221 lines
31 KiB
XML
221 lines
31 KiB
XML
<document id="BD7B10434125B0042CEF24CCD77FA477" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF618319FF2E14C9FA901B10" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia minckleyi Hershler, Liu & Landye, 2011, new species" docType="treatment" docVersion="8" lastPageNumber="24" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="23" updateTime="1698261240372" updateUser="plazi">
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<mods:title id="3506AFB97FDC779FB8614CCC94876B45">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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<mods:namePart id="D1277AF2C83B0C302896F302B2A29027">Hershler, Robert</mods:namePart>
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<mods:roleTerm id="B7BBCCABD7B122D625F5633FF55A882B">Author</mods:roleTerm>
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<mods:namePart id="FBD7F44D145D57D09D0F90B6AC819B2C">Liu, Hsiu-Ping</mods:namePart>
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<mods:roleTerm id="7583C25B7BFE284FE8FA935209390884">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A62710B966E6D00455FF9F56AFABF468">Landye, Jerry</mods:namePart>
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<mods:typeOfResource id="9E58FA132DEABA10EB041AC40F913961">text</mods:typeOfResource>
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<mods:title id="8FB640D18CBBE66DA3FBB34908E350D1">Zootaxa</mods:title>
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<mods:part id="9FC64B27BDFFB8952B2911E93A3D3FBB">
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<mods:date id="F5C41CEC4376F76DB1A97046BAE17E44">2011</mods:date>
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<mods:identifier id="313DEB37F44CEAF20C0A55D9FD98194F" type="DOI">10.5281/zenodo.278442</mods:identifier>
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<mods:identifier id="C1D4880A67E95EA3A779EBF31DB9EBDE" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03E687C5BF618319FF2E14C9FA901B10" ID-DOI="http://doi.org/10.5281/zenodo.5629209" ID-GBIF-Taxon="119406543" ID-Zenodo-Dep="5629209" LSID="urn:lsid:plazi:treatment:03E687C5BF618319FF2E14C9FA901B10" httpUri="http://treatment.plazi.org/id/03E687C5BF618319FF2E14C9FA901B10" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
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<subSubSection id="C3556558BF618318FF2E14C9FB121D33" pageId="22" pageNumber="23" type="nomenclature">
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<paragraph id="8BF036D3BF618318FF2E14C9FE5D1AA3" blockId="22.[151,505,979,1039]" box="[151,505,979,1004]" pageId="22" pageNumber="23">
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<heading id="D0B881BFBF618318FF2E14C9FE5D1AA3" bold="true" box="[151,505,979,1004]" fontSize="10" level="2" pageId="22" pageNumber="23" reason="2">
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<emphasis id="B93BEAC1BF618318FF2E14C9FE5D1AA3" bold="true" box="[151,505,979,1004]" pageId="22" pageNumber="23">
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<taxonomicName id="4C4F4D50BF618318FF2E14C9FECC1AA3" box="[151,360,979,1004]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="minckleyi" status="sp. nov.">
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<emphasis id="B93BEAC1BF618318FF2E14C9FECC1AA3" bold="true" box="[151,360,979,1004]" italics="true" pageId="22" pageNumber="23">Tryonia minckleyi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF618318FED614CEFE5D1AA3" box="[367,505,979,1004]" pageId="22" pageNumber="23" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036D3BF618318FF2E14EDFE471D40" blockId="22.[151,505,979,1039]" box="[151,483,1015,1039]" pageId="22" pageNumber="23">
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(
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<figureCitation id="13742A56BF618318FF2614EDFF411D40" box="[159,229,1015,1039]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="22" pageNumber="23">Figs 5</figureCitation>
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K, 10D–E, H–I, M–
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<collectingCountry id="F3587643BF618318FE7E14EDFE7F1D40" box="[455,475,1015,1039]" name="Iceland" pageId="22" pageNumber="23">O</collectingCountry>
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)
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</paragraph>
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<paragraph id="8BF036D3BF618318FF2E135AFB121D33" blockId="22.[151,1436,1087,2012]" pageId="22" pageNumber="23">
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<emphasis id="B93BEAC1BF618318FF2E135AFF7E1D17" bold="true" box="[151,218,1088,1112]" pageId="22" pageNumber="23">
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<typeStatus id="54F48871BF618318FF2E135AFF7E1D17" box="[151,218,1088,1112]" pageId="22" pageNumber="23">Types</typeStatus>
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</emphasis>
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.
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<typeStatus id="54F48871BF618318FF531325FEF11D18" box="[234,341,1087,1111]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
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, USNM 1001751, Balneario de San Diego de Alcala, Chihuahua,
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<geoCoordinate id="EE7B5014BF618318FB831325FB7E1D18" box="[1082,1242,1087,1111]" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="28.588223">28º35'17.6" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF618318FB5F1325FA3C1D18" box="[1254,1432,1087,1111]" direction="west" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="-105.54745">105º32'50.8" W</geoCoordinate>
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, leg. R.H. and J.J.L.,
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<date id="FFF11013BF618318FEC4137EFE571D33" box="[381,499,1124,1148]" pageId="22" pageNumber="23" value="2001-12-04">4/xii/2001</date>
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.
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<typeStatus id="54F48871BF618318FE47137FFDCF1D32" box="[510,619,1125,1149]" pageId="22" pageNumber="23" type="paratype">Paratypes</typeStatus>
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(from same lot), USNM 1001755, USNM 1153673.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF618318FF7E1392FD1B1DA8" pageId="22" pageNumber="23" type="etymology">
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<paragraph id="8BF036D3BF618318FF7E1392FD1B1DA8" blockId="22.[151,1436,1087,2012]" pageId="22" pageNumber="23">
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<emphasis id="B93BEAC1BF618318FF7E1392FEE21DEF" bold="true" box="[199,326,1160,1184]" pageId="22" pageNumber="23">Etymology</emphasis>
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. A patronym honoring the late Wendell L. Minckley for his remarkable record of research on aquatic biota of the Chihuahuan Desert (
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<bibRefCitation id="EFDE4B22BF618318FDE213B6FC891D8B" author="Collins" box="[603,813,1196,1220]" pageId="22" pageNumber="23" refString="Collins, J. P., Deacon, J., Dowling, T. & Marsh, P. (2002) Wendell Lee Minckley 1935 - 2001. Copeia, 2002, 258 - 262." type="journal article" year="2002">
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Collins
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<emphasis id="B93BEAC1BF618318FD0D13B7FD411D8B" box="[692,741,1196,1220]" italics="true" pageId="22" pageNumber="23">et al</emphasis>
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. 2002
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</bibRefCitation>
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) and for leading the collecting expeditions that resulted in the discovery of this (and many other) species.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF618318FF7E13E9FE831C60" pageId="22" pageNumber="23" type="materials_examined">
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<paragraph id="8BF036D3BF618318FF7E13E9FE831C60" blockId="22.[151,1436,1087,2012]" pageId="22" pageNumber="23">
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<emphasis id="B93BEAC1BF618318FF7E13E9FE3B1C43" bold="true" box="[199,415,1267,1292]" pageId="22" pageNumber="23">Referred material</emphasis>
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. CHIHUAHUA.
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<collectionCode id="ED5EAE16BF618318FDDE13EEFD1C1C43" LSID="urn:lsid:biocol.org:col:34871" box="[615,696,1268,1292]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23">USNM</collectionCode>
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873200,
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<collectionCode id="ED5EAE16BF618318FC9A13EEFCD01C43" LSID="urn:lsid:biocol.org:col:34871" box="[803,884,1268,1292]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23">USNM</collectionCode>
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905268,
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<collectionCode id="ED5EAE16BF618318FC6613EEFB941C43" LSID="urn:lsid:biocol.org:col:34871" box="[991,1072,1268,1292]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="22" pageNumber="23">USNM</collectionCode>
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905269, topotypes, J.J.L.
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<emphasis id="B93BEAC1BF618318FAE513EFFA311C43" box="[1372,1429,1268,1292]" italics="true" pageId="22" pageNumber="23">et al.</emphasis>
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,
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<date id="FFF11013BF618318FF2E120DFE801C60" box="[151,292,1303,1327]" pageId="22" pageNumber="23" value="1971-08-29">29/viii/1971</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF618318FF7E1221FB101F48" pageId="22" pageNumber="23" type="diagnosis">
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<paragraph id="8BF036D3BF618318FF7E1221FB101F48" blockId="22.[151,1436,1087,2012]" pageId="22" pageNumber="23">
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<emphasis id="B93BEAC1BF618318FF7E1221FE9E1C1B" bold="true" box="[199,314,1339,1364]" pageId="22" pageNumber="23">Diagnosis</emphasis>
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. Shell small, ovate-conic or conic; penis having two distal papillae on the inner edge and one–two basal papillae on the outer edge. Contrasted with syntopic
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<taxonomicName id="4C4F4D50BF618318FCF7127BFC511C37" box="[846,1013,1376,1400]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="chuviscarae">
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<emphasis id="B93BEAC1BF618318FCF7127BFC511C37" box="[846,1013,1376,1400]" italics="true" pageId="22" pageNumber="23">T. chuviscarae</emphasis>
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</taxonomicName>
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above. Differs from
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<taxonomicName id="4C4F4D50BF618318FB4A127BFA381C37" box="[1267,1436,1376,1400]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="julimesensis">
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<emphasis id="B93BEAC1BF618318FB4A127BFA381C37" box="[1267,1436,1376,1400]" italics="true" pageId="22" pageNumber="23">T. julimesensis</emphasis>
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</taxonomicName>
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(described next), which is also distributed in the Río Conchos basin, in having basal penial papillae. Readily distinguished from conchologically similar
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<taxonomicName id="4C4F4D50BF618318FDFC12B3FD661C8F" box="[581,706,1448,1472]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="22" pageNumber="23" phylum="Mollusca" rank="species" species="allendae">
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<emphasis id="B93BEAC1BF618318FDFC12B3FD661C8F" box="[581,706,1448,1472]" italics="true" pageId="22" pageNumber="23">T. allendae</emphasis>
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</taxonomicName>
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(described below), which is also distributed in the Río Conchos basin, by its more rounded shell whorls, more elongate cusps on the lateral radular teeth, larger penial stylet, weakly undulating penial duct, and smaller lateral expansion of the terminal bulb of the penis.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF618319FF7E1109FD5418BB" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23" type="description">
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<paragraph id="8BF036D3BF618318FF7E1109FDD71FF3" blockId="22.[151,1436,1087,2012]" pageId="22" pageNumber="23">
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<emphasis id="B93BEAC1BF618318FF7E1109FEF51F63" bold="true" box="[199,337,1555,1580]" pageId="22" pageNumber="23">Description</emphasis>
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. Shell (
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<figureCitation id="13742A56BF618318FE0B110EFDA31F63" box="[434,519,1556,1580]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
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D–E) up to
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<quantity id="4CB79B36BF618318FD22110EFD521F63" box="[667,758,1556,1580]" metricMagnitude="-3" metricUnit="m" metricValue="2.8" pageId="22" pageNumber="23" unit="mm" value="2.8">2.8 mm</quantity>
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tall, large females having 4.75–5.50 whorls, spire height 120–140% width of shell, male shells smaller than those of females. Teleoconch whorls high convex, shouldered, sutures impressed. Aperture rounded adapically, parietal lip complete, usually adnate, rarely slightly disjunct, umbilicus narrow or absent. Outer lip thin, orthocline. Sculpture of strong growth lines, weak spiral thread also sometimes present. Periostracum light tan.
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</paragraph>
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<paragraph id="8BF036D3BF618318FF7E11DDFA881E68" blockId="22.[151,1436,1087,2012]" pageId="22" pageNumber="23">
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<emphasis id="B93BEAC1BF618318FF7E11DDFE6D1FAF" bold="true" box="[199,457,1735,1760]" pageId="22" pageNumber="23">Shell measurements (</emphasis>
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||
mean in parentheses): height
|
||
<quantity id="4CB79B36BF618318FCA011DDFC1A1F90" box="[793,958,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="2.475" metricValueMax="2.7" metricValueMin="2.25" pageId="22" pageNumber="23" unit="mm" value="2.475" valueMax="2.7" valueMin="2.25">2.25–2.70 mm</quantity>
|
||
(2.48), width
|
||
<quantity id="4CB79B36BF618318FBDA11DDFAAC1F90" box="[1123,1288,1735,1759]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" metricValueMax="1.31" metricValueMin="1.17" pageId="22" pageNumber="23" unit="mm" value="1.24" valueMax="1.31" valueMin="1.17">1.17–1.31 mm</quantity>
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||
(1.24), body whorl height
|
||
<quantity id="4CB79B36BF618318FE9411F6FE741E4B" box="[301,464,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="1.53" metricValueMax="1.62" metricValueMin="1.44" pageId="22" pageNumber="23" unit="mm" value="1.53" valueMax="1.62" valueMin="1.44">1.44–1.62 mm</quantity>
|
||
(1.53), body whorl width
|
||
<quantity id="4CB79B36BF618318FD4011F6FC391E4B" box="[761,925,1772,1796]" metricMagnitude="-3" metricUnit="m" metricValue="1.15" metricValueMax="1.24" metricValueMin="1.06" pageId="22" pageNumber="23" unit="mm" value="1.15" valueMax="1.24" valueMin="1.06">1.06–1.24 mm</quantity>
|
||
(1.16), aperture height
|
||
<quantity id="4CB79B36BF618318FB1E11F6FAEE1E4B" box="[1191,1354,1772,1796]" metricMagnitude="-4" metricUnit="m" metricValue="8.8" metricValueMax="9.5" metricValueMin="8.1" pageId="22" pageNumber="23" unit="mm" value="0.88" valueMax="0.95" valueMin="0.81">0.81–0.95 mm</quantity>
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||
(0.87), aperture width
|
||
<quantity id="4CB79B36BF618318FEF91015FE471E68" box="[320,483,1807,1831]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" metricValueMax="7.8" metricValueMin="7.0" pageId="22" pageNumber="23" unit="mm" value="0.74" valueMax="0.78" valueMin="0.7">0.70–0.78 mm</quantity>
|
||
(0.74), total number of whorls 4.75–5.25 (4.93) (USNM 1153673,
|
||
<emphasis id="B93BEAC1BF618318FB6D100BFB461E67" box="[1236,1250,1809,1832]" italics="true" pageId="22" pageNumber="23">n</emphasis>
|
||
= 15).
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF618318FF7E102EFC671E20" blockId="22.[151,1436,1087,2012]" pageId="22" pageNumber="23">
|
||
<emphasis id="B93BEAC1BF618318FF7E102EFDA71E03" bold="true" box="[199,515,1843,1868]" pageId="22" pageNumber="23">
|
||
Measurements of
|
||
<typeStatus id="54F48871BF618318FE25102EFDA71E03" box="[412,515,1844,1868]" pageId="22" pageNumber="23" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
: height
|
||
<quantity id="4CB79B36BF618318FDD9102EFD631E03" box="[608,711,1844,1868]" metricMagnitude="-3" metricUnit="m" metricValue="2.69" pageId="22" pageNumber="23" unit="mm" value="2.69">2.69 mm</quantity>
|
||
, width
|
||
<quantity id="4CB79B36BF618318FCA0102EFC241E03" box="[793,896,1844,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.28" pageId="22" pageNumber="23" unit="mm" value="1.28">1.28 mm</quantity>
|
||
, body whorl height
|
||
