196 lines
29 KiB
XML
196 lines
29 KiB
XML
<document id="E13281F7A98431562134CFC4FABD644A" ID-DOI="10.5281/zenodo.278442" ID-GBIF-Dataset="6b9505e0-ac6e-4baa-9066-6f729d38e9e2" ID-ISSN="1175-5326" ID-Zenodo-Dep="278442" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460341493798" checkinUser="plazi" docAuthor="Hershler, Robert, Liu, Hsiu-Ping & Landye, Jerry" docDate="2011" docId="03E687C5BF608319FF2E15AEFE981E95" docLanguage="en" docName="zt03001p032.pdf" docOrigin="Zootaxa 3001" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Tryonia julimesensis Hershler, Liu & Landye, 2011, new species" docType="treatment" docVersion="8" lastPageNumber="24" masterDocId="FFDFFFBDBF77830EFFB9171AFFA4194F" masterDocTitle="New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot" masterLastPageNumber="32" masterPageNumber="1" pageNumber="24" updateTime="1698261240372" updateUser="plazi">
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<mods:mods id="288EE83F6009EF71E57CA07ED070C74A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A68B77E42540993B82EE16DBBF9A8B2A">
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<mods:title id="5E82DD79A9519CD146BF5A4DD9E826BD">New species and records of springsnails (Caenogastropoda: Cochliopidae: Tryonia) from the Chihuahuan Desert (Mexico and United States), an imperiled biodiversity hotspot</mods:title>
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</mods:titleInfo>
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<mods:name id="2C2C494D9A4AA9D7756529B2FD87218F" type="personal">
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<mods:roleTerm id="9AD88EF7B0671335C381A33D31DD887B">Author</mods:roleTerm>
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<mods:namePart id="9E21BC6A228E526417B8BF4106FA2B7E">Hershler, Robert</mods:namePart>
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</mods:name>
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<mods:name id="4AC6DF7B68B3D5F114A64AB41F0F5E6D" type="personal">
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<mods:roleTerm id="174B9C862B1236417515998984584871">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="849D6AA74718059F7F7E54DAEA362F58">Liu, Hsiu-Ping</mods:namePart>
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</mods:name>
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<mods:name id="3261E223CCA5A18C29108E891D81C5B2" type="personal">
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<mods:roleTerm id="278114C647A35A59D093C802776F4BB0">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="01C24D582477B7BC69BB35A1FB750578">Landye, Jerry</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8AD0471669AD4BAC289EADE62E551C1E">text</mods:typeOfResource>
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<mods:title id="CFBE102B1A02FE3FD881D85D67075A0F">Zootaxa</mods:title>
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<mods:part id="1166ADA435A61E30EE390403B1A27E90">
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<mods:date id="75CE37B4F48D5BE013B180967E2C5D1B">2011</mods:date>
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<mods:detail id="A19C033C5B36096F94A303FE5ED2D088" type="volume">
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<mods:number id="477BEBA275C50E9F6442298B4F399614">3001</mods:number>
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<mods:classification id="69F77C68CE7042E540CA45CA9AE1CC07">journal article</mods:classification>
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<mods:identifier id="8069F4C28B68E691F432312586DED89B" type="DOI">10.5281/zenodo.278442</mods:identifier>
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<mods:identifier id="9415EBE403624BDF600F2B6FC6D42BB8" type="GBIF-Dataset">6b9505e0-ac6e-4baa-9066-6f729d38e9e2</mods:identifier>
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<mods:identifier id="D45CFC2AED4EF2282EDA4EC17D6A2FEF" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9EA2ABA29A3D974A473C554FD0E82DB0" type="Zenodo-Dep">278442</mods:identifier>
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<treatment id="03E687C5BF608319FF2E15AEFE981E95" ID-DOI="http://doi.org/10.5281/zenodo.5629211" ID-GBIF-Taxon="119406544" ID-Zenodo-Dep="5629211" LSID="urn:lsid:plazi:treatment:03E687C5BF608319FF2E15AEFE981E95" httpUri="http://treatment.plazi.org/id/03E687C5BF608319FF2E15AEFE981E95" lastPageNumber="24" pageId="23" pageNumber="24">
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<subSubSection id="C3556558BF608319FF2E15AEFF5B1A31" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BF036D3BF608319FF2E15AEFD8C1B82" blockId="23.[151,552,691,750]" box="[151,552,691,718]" pageId="23" pageNumber="24">
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<heading id="D0B881BFBF608319FF2E15AEFD8C1B82" bold="true" box="[151,552,691,718]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B93BEAC1BF608319FF2E15AEFD8C1B82" bold="true" box="[151,552,691,718]" pageId="23" pageNumber="24">
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<taxonomicName id="4C4F4D50BF608319FF2E15AEFE351B81" box="[151,401,692,718]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="julimesensis" status="sp. nov.">
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<emphasis id="B93BEAC1BF608319FF2E15AEFE351B81" bold="true" box="[151,401,692,718]" italics="true" pageId="23" pageNumber="24">Tryonia julimesensis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857BABF608319FE2015A9FD8C1B82" box="[409,552,691,717]" pageId="23" pageNumber="24" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF036D3BF608319FF2E15CCFE761BA1" blockId="23.[151,552,691,750]" box="[151,466,726,750]" pageId="23" pageNumber="24">
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(
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<figureCitation id="13742A56BF608319FF2615CCFF411BA1" box="[159,229,726,750]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="23" pageNumber="24">Figs 5</figureCitation>
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L, 11A–C, F–G, J–L)
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</paragraph>
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<paragraph id="8BF036D3BF608319FF2E1405FF5B1A31" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
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<emphasis id="B93BEAC1BF608319FF2E1405FF7E1A78" bold="true" box="[151,218,799,823]" pageId="23" pageNumber="24">
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<typeStatus id="54F48871BF608319FF2E1405FF7E1A78" box="[151,218,799,823]" pageId="23" pageNumber="24">Types</typeStatus>
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</emphasis>
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.
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<typeStatus id="54F48871BF608319FF541404FEFC1A79" box="[237,344,798,822]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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, USNM 874130, unnamed springs along east side of Río Conchos, south-southeast of Julimes, Chihuahua,
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<geoCoordinate id="EE7B5014BF608319FE9C1458FE6F1A15" box="[293,459,834,858]" direction="north" orientation="latitude" pageId="23" pageNumber="24" precision="1" value="28.410334">28º24'37.2" N</geoCoordinate>
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,
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<geoCoordinate id="EE7B5014BF608319FE641458FD001A15" box="[477,676,834,858]" direction="west" orientation="longitude" pageId="23" pageNumber="24" precision="1" value="-105.42266">105º25'21.6 " W</geoCoordinate>
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, leg. J.J.L.
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<emphasis id="B93BEAC1BF608319FC8C145EFCC91A14" box="[821,877,835,859]" italics="true" pageId="23" pageNumber="24">et al</emphasis>
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.,
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<date id="FFF11013BF608319FC311458FC541A15" box="[904,1008,834,858]" pageId="23" pageNumber="24" value="1991-05-05">5/v/1991</date>
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.
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<typeStatus id="54F48871BF608319FBB91459FBC91A14" box="[1024,1133,835,859]" pageId="23" pageNumber="24" type="paratype">Paratypes</typeStatus>
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(from same lot), USNM 1153671.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF608319FF7E1491FF631A89" pageId="23" pageNumber="24" type="etymology">
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<paragraph id="8BF036D3BF608319FF7E1491FF631A89" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
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<emphasis id="B93BEAC1BF608319FF7E1491FEE21AEC" bold="true" box="[199,326,907,931]" pageId="23" pageNumber="24">Etymology</emphasis>
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. A geographic epithet referring to the distribution of this species near the town of Julimes, Chihuahua.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF608319FF7E14C8FB181AA5" box="[199,1212,978,1003]" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8BF036D3BF608319FF7E14C8FB181AA5" blockId="23.[151,1436,798,2010]" box="[199,1212,978,1003]" pageId="23" pageNumber="24">
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<emphasis id="B93BEAC1BF608319FF7E14C8FE3A1AA4" bold="true" box="[199,414,978,1003]" pageId="23" pageNumber="24">Referred material</emphasis>
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. CHIHUAHUA.
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<collectionCode id="ED5EAE16BF608319FDDA14C8FD101AA5" LSID="urn:lsid:biocol.org:col:34871" box="[611,692,978,1002]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="23" pageNumber="24">USNM</collectionCode>
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854940, topotype, R.H. and J.J.L.,
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<date id="FFF11013BF608319FBFB14C8FB1C1AA5" box="[1090,1208,978,1002]" pageId="23" pageNumber="24" value="1998-12-09">9/xii/1998</date>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF608319FF7E14ECFD251D7D" pageId="23" pageNumber="24" type="diagnosis">
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<paragraph id="8BF036D3BF608319FF7E14ECFD251D7D" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
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<emphasis id="B93BEAC1BF608319FF7E14ECFE9E1D40" bold="true" box="[199,314,1014,1039]" pageId="23" pageNumber="24">Diagnosis</emphasis>
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. Shell small, broad- or ovate-conic; penis having two distal papillae on the inner edge. Contrasted with
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<taxonomicName id="4C4F4D50BF608319FF761306FED61D7C" box="[207,370,1051,1075]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="chuviscarae">
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<emphasis id="B93BEAC1BF608319FF761306FED61D7C" box="[207,370,1051,1075]" italics="true" pageId="23" pageNumber="24">T. chuviscarae</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C4F4D50BF608319FE101306FD8A1D7C" box="[425,558,1051,1075]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="minckleyi">
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<emphasis id="B93BEAC1BF608319FE101306FD8A1D7C" box="[425,558,1051,1075]" italics="true" pageId="23" pageNumber="24">T. minckleyi</emphasis>
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</taxonomicName>
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above.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556558BF608319FF7E1324FA991E4D" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BF036D3BF608319FF7E1324FE621C45" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
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<emphasis id="B93BEAC1BF608319FF7E1324FEF51D18" bold="true" box="[199,337,1086,1111]" pageId="23" pageNumber="24">Description</emphasis>
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. Shell (
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<figureCitation id="13742A56BF608319FE0B1324FDA11D19" box="[434,517,1086,1110]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
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A–C) up to 2.0 mm tall, large females having 4.00–4.50 whorls, spire height 80–99% width of shell, male shells smaller than those of females. Teleoconch whorls high convex, frequently shouldered, sutures impressed, last 0.25 whorl sometimes loosened. Aperture usually rounded adapically, rarely weakly angled, inner lip complete, slightly thickened, adnate or disjunct, umbilicus narrow or open. Outer lip thin, orthocline or prosocline, weakly sinuate. Sculpture of strong growth lines, weak spiral thread also sometimes present. Periostracum light tan.
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</paragraph>
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<paragraph id="8BF036D3BF608319FF7E120CFA881C39" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
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<emphasis id="B93BEAC1BF608319FF7E120CFE6D1C60" bold="true" box="[199,457,1302,1327]" pageId="23" pageNumber="24">Shell measurements (</emphasis>
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mean in parentheses): height
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<quantity id="4CB79B36BF608319FCA0120CFC1A1C61" box="[793,958,1302,1326]" metricMagnitude="-3" metricUnit="m" metricValue="1.76" metricValueMax="1.98" metricValueMin="1.54" pageId="23" pageNumber="24" unit="mm" value="1.76" valueMax="1.98" valueMin="1.54">1.54–1.98 mm</quantity>
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(1.69), width
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<quantity id="4CB79B36BF608319FBDA120CFAAC1C61" box="[1123,1288,1302,1326]" metricMagnitude="-3" metricUnit="m" metricValue="1.12" metricValueMax="1.21" metricValueMin="1.03" pageId="23" pageNumber="24" unit="mm" value="1.12" valueMax="1.21" valueMin="1.03">1.03–1.21 mm</quantity>
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(1.09), body whorl height
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<quantity id="4CB79B36BF608319FE941220FE741C1D" box="[301,464,1338,1362]" metricMagnitude="-3" metricUnit="m" metricValue="1.245" metricValueMax="1.34" metricValueMin="1.15" pageId="23" pageNumber="24" unit="mm" value="1.245" valueMax="1.34" valueMin="1.15">1.15–1.34 mm</quantity>
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(1.21), body whorl width
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<quantity id="4CB79B36BF608319FD401220FC391C1D" box="[761,925,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="9.950000000000001" metricValueMax="10.700000000000001" metricValueMin="9.200000000000001" pageId="23" pageNumber="24" unit="mm" value="0.995" valueMax="1.07" valueMin="0.92">0.92–1.07 mm</quantity>
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(0.97), aperture height
|
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<quantity id="4CB79B36BF608319FB1E1220FAEE1C1D" box="[1191,1354,1338,1362]" metricMagnitude="-4" metricUnit="m" metricValue="7.25" metricValueMax="7.9" metricValueMin="6.6" pageId="23" pageNumber="24" unit="mm" value="0.725" valueMax="0.79" valueMin="0.66">0.66–0.79 mm</quantity>
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(0.71), aperture width
|
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<quantity id="4CB79B36BF608319FEF91244FE471C39" box="[320,483,1374,1398]" metricMagnitude="-4" metricUnit="m" metricValue="6.6" metricValueMax="7.1" metricValueMin="6.1" pageId="23" pageNumber="24" unit="mm" value="0.6599999999999999" valueMax="0.71" valueMin="0.61">0.61–0.71 mm</quantity>
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(0.65), total number of whorls 4.00–4.50 (4.18) (USNM 1153671,
|
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<emphasis id="B93BEAC1BF608319FB6D127AFB461C38" box="[1236,1250,1376,1399]" italics="true" pageId="23" pageNumber="24">n</emphasis>
|
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= 15).
|
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</paragraph>
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<paragraph id="8BF036D3BF608319FF7E1299FC751CF1" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
|
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<emphasis id="B93BEAC1BF608319FF7E1299FDA71CD4" bold="true" box="[199,515,1410,1435]" pageId="23" pageNumber="24">
|
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Measurements of
|
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<typeStatus id="54F48871BF608319FE251299FDA71CD4" box="[412,515,1411,1435]" pageId="23" pageNumber="24" type="holotype">holotype</typeStatus>
|
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</emphasis>
|
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: height
|
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<quantity id="4CB79B36BF608319FDD91298FD631CD5" box="[608,711,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="23" pageNumber="24" unit="mm" value="1.9">1.90 mm</quantity>
|
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, width
|
||
<quantity id="4CB79B36BF608319FCA01298FC241CD5" box="[793,896,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="1.24" pageId="23" pageNumber="24" unit="mm" value="1.24">1.24 mm</quantity>
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||
, body whorl height
|
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<quantity id="4CB79B36BF608319FBD81298FB6D1CD5" box="[1121,1225,1410,1434]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" pageId="23" pageNumber="24" unit="mm" value="1.34">1.34 mm</quantity>
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||
, body whorl width
|
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<quantity id="4CB79B36BF608319FF2E12BCFF591CF1" box="[151,253,1446,1470]" metricMagnitude="-3" metricUnit="m" metricValue="1.07" pageId="23" pageNumber="24" unit="mm" value="1.07">1.07 mm</quantity>
|
||
, aperture height
|
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<quantity id="4CB79B36BF608319FE0112BCFDBA1CF1" box="[440,542,1446,1470]" metricMagnitude="-4" metricUnit="m" metricValue="8.1" pageId="23" pageNumber="24" unit="mm" value="0.81">0.81 mm</quantity>
|
||
, aperture width
|
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<quantity id="4CB79B36BF608319FD6A12BCFC9D1CF1" box="[723,825,1446,1470]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="23" pageNumber="24" unit="mm" value="0.69">0.69 mm</quantity>
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, 4.25 whorls.
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</paragraph>
|
||
<paragraph id="8BF036D3BF608319FF7E12D1FD321FD9" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
|
||
Inner and outer sides of operculum smooth (
|
||
<figureCitation id="13742A56BF608319FD7B12D0FCB51CAD" box="[706,785,1482,1506]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
|
||
F–G). Radula (
|
||
<figureCitation id="13742A56BF608319FC0412D0FBA91CAD" box="[957,1037,1482,1506]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
|
||
J-L): dorsal edge of central radular teeth strongly concave, basal tongue rounded or V-shaped, median cusps distally pointed, parallel-sided proximally, lateral cusps four–five, basal cusps one–three (roughly equal-sized) (
|
||
<figureCitation id="13742A56BF608319FB9C1108FBDF1F65" box="[1061,1147,1554,1578]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
|
||
K). Lateral teeth having three–four cusps on inner and four–five cusps on outer sides, length of outer wing about 160% width of cutting edge, central cusp pointed (
|
||
<figureCitation id="13742A56BF608319FE721140FDBE1F3D" box="[459,538,1626,1650]" captionStart="FIGURE 11" captionStartId="24.[151,244,1770,1791]" captionTargetBox="[151,1436,151,1743]" captionTargetId="figure@24.[151,1436,151,1743]" captionTargetPageId="24" captionText="FIGURE 11. Tryonia julimesensis sp. nov. and Tryonia allendae sp. nov. A. Holotype, T. julimesensis, USNM 874130. B – C. Male shells, T. julimesensis, USNM 1153671. D. Holotype, T. allendae, USNM 892115. E. Shell, T. allendae, USNM 1153667. F – G. Operculum (outer, inner sides), T. julimesensis, USNM 1153671. H – I. Operculum (outer, inner sides), T. allendae, USNM 1153671. J. Portion of radular ribbon, T. julimesensis, USNM 1153671. K. Central radular teeth, T. julimesensis, USNM 1153671. L. Lateral tooth, T. julimesensis, USNM 1153671. M. Portion of radular ribbon, T. allendae, USNM 1153667. N. Central radular teeth, T. allendae, USNM 1153667. R. Lateral teeth, T. allendae, USNM 1153667. Scale bars A – E = 1.0 mm; F – I = 250 µm; J, K – L, N – O = 10 µm; M = 25 µm." httpUri="https://zenodo.org/record/278453/files/figure.png" pageId="23" pageNumber="24">Fig. 11</figureCitation>
|
||
L). Inner marginal teeth having 16–21 cusps, outer marginal teeth having 15–20 cusps. Radula data are from USNM 1153671.
|
||
</paragraph>
|
||
<paragraph id="8BF036D3BF608319FF7E11B8FA991E4D" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
|
||
Animal darkly pigmented. Penis (
|
||
<figureCitation id="13742A56BF608319FDFF11B8FD2C1FF5" box="[582,648,1698,1722]" captionStart="FIGURE 5" captionStartId="10.[151,244,1857,1878]" captionTargetBox="[241,1348,158,1832]" captionTargetId="figure@10.[229,1363,145,1846]" captionTargetPageId="10" captionText="FIGURE 5. Penes. A. T. peregrina sp. nov., USNM 1153677. B. T. zaragozae sp. nov., USNM 1153679. C. T. ovata sp. nov., USNM 1153676. D. T. angosturae sp. nov., USNM 874174. E. T. molinae sp. nov., USNM 1153674. F. T. contrerasi sp. nov., USNM 1153670. G. T. taylori sp. nov., USNM 874047. H. T. taylori sp. nov., USNM 873252. I. T. metcalfi sp. nov., USNM 1153672. J. T. chuviscarae sp. nov., USNM 1001756. K. T. minckleyi sp. nov., USNM 1153673. L. T. julimesensis sp. nov., USNM 1153671. M. T. allendae sp. nov., USNM 1153667. N. T. oasiensis sp. nov., USNM 1153675. O. T. seemani, USNM 874125. P. T. seemani, USNM 874123. Scale bars = 250 µm. Db = distal bulb, Pa = papilla, Pd = penial duct, St = stylet." httpUri="https://zenodo.org/record/278447/files/figure.png" pageId="23" pageNumber="24">Fig. 5</figureCitation>
|
||
L) having two distal papillae on the inner edge (26 of
|
||
<specimenCount id="9D49FD5ABF608319FB4B11B8FA341FF5" box="[1266,1424,1698,1722]" pageId="23" pageNumber="24" type="generic">30 specimens</specimenCount>
|
||
), four specimens differed in having a single distal papilla. Distal bulb of penis expanded laterally on inner side, black; stylet large. Penial duct weakly undulating in medial section. Penial data are from USNM 1153671.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556558BF608319FF7E1014FE981E95" pageId="23" pageNumber="24" type="distribution">
|
||
<paragraph id="8BF036D3BF608319FF7E1014FE981E95" blockId="23.[151,1436,798,2010]" pageId="23" pageNumber="24">
|
||
<emphasis id="B93BEAC1BF608319FF7E1014FE4E1E68" bold="true" box="[199,490,1806,1831]" pageId="23" pageNumber="24">Distribution and habitat</emphasis>
|
||
. Endemic to the
|
||
<typeStatus id="54F48871BF608319FD121015FD7F1E68" box="[683,731,1807,1831]" pageId="23" pageNumber="24">type</typeStatus>
|
||
locality, a warm spring complex along the east side of the Río Conchos (
|
||
<figureCitation id="13742A56BF608319FEB21028FEEB1E05" box="[267,335,1842,1866]" captionStart="FIGURE 1" captionStartId="2.[151,244,1795,1816]" captionTargetBox="[179,1402,157,1789]" captionTargetId="figure@2.[179,1423,125,1806]" captionTargetPageId="2" captionText="FIGURE 1. Map showing the distribution of Tryonia species in the Chihuahuan Desert. 1. Tryonia hertleini, Las Palomas. 2. Tryonia peregrina sp. nov., Ojo Vareleño, northwest of Casas Grandes. 3. Tryonia peregrina sp. nov., Yepomera. 4. Tryonia zaragozae sp. nov., northeast of Ignacio Zaragoza. 5. Tryonia angostura sp. nov., T. ovata sp. nov., south of Galeana. 6. Tryonia molinae sp. nov., El Molino. 7. Tryonia taylori sp. nov., vicinity of Rancho Ojo Caliente. 8. Tryonia contrerasi sp. nov., vicinity of Rancho Nuevo. 9. Tryonia chuviscarae sp. nov., T. minckleyi sp. nov., Balneario de San Diego de Alcala. 10. Tryonia julimesensis sp. nov., south of Julimes. 11. Tryonia allendae sp. nov., southwest of Talamantes. 12. Tryonia metcalfi sp. nov., La Cienega, south of Candelaria. 13. Tryonia oasiensis sp. nov., Caroline Spring, Oasis Ranch. 14. Tryonia cheatumi, Phantom Lake Spring. 15. Tryonia circumstriata, Diamond Y Draw. 16. Tryonia seemani, Ojo de Agua del Obispo, Durango City. 17. Tryonia seemani, Nombre de Dios. 18. Tryonia seemani, Amado Nervo." httpUri="https://zenodo.org/record/278443/files/figure.png" pageId="23" pageNumber="24">Fig. 1</figureCitation>
|
||
, locality 10).
|
||
<taxonomicName id="4C4F4D50BF608319FE4A102EFD781E04" box="[499,732,1843,1867]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="julimesensis">
|
||
<emphasis id="B93BEAC1BF608319FE4A102EFD781E04" box="[499,732,1843,1867]" italics="true" pageId="23" pageNumber="24">Tryonia julimesensis</emphasis>
|
||
</taxonomicName>
|
||
was abundant (on hard substrate and in detritus) in the main spring runs (44°C) (
|
||
<figureCitation id="13742A56BF608319FEC1104CFE1F1E21" box="[376,443,1878,1902]" captionStart="FIGURE 6" captionStartId="11.[151,244,1710,1731]" captionTargetBox="[221,1371,156,1678]" captionTargetId="figure@11.[212,1381,149,1685]" captionTargetPageId="11" captionText="FIGURE 6. Photographs of habitats of species treated in this paper. A. Ojo Vareleño, Chihuahua, type locality of T. peregrina sp. nov. (photograph taken on 16 / ix / 1990). B. Main spring in Yepomera, Chihuahua, T. peregrina habitat (18 / ix / 1990). C. Springs northeast of Ignacio Zaragoza, Chihuaua, type locality of T. zaragozae sp. nov. (8 / xii / 1998). D. Western-most head spring, Ojos de Arrey, Chihuahua, type locality of T. angosturae sp. nov. and T. ovata sp. nov. (7 / xii / 1998). E. Spring in northern part of El Molino, Chihuahua, type locality of T. molinae sp. nov. (26 / iv / 2010). F. Ojo del Apache, Chihuahua, type locality of T. contrerasi sp. nov. (14 / xii / 1998). G. Ojo Carbonero, Chihuahua, T. contrerasi habitat (2 / iv / 1991). H. Outflow of Ojo Caliente, Chihuahua, type locality of T. taylori sp. nov. (7 / xii / 1998). I – J. El Ojo de Servin, Chihuahua, T. taylori habitat (27 - viii- 1971, 7 / xii / 1998, respectively). K. Seeps in La Cienega, Texas, type locality of T. metcalfi sp. nov. (6 / xii / 2001). L – M. Balneario de San Diego de Alcala, Chihuahua, type locality of T. chuviscarae sp. nov. and T. minckleyi sp. nov. (4 / xii / 2001). N. Springs southeast of Julimes, Chihuahua, type locality of T. julimesensis sp. nov. (5 / iv / 1991). O. El Ojito (outflow), southwest of Talamantes, Chihuahua, type locality of T. allendae sp. nov. (10 / xii / 1998). P. Caroline Spring, Oasis Ranch, Texas, type locality of T. oasiensis sp. nov. (30 / iii / 2009). Q. Spring in Nombre de Dios, Durango, T. seemani habitat (2 / ix / 1971). R. Spring in Amado Nervo, Durango, T. seemani habitat (7 / iv / 1991)." httpUri="https://zenodo.org/record/278448/files/figure.png" pageId="23" pageNumber="24">Fig. 6</figureCitation>
|
||
N) when first discovered in 1991. It was also found in a small rheocrene just to the west of the main spring pools (44°C) as well as in semi-aquatic habitats on exposed rootlets and stones. When visited in 2001 the springs had been draglined (excavated) and
|
||
<taxonomicName id="4C4F4D50BF608319FD5010BAFCD21EF9" box="[745,886,1950,1975]" class="Gastropoda" family="Hydrobiidae" genus="Tryonia" kingdom="Animalia" order="Littorinimorpha" pageId="23" pageNumber="24" phylum="Mollusca" rank="species" species="julimensis">
|
||
<emphasis id="B93BEAC1BF608319FD5010BAFCD21EF9" box="[745,886,1950,1975]" italics="true" pageId="23" pageNumber="24">T. julimensis</emphasis>
|
||
</taxonomicName>
|
||
could not be found. We presume that this species is now extinct.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |