320,1472,2022,2046]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" metricValueMax="11.0" metricValueMin="8.299999999999999" pageId="10" pageNumber="11" unit="mm" value="0.965" valueMax="1.1" valueMin="0.83">0.83–1.1 mm</quantity>
(smallest and largest specimens measured).
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC50F1402FFABFF7DFF501F3D" pageId="11" pageNumber="12" type="etymology">
<paragraph id="8BF8A831C50F1402FFABFF7DFF501F3D" blockId="11.[113,752,189,277]" pageId="11" pageNumber="12">
<emphasis id="B9337423C50F1402FFABFF7DFF6C1EFD" bold="true" box="[113,250,189,213]" pageId="11" pageNumber="12">Etymology.</emphasis>
The specific epithet refers to the extreme variability of shell shape between localities of this new species.
</paragraph>
</subSubSection>
<subSubSection id="C35DFBBAC50F1402FFABFEFDFE021A9D" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BF8A831C50F1402FFABFEFDFEC21C7D" blockId="11.[112,752,317,597]" pageId="11" pageNumber="12">
<emphasis id="B9337423C50F1402FFABFEFDFF761F7D" bold="true" box="[113,224,317,341]" pageId="11" pageNumber="12">Remarks.</emphasis>
<emphasis id="B9337423C50F1402FF33FEFDFE551F7D" box="[233,451,317,341]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="4C47D3B2C50F1402FF33FEFDFECE1F7C" authorityName="Kobelt" authorityYear="1904" box="[233,344,317,340]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="genus">Sinoennea</taxonomicName>
atomaria
</emphasis>
(
<bibRefCitation id="EFD6D5C0C50F1402FE09FEFDFD321F7D" author="Dautzenberg P" box="[467,676,317,341]" pageId="11" pageNumber="12" pagination="157 - 165" refId="ref7731" refString="Dautzenberg P (1893) Mollusques nouveaux recueillis au Tonkin par M. le capitaine Em. Dorr et decrits. Journal de Conchyliologie, 41: 157 - 165." type="journal article" year="1893">Dautzenberg, 1893</bibRefCitation>
) is
<quantity id="4CBF05D4C50F1402FD15FEFDFF0C1F5C" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="11" pageNumber="12" unit="mm" value="2.5">2.5 mm</quantity>
high according to the original description, therefore a comparison with this new species should be given. However, the examined
<typeStatus id="54FC1693C50F1402FED3FE5DFEF01F9D" box="[265,358,413,437]" pageId="11" pageNumber="12" type="holotype">holotype</typeStatus>
(MNHN-IM-2000-30895) is a shell fragment of only the apical whorls. The drawing in the original description, however, shows a shell with a last whorl that is partly free from penultimate whorl, whereas in
<taxonomicName id="4C47D3B2C50F1402FF55FDDDFE871C1D" authority="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi, 2020" authorityName="Páll-Gergely &amp; Reischütz &amp; Maassen &amp; Grego &amp; Hunyadi" authorityYear="2020" box="[143,273,541,565]" class="Gastropoda" family="Streptaxidae" genus="Sinoennea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Stylommatophora" pageId="11" pageNumber="12" phylum="Mollusca" rank="species" species="variabilis" status="sp. nov.">
<emphasis id="B9337423C50F1402FF55FDDDFE871C1D" box="[143,273,541,565]" italics="true" pageId="11" pageNumber="12">S. variabilis</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A200C958C50F1402FEFBFDDDFE301C1D" box="[289,422,541,565]" pageId="11" pageNumber="12" rank="species">new species</taxonomicNameLabel>
, the body whorl is attached to the preceding whorl.
</paragraph>
<paragraph id="8BF8A831C50F1402FFABFDBDFE021A9D" blockId="11.[113,753,637,1205]" pageId="11" pageNumber="12">
This species is known from three sites. Two of them (3L07 and 4L07) are situated
<quantity id="4CBF05D4C50F1402FEB3FD5DFE251C9D" box="[361,435,669,693]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="11" pageNumber="12" unit="km" value="2.7">2.7 km</quantity>
from each other, whereas the third sample (JG2B) lies
<quantity id="4CBF05D4C50F1402FE5DFD7DFE5A1CFD" box="[391,460,701,725]" metricMagnitude="4" metricUnit="m" metricValue="3.1" pageId="11" pageNumber="12" unit="km" value="31.0">31 km</quantity>
and
<quantity id="4CBF05D4C50F1402FDDFFD7DFDDC1CFD" box="[517,586,701,725]" metricMagnitude="4" metricUnit="m" metricValue="3.4" pageId="11" pageNumber="12" unit="km" value="34.0">34 km</quantity>
from the other two sites, respectively. The
<typeStatus id="54FC1693C50F1402FE41FD1DFE5F1CDD" box="[411,457,733,757]" pageId="11" pageNumber="12">type</typeStatus>
series contained four shells that were very similar to each other. Sample from site 4L07 (
<figureCitation id="137CB4B4C50F1402FFA3FCDDFF581D1D" box="[121,206,797,821]" captionStart="Fig" captionStartId="9.[113,148,1915,1937]" captionTargetBox="[113,1451,338,1900]" captionTargetId="figure-11@9.[113,1451,338,1900]" captionTargetPageId="9" captionText="Fig. 5. Sinoennea variabilis Páll-Gergely &amp; Grego, new species. A, holotype (3L07); B, 4L07 (not paratype); C, JG2B (not paratype). Scale bar represents 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4577177" httpUri="https://zenodo.org/record/4577177/files/figure.png" pageId="11" pageNumber="12">Fig. 5B</figureCitation>
) contained a single shell. Sample from site JG2B contained two shells, of which one was slightly smaller than the other and had denser ribs, but their shell shapes were similar. While the shell shape shows conspicuous variability between populations, and rib density differed within and between populations as well, the apertural denticles usually used to distinguish species were indistinguishable in all examined shells. Since our sampling is rather insufficient due to low number of localities and specimens per site, we do not describe the local forms as distinct species, but interpret the variable shell shape as a character irrelevant for species recognition. More detailed sampling should clarify this question in the future.
</paragraph>
</subSubSection>
</treatment>
</document>
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