<quantity id="4CB79B36BF618318FBD8102EFB6D1E03" box="[1121,1225,1844,1868]" metricMagnitude="-3" metricUnit="m" metricValue="1.62" pageId="22" pageNumber="23" unit="mm" value="1.62">1.62 mm</quantity>
|
||
, body whorl width
|
||
<quantity id="4CB79B36BF618318FF2E104DFF591E20" box="[151,253,1879,1903]" metricMagnitude="-3" metricUnit="m" metricValue="1.19" pageId="22" pageNumber="23" unit="mm" value="1.19">1.19 mm</quantity>
|
||
, aperture height
|
||
<quantity id="4CB79B36BF618318FE01104DFDBA1E20" box="[440,542,1879,1903]" metricMagnitude="-4" metricUnit="m" metricValue="9.399999999999999" pageId="22" pageNumber="23" unit="mm" value="0.94">0.94 mm</quantity>
|
||
, aperture width
|
||
<quantity id="4CB79B36BF618318FD6A104DFC9D1E20" box="[723,825,1879,1903]" metricMagnitude="-4" metricUnit="m" metricValue="7.4" pageId="22" pageNumber="23" unit="mm" value="0.74">0.74 mm</quantity>
|
||
, 5.0 whorls.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF618319FF7E1067FDEE19B8" blockId="22.[151,1436,1087,2012]" lastBlockId="23.[151,1436,151,608]" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
|
||
Inner and outer sides of operculum smooth (
|
||
<figureCitation id="13742A56BF618318FD051066FCA91EDB" box="[700,781,1916,1940]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
H–I). Radula (
|
||
<figureCitation id="13742A56BF618318FC091066FBA61EDB" box="[944,1026,1916,1940]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
M-O): dorsal edge of central radular teeth weakly concave, basal tongue rounded, median cusps distally pointed, parallel-sided proximally, lateral cusps four–five, basal cusps two (equal-sized or innermost larger), small (
|
||
<figureCitation id="13742A56BF618318FC3110DEFC7C1E93" box="[904,984,1988,2012]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="22" pageNumber="23">Fig. 10</figureCitation>
|
||
N). Lateral teeth having two–three cusps on inner and four–five cusps on outer sides, length of outer wing about 180% width of cutting edge, central cusp pointed, curved (
|
||
<figureCitation id="13742A56BF608319FEE317A6FE0B199B" box="[346,431,188,212]" captionStart="FIGURE 10" captionStartId="21.[151,244,1801,1822]" captionTargetBox="[151,1436,151,1774]" captionTargetId="figure@21.[151,1436,151,1775]" captionTargetPageId="21" captionText="FIGURE 10. Tryonia chuviscarae sp. nov. and Tryonia minckleyi sp. nov. A. Holotype, T. chuviscarae, USNM 873270. B. Male shell, T. chuviscarae, USNM 1153669. C. Male shell, T. chuviscarae, USNM 1001750. D. Holotype, T. minckleyi, USNM 1001751. E. Male shell, T. minckleyi, USNM 1153673. F – G. Operculum (outer, inner sides), T. chuviscarae, USNM 1001750. H – I. Operculum (outer, inner sides), T. minckleyi, USNM 1153673. J. Portion of radular ribbon, T. chuviscarae, USNM 0 0 1750. K. Central radular teeth, T. chuviscarae, USNM 1001750. L. Lateral and inner marginal teeth, T. chuviscarae, USNM 1001750. M. Portion of radular ribbon, T. minckleyi, USNM 1153673. N. Central radular teeth, T. minckleyi, USNM 1153673. O. Lateral tooth, T. minckleyi, USNM 1153673. Scale bars A – E = 1.0 mm, F – I = 100 µm, J – O = 10 µm." httpUri="https://zenodo.org/record/278452/files/figure.png" pageId="23" pageNumber="24">Fig. 10</figureCitation>
|
||
<collectingCountry id="F3587643BF608319FE1717A6FE60199B" box="[430,452,188,212]" name="Iceland" pageId="23" pageNumber="24">O</collectingCountry>
|
||
). Inner marginal teeth having 19–30 cusps, outer marginal teeth having 25–31 cusps. Radula data are from USNM 1153673.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF608319FF7E161EFD5418BB" blockId="23.[151,1436,151,608]" pageId="23" pageNumber="24">
|
||
Animal darkly pigmented. Penis (
|
||
<figureCitation id="13742A56BF608319FDF1161EFD2F1853" box="[584,651,260,284]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="23" pageNumber="24">Fig. 5</figureCitation>
|
||
K) usually having two distal papillae on the inner edge and one–two basal papillae on the outer edge (19 of
|
||
<specimenCount id="9D49FD5ABF608319FDE8163DFD4A1870" box="[593,750,295,319]" pageId="23" pageNumber="24" type="generic">30 specimens</specimenCount>
|
||
). Two specimens differed in having a single distal papilla on the inner edge, one specimen differed in having three distal papillae on the inner edge, two specimens lacked a papilla on the outer edge, two specimens had three basal papillae on the outer edge, one specimen had five basal papillae on the outer edge, and one specimen had two distal and four basal papillae on the outer edge. Distal bulb of penis weakly expanded laterally on inner side, black; stylet large. Penial duct straight or weakly undulating along most of length. Penial data are from USNM 1153673.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF608319FF7E16E5FE541B73" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="8BF036D3BF608319FF7E16E5FE541B73" blockId="23.[151,1436,151,608]" pageId="23" pageNumber="24">
|
||
<emphasis id="B93BEAC1BF608319FF7E16E5FE4C1B57" bold="true" box="[199,488,511,536]" pageId="23" pageNumber="24">Distribution and habitat</emphasis>
|
||
. Endemic to the
|
||
<typeStatus id="54F48871BF608319FD1C151AFD711B57" box="[677,725,512,536]" pageId="23" pageNumber="24">type</typeStatus>
|
||
locality in the Río Conchos basin (
|
||
<figureCitation id="13742A56BF608319FBE416E5FB3B1B58" box="[1117,1183,511,535]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="23" pageNumber="24">Fig. 1</figureCitation>
|
||
, locality 9). Habitat as for
|
||
<taxonomicName id="4C4F4D50BF608319FF07153FFEFB1B73" box="[190,351,548,572]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="chuviscarae">
|
||
<emphasis id="B93BEAC1BF608319FF07153FFEFB1B73" box="[190,351,548,572]" italics="true" pageId="23" pageNumber="24">T. chuviscarae</emphasis>
|
||
</taxonomicName>
|
||
(see above).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF608319FF7E1552FA901B10" box="[199,1332,583,608]" pageId="23" pageNumber="24" type="discussion">
|
||
<paragraph id="8BF036D3BF608319FF7E1552FA901B10" blockId="23.[151,1436,151,608]" box="[199,1332,583,608]" pageId="23" pageNumber="24">
|
||
<emphasis id="B93BEAC1BF608319FF7E1552FE901B2F" bold="true" box="[199,308,584,608]" pageId="23" pageNumber="24">Remarks</emphasis>
|
||
.
|
||
<taxonomicName id="4C4F4D50BF608319FEFB1553FDAF1B2F" box="[322,523,584,608]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="minckleyi">
|
||
<emphasis id="B93BEAC1BF608319FEFB1553FDAF1B2F" box="[322,523,584,608]" italics="true" pageId="23" pageNumber="24">Tryonia minckleyi</emphasis>
|
||
</taxonomicName>
|
||
was delineated as a well supported lineage in the Bayesian tree (
|
||
<figureCitation id="13742A56BF608319FB5B155DFA831B10" box="[1250,1319,583,607]" captionStart="FIGURE 2" captionStartId="5.[151,244,1938,1959]" captionTargetBox="[244,1345,828,1915]" captionTargetId="figure@5.[244,1346,828,1920]" captionTargetPageId="5" captionText="FIGURE 2. Bayesian tree based on the COI dataset. Posterior probabilities are provided when> 95 %. The names of the species treated herein are in bold face. Terminals and specimen codes are labeled as in Table 1. The two well supported clades composed of new species described herein are labeled A – B." httpUri="https://zenodo.org/record/278444/files/figure.png" pageId="23" pageNumber="24">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |