treatments-xml/data/03/E6/87/03E687BAFFD2F339FF5EF96C0ECDC6E3.xml
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<document ID-DOI="10.11646/zootaxa.3219.1.1" ID-GBIF-Dataset="3fa772dc-c5b2-40c9-b28e-cd067ca9a3f6" ID-ISSN="1175-5326" ID-Zenodo-Dep="280223" checkinTime="1460447300036" checkinUser="plazi" docAuthor="Fikáček, Martin, Jia, Fenglong &amp; Prokin, Alexander" docDate="2012" docId="03E687BAFFD2F339FF5EF96C0ECDC6E3" docLanguage="en" docName="zt03219p053.pdf" docOrigin="Zootaxa 3219" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.3:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="3" docTitle="Pachysternum kubani Fikáček, Jia &amp; Prokin, 2012, sp. nov." docType="treatment" docVersion="12" lastPageNumber="34" masterDocId="FFDFFFC2FFCCF318FFC9FFDB0C07C424" masterDocTitle="A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae)" masterLastPageNumber="53" masterPageNumber="1" pageNumber="31" updateTime="1678199402054" updateUser="tatiana">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A review of the Asian species of the genus Pachysternum (Coleoptera: Hydrophilidae: Sphaeridiinae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fikáček, Martin</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jia, Fenglong</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Prokin, Alexander</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Zootaxa</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2012</mods:date>
<mods:detail type="pubDate">
<mods:number>2012-03-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3219</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>53</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.11646/zootaxa.3219.1.1</mods:identifier>
<mods:identifier type="GBIF-Dataset">3fa772dc-c5b2-40c9-b28e-cd067ca9a3f6</mods:identifier>
<mods:identifier type="ISSN">1175-5326</mods:identifier>
<mods:identifier type="Zenodo-Dep">280223</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6180440" ID-GBIF-Taxon="119509790" ID-Zenodo-Dep="6180440" LSID="urn:lsid:plazi:treatment:03E687BAFFD2F339FF5EF96C0ECDC6E3" httpUri="http://treatment.plazi.org/id/03E687BAFFD2F339FF5EF96C0ECDC6E3" lastPageId="33" lastPageNumber="34" pageId="30" pageNumber="31">
<subSubSection box="[151,518,1719,1745]" pageId="30" pageNumber="31" type="nomenclature">
<paragraph blockId="30.[151,518,1719,1778]" box="[151,518,1719,1745]" pageId="30" pageNumber="31">
<heading bold="true" box="[151,518,1719,1745]" fontSize="11" level="1" pageId="30" pageNumber="31" reason="1">
<emphasis bold="true" box="[151,518,1719,1745]" pageId="30" pageNumber="31">
<taxonomicName ID-CoL="4BV58" authorityName="Fikáček &amp; Jia &amp; Prokin" box="[151,417,1719,1745]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="kubani" status="sp. nov.">
<emphasis bold="true" box="[151,417,1719,1745]" italics="true" pageId="30" pageNumber="31">Pachysternum kubani</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[424,518,1719,1745]" pageId="30" pageNumber="31" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[151,386,1754,1778]" pageId="30" pageNumber="31" type="description">
<paragraph blockId="30.[151,518,1719,1778]" box="[151,386,1754,1778]" pageId="30" pageNumber="31">
(
<figureCitation box="[159,252,1754,1778]" captionStart="FIGURES 26 28" captionStartId="6.[151,264,735,758]" captionTargetBox="[154,1436,187,727]" captionTargetId="figure@6.[151,1436,187,728]" captionTargetPageId="6" captionText="FIGURES 26 28. Characters of male genitalia of possible phylogenetic significance (a: detail of apex, b: detail of basal struts). 26: median lobe of P. haemorrhoum; 27: median lobe of P. kubani; 28: apical portion of the aedeagus of P. nigrovittatum. Abbreviations: lr: lateral rim." httpUri="https://zenodo.org/record/280226/files/figure.png" pageId="30" pageNumber="31">Figs. 27</figureCitation>
,
<figureCitation box="[264,294,1754,1778]" captionStart="FIGURES 3335" captionStartId="9.[151,264,1686,1709]" captionTargetBox="[166,1425,193,1639]" captionTargetId="figure@9.[151,1436,193,1645]" captionText="FIGURES 3335. Male genitalia of Asian Pachysternum. 33: P. kubani (ac, f: holotype; de: dark specimen from Sichuan; g: specimen from Laos, Ban Nape; h: terratological specimen from Laos, Phou Pane Mt.; a: tegmen, b, d: median lobe, c, e: apical portion of median lobe, fh: sternite 9); 34: P. stevensi (a: tegmen, b-c: median lobe, de: variation of the shape of sternite 9); 35: P. sulawesicum (a: tegmen, b: median lobe, c: apical portion of median lobe, d: sternite 9)." pageId="30" pageNumber="31">33</figureCitation>
, 45,
<figureCitation box="[348,377,1754,1778]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="30" pageNumber="31">60</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31" type="materials_examined">
<paragraph blockId="30.[151,1436,1825,2030]" pageId="30" pageNumber="31">
<emphasis bold="true" box="[151,305,1826,1851]" pageId="30" pageNumber="31">
<typeStatus box="[151,207,1827,1851]" pageId="30" pageNumber="31">Type</typeStatus>
locality.
</emphasis>
<collectingCountry box="[313,369,1827,1850]" name="Laos" pageId="30" pageNumber="31">Laos</collectingCountry>
, Houaphan province, Mt. Phou Pane, Phongsali env.,
<number box="[963,994,1826,1850]" pageId="30" pageNumber="31" value="20.0">20</number>
°
<number box="[1003,1030,1826,1850]" pageId="30" pageNumber="31" value="13.0">13</number>
ʹ09
<number box="[1081,1107,1826,1850]" pageId="30" pageNumber="31" value="19.0">19</number>
ʺN
<number box="[1146,1188,1826,1850]" pageId="30" pageNumber="31" value="103.0">103</number>
°
<number box="[1197,1227,1826,1850]" pageId="30" pageNumber="31" value="59.0">59</number>
ʹ
<number box="[1233,1261,1826,1850]" pageId="30" pageNumber="31" value="54.0">54</number>
ʺ–
<number box="[1285,1328,1826,1850]" pageId="30" pageNumber="31" value="104.0">104</number>
°00ʹ0
<number box="[1387,1400,1826,1850]" pageId="30" pageNumber="31" value="3.0">3</number>
ʺE,
<quantity box="[151,302,1862,1886]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.51" metricValueMin="1.48" pageId="30" pageNumber="31" unit="m" value="1495.0" valueMax="1510.0" valueMin="1480.0">14801510m</quantity>
.
</paragraph>
<paragraph blockId="30.[151,1436,1825,2030]" lastBlockId="31.[151,1437,151,1965]" lastPageId="31" lastPageNumber="32" pageId="30" pageNumber="31">
<emphasis bold="true" box="[199,375,1898,1923]" pageId="30" pageNumber="31">
<typeStatus box="[199,255,1899,1923]" pageId="30" pageNumber="31">Type</typeStatus>
material.
</emphasis>
<materialsCitation ID-GBIF-Occurrence="3499776473" collectionCode="NMPC, PHOU, PANE, NHMB" elevation="1495" pageId="30" pageNumber="31" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus box="[391,493,1898,1922]" pageId="30" pageNumber="31">Holotype</typeStatus>
:
<specimenCount box="[515,568,1898,1922]" pageId="30" pageNumber="31" type="male">male</specimenCount>
(
<collectionCode box="[590,675,1898,1922]" collectionName="Czech Republic, Prague, National Museum (Natural History)" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="30" pageNumber="31">NMPC</collectionCode>
): “LAOS-NE, Houa Phan prov. /
<number box="[1087,1118,1898,1922]" pageId="30" pageNumber="31" value="20.0">20</number>
°
<number box="[1127,1154,1898,1922]" pageId="30" pageNumber="31" value="13.0">13</number>
ʹ09
<number box="[1204,1230,1898,1922]" pageId="30" pageNumber="31" value="19.0">19</number>
ʺN
<number box="[1277,1319,1898,1922]" pageId="30" pageNumber="31" value="103.0">103</number>
°
<number box="[1329,1359,1898,1922]" pageId="30" pageNumber="31" value="59.0">59</number>
ʹ
<number box="[1364,1392,1898,1922]" pageId="30" pageNumber="31" value="54.0">54</number>
ʺ- /
<number box="[151,194,1934,1958]" pageId="30" pageNumber="31" value="104.0">104</number>
°00ʹ0
<number box="[253,266,1934,1958]" pageId="30" pageNumber="31" value="3.0">3</number>
ʺE,
<quantity box="[309,457,1934,1958]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.51" metricValueMin="1.48" pageId="30" pageNumber="31" unit="m" value="1495.0" valueMax="1510.0" valueMin="1480.0">
<elevation box="[309,457,1934,1958]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" metricValueMax="1.51" metricValueMin="1.48" pageId="30" pageNumber="31" unit="m" value="1495.0" valueMax="1510.0" valueMin="1480.0">14801510m</elevation>
</quantity>
/
<collectionCode box="[480,556,1934,1958]" pageId="30" pageNumber="31">PHOU</collectionCode>
<collectionCode box="[563,634,1935,1959]" pageId="30" pageNumber="31">PANE</collectionCode>
Mt.,
<date box="[696,869,1934,1958]" pageId="30" pageNumber="31" value="2009-06-01" valueMax="2009-06-16" valueMin="2009-06-01">
<number box="[696,755,1934,1958]" pageId="30" pageNumber="31" value="8.5" valueMax="16.0" valueMin="1.0">116</number>
.vi. /
<number box="[810,869,1934,1958]" pageId="30" pageNumber="31" value="2009.0">2009</number>
</date>
, Vít Kubáň leg. // Primary mountain forest / flight intercept trap / Laos
<number box="[380,436,1970,1994]" pageId="30" pageNumber="31" value="2009.0">2009</number>
Expedition /
<collectionCode box="[586,670,1971,1994]" collectionName="Switzerland, Basel, Naturhistorisches Museum" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="30" pageNumber="31">NHMB</collectionCode>
Basel / NMPC Prague”.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3499776328" collectionCode="NMPC" pageId="30" pageNumber="31" specimenCount="2" specimenCount-female="1" typeStatus="paratype">
<typeStatus box="[949,1058,1971,1995]" pageId="30" pageNumber="31">Paratypes</typeStatus>
:
<specimenCount box="[1072,1172,1970,1994]" pageId="30" pageNumber="31" type="female">
<number box="[1072,1086,1970,1994]" pageId="30" pageNumber="31" value="1.0">1</number>
female
</specimenCount>
,
<specimenCount box="[1182,1260,1970,1994]" pageId="30" pageNumber="31" type="generic">
<number box="[1182,1196,1970,1994]" pageId="30" pageNumber="31" value="1.0">1</number>
spec.
</specimenCount>
(
<collectionCode box="[1274,1360,1970,1994]" collectionName="Czech Republic, Prague, National Museum (Natural History)" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="30" pageNumber="31">NMPC</collectionCode>
): same data as the holotype;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3499776397" collectingDate="2009-05-10" collectingDateMax="2009-05-30" collectingDateMin="2009-05-10" collectionCode="NMPC, PHOU, SANE, NHMB" country="Laos" elevation="1420" lastPageId="31" lastPageNumber="32" latitude="19.636667" location="Basel" longLatPrecision="12" longitude="103.33667" municipality="Expedition" pageId="30" pageNumber="31" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Xieng Khouang">
<specimenCount box="[402,482,2006,2030]" pageId="30" pageNumber="31" type="male">
<number box="[402,416,2006,2030]" pageId="30" pageNumber="31" value="1.0">1</number>
male
</specimenCount>
,
<specimenCount box="[496,595,2006,2030]" pageId="30" pageNumber="31" type="female">
<number box="[496,510,2006,2030]" pageId="30" pageNumber="31" value="1.0">1</number>
female
</specimenCount>
(
<collectionCode box="[613,698,2006,2030]" collectionName="Czech Republic, Prague, National Museum (Natural History)" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="30" pageNumber="31">NMPC</collectionCode>
): “LAOS-NE,
<collectingRegion box="[867,1045,2006,2030]" country="Laos" name="Xiangkhouang" pageId="30" pageNumber="31">Xieng Khouang</collectingRegion>
/ prov.,
<geoCoordinate box="[1145,1271,2005,2030]" degrees="19" direction="north" minutes="38.20" orientation="latitude" pageId="30" pageNumber="31" precision="9" value="19.636667">
<number box="[1145,1171,2006,2030]" pageId="30" pageNumber="31" value="19.0">19</number>
°
<number box="[1179,1245,2006,2030]" pageId="30" pageNumber="31" value="38.2">38.20</number>
ʹN
</geoCoordinate>
<geoCoordinate box="[1281,1419,2005,2030]" degrees="103" direction="east" minutes="20.20" orientation="longitude" pageId="30" pageNumber="31" precision="9" value="103.33667">
<number box="[1281,1321,2006,2030]" pageId="30" pageNumber="31" value="103.0">103</number>
°
<number box="[1330,1396,2006,2030]" pageId="30" pageNumber="31" value="20.2">20.20</number>
ʹE
</geoCoordinate>
/ Phonsavan (
<locationDeviation bearing="45.0" box="[287,411,151,175]" distance="30000.0" pageId="31" pageNumber="32">
<quantity box="[287,361,151,175]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="km" value="30.0">30 km</quantity>
NE
</locationDeviation>
): /
<collectionCode box="[448,523,151,175]" pageId="31" pageNumber="32">PHOU</collectionCode>
<collectionCode box="[532,604,151,175]" pageId="31" pageNumber="32">SANE</collectionCode>
Mt.,
<quantity box="[668,745,151,175]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="31" pageNumber="32" unit="m" value="1420.0">
<elevation box="[668,745,151,175]" metricMagnitude="3" metricUnit="m" metricValue="1.42" pageId="31" pageNumber="32" unit="m" value="1420.0">1420m</elevation>
</quantity>
/
<date box="[770,927,151,175]" pageId="31" pageNumber="32" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<collectingDate box="[770,927,151,175]" pageId="31" pageNumber="32" value="2009-05-10" valueMax="2009-05-30" valueMin="2009-05-10">
<number box="[770,800,151,175]" pageId="31" pageNumber="32" value="10.0">10</number>
.
<number box="[817,846,151,175]" pageId="31" pageNumber="32" value="30.0">30</number>
.v.
<number box="[867,927,151,175]" pageId="31" pageNumber="32" value="2009.0">2009</number>
</collectingDate>
</date>
, V. Kubáň lgt. // secondary mountain forest / flight intercept trap /
<collectingCountry box="[386,440,189,212]" name="Laos" pageId="31" pageNumber="32">Laos</collectingCountry>
<number box="[448,503,188,212]" pageId="31" pageNumber="32" value="2009.0">2009</number>
<collectingMunicipality box="[510,631,188,212]" pageId="31" pageNumber="32">Expedition</collectingMunicipality>
/
<collectionCode box="[654,737,189,212]" collectionName="Switzerland, Basel, Naturhistorisches Museum" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="31" pageNumber="32">NHMB</collectionCode>
<location LSID="urn:lsid:plazi:treatment:03E687BAFFD2F339FF5EF96C0ECDC6E3:8E906077FFD3F307FD20FF670F21C4F0" box="[745,806,188,212]" country="Laos" latitude="19.636667" longLatPrecision="12" longitude="103.33667" municipality="Expedition" name="Basel" pageId="31" pageNumber="32" stateProvince="Xieng Khouang">Basel</location>
/
<collectionCode box="[828,906,188,212]" collectionName="Czech Republic, Prague, National Museum (Natural History)" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="31" pageNumber="32">NMPC</collectionCode>
<location LSID="urn:lsid:plazi:treatment:03E687BAFFD2F339FF5EF96C0ECDC6E3:8E906077FFD3F307FC5BFF670FE3C4F0" box="[914,996,188,212]" country="Laos" latitude="19.636667" longLatPrecision="12" longitude="103.33667" municipality="Expedition" name="Prague" pageId="31" pageNumber="32" stateProvince="Xieng Khouang">Prague</location>
”;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3499776324" collectingDate="2004-05-06" collectingDateMax="2004-05-17" collectingDateMin="2004-05-06" collectionCode="NHMB, BMNH, NHMW" collectorName="Vit Kuban" country="Laos" elevation="1500" latitude="21.683332" location="PHONGSALY env." longLatPrecision="1263" longitude="102.1" municipality="Phongsaly" pageId="31" pageNumber="32" specimenCount="5" specimenCount-female="2" specimenCount-male="3">
<specimenCount box="[1018,1106,188,212]" pageId="31" pageNumber="32" type="male">
<number box="[1018,1032,188,212]" pageId="31" pageNumber="32" value="3.0">3</number>
males
</specimenCount>
,
<specimenCount box="[1118,1228,188,212]" pageId="31" pageNumber="32" type="female">
<number box="[1118,1132,188,212]" pageId="31" pageNumber="32" value="2.0">2</number>
females
</specimenCount>
. (
<collectionCode box="[1247,1334,188,212]" collectionName="Switzerland, Basel, Naturhistorisches Museum" httpUri="http://grbio.org/cool/1d5a-4efs" name="Naturhistorisches Museum, Basel" pageId="31" pageNumber="32">NHMB</collectionCode>
,
<collectionCode box="[1346,1432,189,212]" collectionName="United Kingdom, London, The Natural History Museum [formerly British Museum (Natural History)]" pageId="31" pageNumber="32">BMNH</collectionCode>
,
<collectionCode box="[151,247,223,247]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="32">NHMW</collectionCode>
): “
<collectingCountry box="[278,339,223,247]" name="Laos" pageId="31" pageNumber="32">LAO</collectingCountry>
,
<collectingMunicipality box="[354,469,223,247]" pageId="31" pageNumber="32">Phongsaly</collectingMunicipality>
prov. /
<geoCoordinate box="[562,654,222,247]" degrees="21" direction="north" minutes="41" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="21.683332">
<number box="[562,590,223,247]" pageId="31" pageNumber="32" value="21.0">21</number>
°
<number box="[599,628,223,247]" pageId="31" pageNumber="32" value="41.0">41</number>
ʹN
</geoCoordinate>
<geoCoordinate box="[664,753,222,247]" degrees="102" direction="east" minutes="6" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="102.1">
<number box="[664,707,223,247]" pageId="31" pageNumber="32" value="102.0">102</number>
°
<number box="[716,730,223,247]" pageId="31" pageNumber="32" value="6.0">6</number>
ʹE
</geoCoordinate>
/
<location LSID="urn:lsid:plazi:treatment:03E687BAFFD2F339FF5EF96C0ECDC6E3:8E906077FFD3F307FCC5FF040FEDC4D3" box="[780,1002,223,247]" country="Laos" latitude="21.683332" longLatPrecision="1263" longitude="102.1" municipality="Phongsaly" name="PHONGSALY env." pageId="31" pageNumber="32">PHONGSALY env.</location>
, /
<date box="[1034,1177,223,247]" pageId="31" pageNumber="32" value="2004-05-06" valueMax="2004-05-17" valueMin="2004-05-06">
<collectingDate box="[1034,1177,223,247]" pageId="31" pageNumber="32" value="2004-05-06" valueMax="2004-05-17" valueMin="2004-05-06">
<number box="[1034,1050,223,247]" pageId="31" pageNumber="32" value="6.0">6</number>
.
<number box="[1066,1093,223,247]" pageId="31" pageNumber="32" value="17.0">17</number>
.v.
<number box="[1115,1177,223,247]" pageId="31" pageNumber="32" value="2004.0">2004</number>
</collectingDate>
</date>
, ~
<quantity box="[1205,1285,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="1500.0">
<elevation box="[1205,1285,223,247]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="31" pageNumber="32" unit="m" value="1500.0">1500m</elevation>
</quantity>
, /
<collectorName box="[1317,1436,223,247]" pageId="31" pageNumber="32">Vít Kubáň</collectorName>
leg.”;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3499776415" collectingDate="2001-05-01" collectingDateMax="2001-05-18" collectingDateMin="2001-05-01" collectionCode="NMHB, BMNH, NMPC" collectorName="Vit Kuban" country="Laos" elevation="600" latitude="18.35" location="Ban Nape" longLatPrecision="1275" longitude="105.13333" municipality="Bolikhamsai" pageId="31" pageNumber="32" specimenCount="8" specimenCount-male="8">
<specimenCount box="[218,307,260,284]" pageId="31" pageNumber="32" type="male">
<number box="[218,232,260,284]" pageId="31" pageNumber="32" value="8.0">8</number>
males
</specimenCount>
,
<number box="[319,346,260,284]" pageId="31" pageNumber="32" value="12.0">12</number>
spec. (
<collectionCode box="[426,513,260,284]" pageId="31" pageNumber="32">NMHB</collectionCode>
,
<collectionCode box="[525,611,261,284]" pageId="31" pageNumber="32">BMNH</collectionCode>
,
<collectionCode box="[622,706,260,284]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="31" pageNumber="32">NMPC</collectionCode>
): “
<collectingCountry box="[735,812,260,284]" name="Laos" pageId="31" pageNumber="32">LAOS</collectingCountry>
,
<date box="[824,965,260,284]" pageId="31" pageNumber="32" value="2001-05-01" valueMax="2001-05-18" valueMin="2001-05-01">
<collectingDate box="[824,965,260,284]" pageId="31" pageNumber="32" value="2001-05-01" valueMax="2001-05-18" valueMin="2001-05-01">
<number box="[824,839,260,284]" pageId="31" pageNumber="32" value="1.0">1</number>
.
<number box="[857,884,260,284]" pageId="31" pageNumber="32" value="18.0">18</number>
.v.
<number box="[905,965,260,284]" pageId="31" pageNumber="32" value="2001.0">2001</number>
</collectingDate>
</date>
/
<collectingMunicipality box="[988,1127,260,284]" pageId="31" pageNumber="32">Bolikhamsai</collectingMunicipality>
prov. /
<geoCoordinate box="[1213,1306,260,284]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="925" value="18.35">
<number box="[1213,1241,260,284]" pageId="31" pageNumber="32" value="18.0">18</number>
°
<number box="[1251,1280,260,284]" pageId="31" pageNumber="32" value="21.0">21</number>
'N
</geoCoordinate>
,
<geoCoordinate box="[1319,1420,260,284]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="925" value="105.13333">
<number box="[1319,1361,260,284]" pageId="31" pageNumber="32" value="105.0">105</number>
°08'E
</geoCoordinate>
/
<location LSID="urn:lsid:plazi:treatment:03E687BAFFD2F339FF5EF96C0ECDC6E3:8E906077FFD3F307FF5EFEF30D02C51B" box="[151,261,296,319]" country="Laos" latitude="18.35" longLatPrecision="1275" longitude="105.13333" municipality="Bolikhamsai" name="Ban Nape" pageId="31" pageNumber="32">Ban Nape</location>
(
<locationDeviation bearing="45.0" box="[278,381,295,319]" distance="8000.0" pageId="31" pageNumber="32">
<quantity box="[278,335,295,319]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="31" pageNumber="32" unit="km" value="8.0">8 km</quantity>
NE
</locationDeviation>
) / ~
<quantity box="[435,507,295,319]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="32" unit="m" value="600.0">
<elevation box="[435,507,295,319]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="31" pageNumber="32" unit="m" value="600.0">600 m</elevation>
</quantity>
,
<collectorName box="[520,637,295,319]" pageId="31" pageNumber="32">Vít Kubáň</collectorName>
lgt.”;
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="3499776346" collectionCode="NHMW, NMPC, SYSU, KSEM" collectorName="A. Putz" country="China" elevation="900" latitude="30.53" location="Qingcheng Shan" longLatPrecision="732" longitude="103.33" municipality="Sichuan" pageId="31" pageNumber="32" specimenCount="12" specimenCount-male="1">
<specimenCount box="[707,784,295,319]" pageId="31" pageNumber="32" type="male">
<number box="[707,721,295,319]" pageId="31" pageNumber="32" value="1.0">1</number>
male
</specimenCount>
,
<specimenCount box="[797,883,295,319]" pageId="31" pageNumber="32">
<number box="[797,824,295,319]" pageId="31" pageNumber="32" value="11.0">11</number>
spec
</specimenCount>
. (
<collectionCode box="[902,995,295,319]" httpUri="http://grbio.org/cool/91g3-0mnw" name="Naturhistorisches Museum, Wien" pageId="31" pageNumber="32">NHMW</collectionCode>
,
<collectionCode box="[1006,1087,295,319]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="31" pageNumber="32">NMPC</collectionCode>
,
<collectionCode LSID="urn:lsid:biocol.org:col:13728" box="[1099,1172,295,319]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13728" name="National Sun Yat-Sen University, Department of Biological Sciences" pageId="31" pageNumber="32">SYSU</collectionCode>
,
<collectionCode box="[1184,1266,295,319]" pageId="31" pageNumber="32">KSEM</collectionCode>
): “
<collectingCountry box="[1296,1366,295,319]" name="China" pageId="31" pageNumber="32">China</collectingCountry>
,
<collectingMunicipality pageId="31" pageNumber="32">Sichuan</collectingMunicipality>
,
<location LSID="urn:lsid:plazi:treatment:03E687BAFFD2F339FF5EF96C0ECDC6E3:8E906077FFD3F307FF07FE970D81C540" box="[206,390,332,356]" country="China" latitude="30.53" longLatPrecision="732" longitude="103.33" municipality="Sichuan" name="Qingcheng Shan" pageId="31" pageNumber="32">Qingcheng Shan</location>
/
<locationDeviation box="[414,648,332,356]" pageId="31" pageNumber="32">
<quantity box="[414,485,332,356]" metricMagnitude="4" metricUnit="m" metricValue="6.5" pageId="31" pageNumber="32" unit="km" value="65.0">65 km</quantity>
NW Chengdu
</locationDeviation>
,
<geoCoordinate box="[660,755,332,356]" direction="east" orientation="longitude" pageId="31" pageNumber="32" precision="555" value="103.33">
<number box="[660,737,332,356]" pageId="31" pageNumber="32" value="103.33">103.33</number>
E
</geoCoordinate>
<geoCoordinate box="[763,845,332,356]" direction="north" orientation="latitude" pageId="31" pageNumber="32" precision="555" value="30.53">
<number box="[763,825,332,356]" pageId="31" pageNumber="32" value="30.53">30.53</number>
N
</geoCoordinate>
/
<number box="[870,899,332,356]" pageId="31" pageNumber="32" value="18.0">18</number>
.v./
<date box="[933,1068,332,356]" pageId="31" pageNumber="32" value="1997-06-03" valueMax="1997-06-04" valueMin="1997-06-03">
<number box="[933,947,332,356]" pageId="31" pageNumber="32" value="3.0">3</number>
.
<number box="[962,977,332,356]" pageId="31" pageNumber="32" value="4.0">4</number>
.vi.1997
</date>
,
<locationDeviation box="[1081,1265,332,356]" pageId="31" pageNumber="32">8 km W Taiping</locationDeviation>
/
<quantity box="[1289,1432,332,356]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="31" pageNumber="32" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">
<elevation box="[1289,1432,332,356]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="31" pageNumber="32" unit="m" value="900.0" valueMax="1000.0" valueMin="800.0">8001000 m</elevation>
</quantity>
, leg.
<collectorName box="[198,285,367,392]" pageId="31" pageNumber="32">A. Pütz</collectorName>
”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="31" pageNumber="32" type="diagnosis">
<paragraph blockId="31.[151,1437,151,1965]" pageId="31" pageNumber="32">
<emphasis bold="true" box="[199,462,403,428]" pageId="31" pageNumber="32">Differential diagnosis.</emphasis>
Pale specimens of
<taxonomicName box="[679,923,404,428]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kubani" status="sp. nov.">
<emphasis box="[679,923,404,428]" italics="true" pageId="31" pageNumber="32">Pachysternum kubani</emphasis>
</taxonomicName>
<emphasis bold="true" box="[931,1021,404,428]" pageId="31" pageNumber="32">
<taxonomicNameLabel box="[931,1021,404,428]" pageId="31" pageNumber="32" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation box="[1036,1163,404,428]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="31" pageNumber="32">
Figs.
<number box="[1100,1128,404,428]" pageId="31" pageNumber="32" value="60.0">60</number>
af
</figureCitation>
) are easily recognizable from other
<taxonomicName box="[278,438,439,463]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus">
<emphasis box="[278,438,439,463]" italics="true" pageId="31" pageNumber="32">Pachysternum</emphasis>
</taxonomicName>
species by pale yellowish elytra with irregularly lobate transverse dark stripe in anterior third (
<figureCitation box="[218,346,475,499]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="31" pageNumber="32">
Figs.
<number box="[281,309,475,499]" pageId="31" pageNumber="32" value="60.0">60</number>
ae
</figureCitation>
) or by very distinct pale humeral spots (
<figureCitation box="[794,886,475,499]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="31" pageNumber="32">
Fig.
<number box="[846,876,475,499]" pageId="31" pageNumber="32" value="60.0">60</number>
f
</figureCitation>
), and distinctly bisinuate outer margin of anterior tibiae. By the coloration, these specimens may be confused with
<taxonomicName box="[882,998,512,535]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="stevensi">
<emphasis box="[882,998,512,535]" italics="true" pageId="31" pageNumber="32">P. stevensi</emphasis>
</taxonomicName>
, from which they differ by the characters given in couplet
<number box="[388,416,546,570]" pageId="31" pageNumber="32" value="12.0">12</number>
of the identification key. The body form of large specimens (and also the presence of pale spots on pronotal disc in pale specimens) resembles
<taxonomicName box="[736,849,582,607]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis box="[736,849,582,607]" italics="true" pageId="31" pageNumber="32">P. cardoni</emphasis>
</taxonomicName>
; in contrast to it, the dark coloration of elytral series
<number box="[151,193,618,642]" pageId="31" pageNumber="32" value="3.0" valueMax="4.0" valueMin="2.0">24</number>
reaches at most the midlength in pale specimens of
<taxonomicName box="[772,875,618,642]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kubani">
<emphasis box="[772,875,618,642]" italics="true" pageId="31" pageNumber="32">P. kubani</emphasis>
</taxonomicName>
(it reaches subapically in
<taxonomicName box="[1166,1275,618,642]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis box="[1166,1275,618,642]" italics="true" pageId="31" pageNumber="32">P cardoni</emphasis>
</taxonomicName>
), both species differ in the color pattern of the pronotum (with dark M-shaped spot or at least distinct pair of pale spots submedially in
<taxonomicName box="[230,345,690,714]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="cardoni">
<emphasis box="[230,345,690,714]" italics="true" pageId="31" pageNumber="32">P. cardoni</emphasis>
</taxonomicName>
, with anteriorly trilobate dark spot or completely dark pronotal disc in
<taxonomicName box="[1152,1256,690,714]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kubani">
<emphasis box="[1152,1256,690,714]" italics="true" pageId="31" pageNumber="32">P. kubani</emphasis>
</taxonomicName>
) and the elytral microsculpture is also frequently different (absent in
<taxonomicName box="[763,869,725,750]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="kubani">
<emphasis box="[763,869,725,750]" italics="true" pageId="31" pageNumber="32">P. kubani</emphasis>
</taxonomicName>
, distinctly developed or rarely missing in
<emphasis italics="true" pageId="31" pageNumber="32">P. c a rdoni</emphasis>
). Dark specimens, which are usually slightly smaller than paler ones, highly resemble the mainland forms of
<taxonomicName box="[151,283,798,821]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="apicatum">
<emphasis box="[151,283,798,821]" italics="true" pageId="31" pageNumber="32">P. apicatum</emphasis>
</taxonomicName>
(with which they sometimes even co-occur) in the coloration and very often also in the shape of anterior tibia; they may be distinguished from
<taxonomicName box="[621,752,834,857]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="apicatum">
<emphasis box="[621,752,834,857]" italics="true" pageId="31" pageNumber="32">P. apicatum</emphasis>
</taxonomicName>
by the characters given in the couplet
<number box="[1181,1209,832,856]" pageId="31" pageNumber="32" value="15.0">15</number>
of the identification key.
</paragraph>
</subSubSection>
<subSubSection lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32" type="description">
<paragraph blockId="31.[151,1437,151,1965]" pageId="31" pageNumber="32">
<emphasis bold="true" box="[199,338,903,928]" pageId="31" pageNumber="32">Description</emphasis>
(based on the
<typeStatus box="[510,613,904,928]" pageId="31" pageNumber="32">holotype</typeStatus>
). Body widely oval, shape of elytra slightly sexually dimorphic, female with distinctly pronounced humeral portion of elytra. Body length
<quantity box="[920,1058,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="2.85" metricValueMax="3.5" metricValueMin="2.2" pageId="31" pageNumber="32" unit="mm" value="2.85" valueMax="3.5" valueMin="2.2">2.23.5 mm</quantity>
(
<typeStatus box="[1075,1179,940,964]" pageId="31" pageNumber="32">holotype</typeStatus>
:
<quantity box="[1192,1284,940,964]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="31" pageNumber="32" unit="mm" value="3.3">3.3 mm</quantity>
); body width
<quantity box="[151,286,975,999]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" metricValueMax="2.2" metricValueMin="1.6" pageId="31" pageNumber="32" unit="mm" value="1.9" valueMax="2.2" valueMin="1.6">1.62.2 mm</quantity>
(
<typeStatus box="[300,400,975,999]" pageId="31" pageNumber="32">holotype</typeStatus>
<quantity box="[407,497,975,999]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" pageId="31" pageNumber="32" unit="mm" value="2.2">2.2 mm</quantity>
).
</paragraph>
<paragraph blockId="31.[151,1437,151,1965]" pageId="31" pageNumber="32">
<emphasis box="[199,322,1011,1035]" italics="true" pageId="31" pageNumber="32">Coloration</emphasis>
(
<figureCitation box="[341,429,1011,1035]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="31" pageNumber="32">
Fig.
<number box="[397,429,1011,1035]" pageId="31" pageNumber="32" value="60.0">60</number>
</figureCitation>
). Clypeus and lateral portions of frons black, transverse ridge between eyes and large median portion of the head pale reddish. Pronotum widely pale reddish along lateral and anterior margins, bearing large posteriorly wide and anteriorly trilobate black spot. General coloration of elytral yellowish; interval
<number box="[1353,1367,1083,1107]" pageId="31" pageNumber="32" value="1.0">1</number>
black throughout, all elytral series black in anterior half, the black coloration slightly extending to intervals from series
<number box="[151,195,1154,1178]" pageId="31" pageNumber="32" value="3.5" valueMax="5.0" valueMin="2.0">25</number>
, but largely extending across intervals
<number box="[634,691,1154,1178]" pageId="31" pageNumber="32" value="8.5" valueMax="11.0" valueMin="6.0">611</number>
, forming large lateral black spot in anterior third of elytra. Ventral surface of head brown, mentum and thoracic sclerites black, abdominal ventrites dark brown anteriorly, largely pale reddish posteriorly. Legs brown, femora black.
</paragraph>
<paragraph blockId="31.[151,1437,151,1965]" pageId="31" pageNumber="32">
<emphasis box="[199,441,1261,1285]" italics="true" pageId="31" pageNumber="32">External morphology</emphasis>
. Clypeus with moderately coarse but rather sparse punctation consisting of rounded punctures of two slightly different sizes. Punctation of frons similar to that on clypeus, with punctures separated by
<number box="[151,194,1333,1357]" pageId="31" pageNumber="32" value="3.0" valueMax="4.0" valueMin="2.0">24</number>
× the puncture diameter. Interstices on head without microsculpture. Larger punctures on pronotum slightly crescent-like, deep, much larger than small punctures; small punctures slightly semicircular, divided from each other by ca.
<number box="[301,363,1404,1428]" pageId="31" pageNumber="32" value="1.25" valueMax="1.5" valueMin="1.0">11.5</number>
× puncture width. Pronotal interstices without microsculpture. Prosternum with very distinct median carina. Elytral series consisting of large, shallow punctures; interval punctation with rather fine semicircular punctures much smaller than serial ones, with sparsely intermixed slightly larger setiferous punctures of the same shape; interstices without microsculpture. Elytral series weakly impressed. Preepisternal elevation of mesothorax without posterolateral pits. Femoral lines on metaventrite indistinctly angulate, median portion of metaventrite bearing moderately coarse and dense punctation, interstices without microsculpture; lateral portions with very coarse and dense, densely pubescent punctation. Anterior tibia rather wide, outer margin distinctly bisinuate, outer series of spines interrupted at the place of emargination.
</paragraph>
<paragraph blockId="31.[151,1437,151,1965]" pageId="31" pageNumber="32">
<emphasis box="[199,364,1690,1714]" italics="true" pageId="31" pageNumber="32">
<collectingCountry box="[199,257,1690,1714]" name="Malta" pageId="31" pageNumber="32">Male</collectingCountry>
genitalia
</emphasis>
(
<figureCitation box="[381,464,1690,1714]" captionStart="FIGURES 3335" captionStartId="9.[151,264,1686,1709]" captionTargetBox="[166,1425,193,1639]" captionTargetId="figure@9.[151,1436,193,1645]" captionText="FIGURES 3335. Male genitalia of Asian Pachysternum. 33: P. kubani (ac, f: holotype; de: dark specimen from Sichuan; g: specimen from Laos, Ban Nape; h: terratological specimen from Laos, Phou Pane Mt.; a: tegmen, b, d: median lobe, c, e: apical portion of median lobe, fh: sternite 9); 34: P. stevensi (a: tegmen, b-c: median lobe, de: variation of the shape of sternite 9); 35: P. sulawesicum (a: tegmen, b: median lobe, c: apical portion of median lobe, d: sternite 9)." pageId="31" pageNumber="32">
Fig.
<number box="[435,464,1690,1714]" pageId="31" pageNumber="32" value="33.0">33</number>
</figureCitation>
). Tegmen
<quantity box="[586,723,1690,1714]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="31" pageNumber="32" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.31.5 mm</quantity>
long, median lobe
<quantity box="[944,1080,1690,1714]" metricMagnitude="-3" metricUnit="m" metricValue="1.4" metricValueMax="1.5" metricValueMin="1.3" pageId="31" pageNumber="32" unit="mm" value="1.4" valueMax="1.5" valueMin="1.3">1.31.5 mm</quantity>
long. Phallobase
<number box="[1284,1298,1690,1714]" pageId="31" pageNumber="32" value="2.0">2</number>
× as long as parameres, bearing small, slightly asymmetrical, indistinctly detached basal manubrium. Median lobe rather narrow, widest in basal third, weakly narrowing apicad, apex narrowly rounded; gonopore subapical, rather indistinct; lateral pubescent lobes indistinct even subapically. Sternite
<number box="[817,831,1797,1821]" pageId="31" pageNumber="32" value="9.0">9</number>
entire, rather narrow.
</paragraph>
<paragraph blockId="31.[151,1437,151,1965]" lastBlockId="33.[151,1436,151,711]" lastPageId="33" lastPageNumber="34" pageId="31" pageNumber="32">
<emphasis bold="true" box="[199,315,1833,1858]" pageId="31" pageNumber="32">Variation.</emphasis>
The species varies considerably in the extent of the black coloration on the head, pronotum and elytra. Coloration of the frons may vary from entirely dark brown to wide pale spot situated medially. Pronotum may bear large black spot which is wide posteriorly, but trilobate anteriorly, or may be completely black on the disc, with widely pale lateral margins (the pale areas widest anteriorly and narrowest posteriorly in that case) and median portion of anterior margin. Elytral coloration varies from largely reddish with irregular dark spots at anterior third of the elytral series (
<figureCitation box="[348,441,187,211]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="33" pageNumber="34">
Fig.
<number box="[400,429,187,211]" pageId="33" pageNumber="34" value="60.0">60</number>
c
</figureCitation>
) through reddish with a more or less extended irregularly shaped transverse dark stripe at anterior third (
<figureCitation box="[314,498,223,247]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="33" pageNumber="34">
Figs.
<number box="[378,406,223,247]" pageId="33" pageNumber="34" value="60.0">60</number>
ab, de
</figureCitation>
) to largely dark brown to black with large to very small apical reddish spot and in some cases also basal pale spots (
<figureCitation box="[523,644,259,283]" captionStart="FIGURE 60" captionStartId="32.[151,250,1042,1065]" captionTargetBox="[210,1357,184,1017]" captionTargetId="figure@32.[203,1390,163,1024]" captionText="FIGURE 60. Pachysternum kubani, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab: Laos, Phou Pane Mt.; c: same locality (holotype, male); d: Laos, Phongsaly env. (female); e: same locality (male); fh: Laos, Ban Nape; i: China, Sichuan, Qingcheng Shan." pageId="33" pageNumber="34">
Figs.
<number box="[586,612,259,283]" pageId="33" pageNumber="34" value="60.0">60</number>
fi
</figureCitation>
). The shape of the male sternite
<number box="[1007,1021,259,283]" pageId="33" pageNumber="34" value="9.0">9</number>
varies from rounded (in the
<typeStatus box="[1340,1436,259,283]" pageId="33" pageNumber="34">holotype</typeStatus>
and majority of the specimens examined;
<figureCitation box="[609,698,295,319]" captionStart="FIGURES 3335" captionStartId="9.[151,264,1686,1709]" captionTargetBox="[166,1425,193,1639]" captionTargetId="figure@9.[151,1436,193,1645]" captionText="FIGURES 3335. Male genitalia of Asian Pachysternum. 33: P. kubani (ac, f: holotype; de: dark specimen from Sichuan; g: specimen from Laos, Ban Nape; h: terratological specimen from Laos, Phou Pane Mt.; a: tegmen, b, d: median lobe, c, e: apical portion of median lobe, fh: sternite 9); 34: P. stevensi (a: tegmen, b-c: median lobe, de: variation of the shape of sternite 9); 35: P. sulawesicum (a: tegmen, b: median lobe, c: apical portion of median lobe, d: sternite 9)." pageId="33" pageNumber="34">
Fig.
<number box="[660,687,295,319]" pageId="33" pageNumber="34" value="33.0">33</number>
f
</figureCitation>
) to shallowly emarginate (rarely found at the same localities where specimens with rounded apex of the sternite
<number box="[642,656,330,354]" pageId="33" pageNumber="34" value="9.0">9</number>
predominate,
<figureCitation box="[814,909,330,354]" captionStart="FIGURES 3335" captionStartId="9.[151,264,1686,1709]" captionTargetBox="[166,1425,193,1639]" captionTargetId="figure@9.[151,1436,193,1645]" captionText="FIGURES 3335. Male genitalia of Asian Pachysternum. 33: P. kubani (ac, f: holotype; de: dark specimen from Sichuan; g: specimen from Laos, Ban Nape; h: terratological specimen from Laos, Phou Pane Mt.; a: tegmen, b, d: median lobe, c, e: apical portion of median lobe, fh: sternite 9); 34: P. stevensi (a: tegmen, b-c: median lobe, de: variation of the shape of sternite 9); 35: P. sulawesicum (a: tegmen, b: median lobe, c: apical portion of median lobe, d: sternite 9)." pageId="33" pageNumber="34">
Fig.
<quantity box="[865,909,330,354]" metricMagnitude="-2" metricUnit="kg" metricValue="3.3" pageId="33" pageNumber="34" unit="g" value="33.0">33g</quantity>
</figureCitation>
); a very deeply emarginate sternite
<number box="[1301,1315,330,354]" pageId="33" pageNumber="34" value="9.0">9</number>
was found in a male from Xiangkhouang (
<figureCitation box="[497,592,366,390]" captionStart="FIGURES 3335" captionStartId="9.[151,264,1686,1709]" captionTargetBox="[166,1425,193,1639]" captionTargetId="figure@9.[151,1436,193,1645]" captionText="FIGURES 3335. Male genitalia of Asian Pachysternum. 33: P. kubani (ac, f: holotype; de: dark specimen from Sichuan; g: specimen from Laos, Ban Nape; h: terratological specimen from Laos, Phou Pane Mt.; a: tegmen, b, d: median lobe, c, e: apical portion of median lobe, fh: sternite 9); 34: P. stevensi (a: tegmen, b-c: median lobe, de: variation of the shape of sternite 9); 35: P. sulawesicum (a: tegmen, b: median lobe, c: apical portion of median lobe, d: sternite 9)." pageId="33" pageNumber="34">
Fig.
<number box="[549,577,366,390]" pageId="33" pageNumber="34" value="33.0">33</number>
h
</figureCitation>
) which also has extremely shortened tibial spines (all remaining specimens from the locality have usually developed sternite
<number box="[700,714,402,426]" pageId="33" pageNumber="34" value="9.0">9</number>
as well as tibial spines, and the specimen is therefore considered as teratological here). This variation of the shape of the sternite
<number box="[876,890,438,462]" pageId="33" pageNumber="34" value="9.0">9</number>
resembles that observed in
<taxonomicName box="[1206,1323,439,462]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="stevensi">
<emphasis box="[1206,1323,439,462]" italics="true" pageId="33" pageNumber="34">P. stevensi</emphasis>
</taxonomicName>
, in which both rounded and emarginate forms are more frequent, however.
</paragraph>
</subSubSection>
<caption pageId="32" pageNumber="33" startId="32.[151,250,1042,1065]" targetBox="[210,1357,184,1017]" targetPageId="32">
<paragraph blockId="32.[151,1436,1042,1131]" pageId="32" pageNumber="33">
<emphasis bold="true" box="[151,287,1042,1065]" pageId="32" pageNumber="33">
FIGURE
<number box="[255,282,1043,1064]" pageId="32" pageNumber="33" value="60.0">60</number>
.
</emphasis>
<taxonomicName box="[293,512,1043,1065]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="kubani">
<emphasis box="[293,512,1043,1065]" italics="true" pageId="32" pageNumber="33">Pachysternum kubani</emphasis>
</taxonomicName>
, general habitus (a, ci: dorsal view, b: laterodorsal view). Localities of the specimens: ab:
<collectingCountry box="[151,203,1077,1098]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
, Phou Pane Mt.; c: same locality (
<typeStatus box="[556,649,1076,1098]" pageId="32" pageNumber="33">holotype</typeStatus>
, male); d:
<collectingCountry box="[759,811,1077,1098]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
, Phongsaly env. (female); e: same locality (male); fh:
<collectingCountry box="[1380,1432,1077,1098]" name="Laos" pageId="32" pageNumber="33">Laos</collectingCountry>
, Ban Nape; i:
<collectingCountry box="[285,348,1109,1131]" name="China" pageId="32" pageNumber="33">China</collectingCountry>
, Sichuan, Qingcheng Shan.
</paragraph>
</caption>
<caption box="[151,751,2012,2035]" pageId="32" pageNumber="33" startId="32.[151,250,2012,2035]" targetBox="[277,1305,1171,2005]" targetPageId="32">
<paragraph blockId="32.[151,751,2012,2035]" box="[151,751,2012,2035]" pageId="32" pageNumber="33">
<emphasis bold="true" box="[151,287,2012,2035]" pageId="32" pageNumber="33">FIGURE 61.</emphasis>
Known distribution of
<taxonomicName box="[525,745,2012,2034]" class="Insecta" family="Hydrophilidae" genus="Pachysternum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="kubani">
<emphasis box="[525,745,2012,2034]" italics="true" pageId="32" pageNumber="33">Pachysternum kubani</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection pageId="33" pageNumber="34" type="etymology">
<paragraph blockId="33.[151,1436,151,711]" pageId="33" pageNumber="34">
<emphasis bold="true" box="[199,334,509,533]" pageId="33" pageNumber="34">Etymology.</emphasis>
The species is dedicated to my colleague Vítĕzslav Kubáň (
<collectionCode box="[1002,1085,509,533]" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="33" pageNumber="34">NMPC</collectionCode>
) who collected most
<typeStatus box="[1322,1370,510,534]" pageId="33" pageNumber="34">type</typeStatus>
specimens of this species.
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="34" type="biology_ecology">
<paragraph blockId="33.[151,1436,151,711]" pageId="33" pageNumber="34">
<emphasis bold="true" box="[199,295,580,605]" pageId="33" pageNumber="34">Biology.</emphasis>
Unknown. Most specimens were collected in primary or secondary mountain forests using flight intercept traps.
</paragraph>
</subSubSection>
<subSubSection pageId="33" pageNumber="34" type="distribution">
<paragraph blockId="33.[151,1436,151,711]" pageId="33" pageNumber="34">
<emphasis bold="true" box="[199,349,652,677]" pageId="33" pageNumber="34">Distribution.</emphasis>
Known from four localities in northern
<collectingCountry box="[806,859,653,676]" name="Laos" pageId="33" pageNumber="34">Laos</collectingCountry>
(provinces Bolikhamsai, Houaphan, Phongsali and Xiangkhouang) and from central
<collectingCountry box="[524,591,687,711]" name="China" pageId="33" pageNumber="34">China</collectingCountry>
(Sichuan).
</paragraph>
</subSubSection>
</treatment>
</document>