231 lines
39 KiB
XML
231 lines
39 KiB
XML
<document id="035D0C142309F9D0ED878681E47DE582" ID-DOI="10.11646/zootaxa.5039.3.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5511544" ID-ZooBank="2F98783C-14F7-46FF-9DB6-69E343F1DA37" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1631776248578" checkinUser="plazi" docAuthor="Gonçalves, Clayton Corrêa & Barbosa, Julianna Freires" docDate="2021" docId="03E6879EFFFFFFC6FF1DD286FB2C3FB0" docLanguage="en" docName="zootaxa.5039.3.6.pdf" docOrigin="Zootaxa 5039 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tichocoelidia marceloi Gonçalves & Barbosa 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="419" masterDocId="FFDFFFE6FFFDFFC4FF8AD314FFAC3928" masterDocTitle="Two new Peruvian species of Tichocoelidia Kramer (Hemiptera: Cicadellidae: Neocoelidiinae) with a key to males" masterLastPageNumber="424" masterPageNumber="417" pageNumber="419" updateTime="1699414663462" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:titleInfo id="B77D28B297F46B8851FE73D18FA33183">
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<mods:title id="718CE9DB1D479AFE9285162740BA9BE3">Two new Peruvian species of Tichocoelidia Kramer (Hemiptera: Cicadellidae: Neocoelidiinae) with a key to males</mods:title>
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<mods:name id="30A504E3825A5274C74F868342371F64" type="personal">
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<mods:roleTerm id="783FDF14E8F20E9251C671063977C584">Author</mods:roleTerm>
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<mods:namePart id="91A8A7D065414A1BD044DBB5530B4C51">Gonçalves, Clayton Corrêa</mods:namePart>
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<mods:nameIdentifier id="CF1E3194EC3DE60A66A414526FECAB1A" type="ORCID">0000-0003-3045-3425</mods:nameIdentifier>
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<mods:affiliation id="1072C3E174BA425D67016E997DA9D6FA">Laboratório de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ, Brasil. & clayton. correa. goncalves @ gmail. com; https: // orcid. org / 0000 - 0003 - 3045 - 3425</mods:affiliation>
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<mods:nameIdentifier id="8AC5025683515833981BD7070AD0C26C" type="email">clayton.correa.goncalves@gmail.com</mods:nameIdentifier>
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<mods:namePart id="5CCB089294839235E504FE4E0FEA926E">Barbosa, Julianna Freires</mods:namePart>
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<mods:nameIdentifier id="E3ADD236D29EC8B876B4142E10B36ACF" type="ORCID">0000-0002-5276-194X</mods:nameIdentifier>
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<mods:affiliation id="4F5978A34FF1A642EAF254A92514443B">Laboratório de Entomologia, Departamento de Zoologia, Instituto de Biologia, Universidade Federal do Rio de Janeiro, Caixa Postal 68044, Rio de Janeiro, 21941 - 971, RJ, Brasil. & julianna. freires @ gmail. com; https: // orcid. org / 0000 - 0002 - 5276 - 194 X</mods:affiliation>
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<mods:nameIdentifier id="2E2E83E685215FC93AFD74F66A008905" type="email">julianna.freires@gmail.com</mods:nameIdentifier>
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<mods:title id="0E08C181C957D1E81FFA7B99EDBA3597">Zootaxa</mods:title>
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<mods:part id="F67BBCEB7344D46DFD352A20650A9454">
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<mods:date id="3ADCC0204662E98859D957B82660F6E1">2021</mods:date>
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<mods:number id="844ADB7F4F8060F8CCCF6344FA31818A">2021-09-16</mods:number>
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<mods:number id="000E088A12A11F7DF3971FC54B6A4F1D">5039</mods:number>
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<mods:identifier id="73CF47E5537F391356B12A4CA601A2DC" type="DOI">10.11646/zootaxa.5039.3.6</mods:identifier>
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<treatment id="03E6879EFFFFFFC6FF1DD286FB2C3FB0" ID-DOI="http://doi.org/10.5281/zenodo.5511550" ID-Zenodo-Dep="5511550" LSID="urn:lsid:plazi:treatment:03E6879EFFFFFFC6FF1DD286FB2C3FB0" httpUri="http://treatment.plazi.org/id/03E6879EFFFFFFC6FF1DD286FB2C3FB0" lastPageNumber="419" pageId="2" pageNumber="419">
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<subSubSection id="C3556503FFFFFFC6FF1DD286FDBD3884" box="[151,529,402,429]" pageId="2" pageNumber="419" type="nomenclature">
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<paragraph id="8BF03688FFFFFFC6FF1DD286FDBD3884" blockId="2.[151,529,402,464]" box="[151,529,402,429]" pageId="2" pageNumber="419">
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<heading id="D0B881E4FFFFFFC6FF1DD286FDBD3884" bold="true" box="[151,529,402,429]" fontSize="11" level="1" pageId="2" pageNumber="419" reason="1">
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<emphasis id="B93BEA9AFFFFFFC6FF1DD286FDBD3884" bold="true" box="[151,529,402,429]" pageId="2" pageNumber="419">
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<taxonomicName id="4C4F4D0BFFFFFFC6FF1DD286FE003885" ID-CoL="8RVJ4" authority="Gonçalves & Barbosa, 2021" authorityName="Gonçalves & Barbosa" authorityYear="2021" box="[151,428,402,429]" class="Insecta" family="Cicadellidae" genus="Tichocoelidia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="419" phylum="Arthropoda" rank="species" species="marceloi" status="sp. nov.">
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<emphasis id="B93BEA9AFFFFFFC6FF1DD286FE003885" bold="true" box="[151,428,402,429]" italics="true" pageId="2" pageNumber="419">Tichocoelidia marceloi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20857E1FFFFFFC6FE39D286FDBD3884" box="[435,529,402,428]" pageId="2" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556503FFFFFFC6FF1DD2A2FEC138F8" box="[151,365,438,464]" pageId="2" pageNumber="419" type="description">
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<paragraph id="8BF03688FFFFFFC6FF1DD2A2FEC138F8" blockId="2.[151,529,402,464]" box="[151,365,438,464]" pageId="2" pageNumber="419">
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||
(
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||
<figureCitation id="13742A0DFFFFFFC6FF15D2A2FEBD38F8" box="[159,273,438,464]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Figs 1–10</figureCitation>
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,
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<figureCitation id="13742A0DFFFFFFC6FE94D2A2FEC938F8" box="[286,357,438,464]" captionStart="FIGURES 21–28" captionStartId="5.[151,264,1926,1950]" captionTargetBox="[162,1426,529,1903]" captionTargetId="figure-151@5.[162,1426,525,1903]" captionTargetPageId="5" captionText="FIGURES 21–28. Habitus, dorsal and lateral views, respectively. 21 and 22, Tichocoelidia marceloi sp. nov., holotype male. 23 and 24, T. vanessae sp. nov., holotype male. 25 and 26, T. clarkei Kramer, 1962, holotype male (USNM). 27 and 28, T. grandis Chiamolera & Cavichioli, 2003, holotype male (INPA)." figureDoi="http://doi.org/10.5281/zenodo.5511554" httpUri="https://zenodo.org/record/5511554/files/figure.png" pageId="2" pageNumber="419">21–22</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556503FFFFFFC6FF1DD2EAFD693BE4" pageId="2" pageNumber="419" type="diagnosis">
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<paragraph id="8BF03688FFFFFFC6FF1DD2EAFD693BE4" blockId="2.[151,1437,510,1689]" pageId="2" pageNumber="419">
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<emphasis id="B93BEA9AFFFFFFC6FF1DD2EAFEBE3B30" bold="true" box="[151,274,510,536]" pageId="2" pageNumber="419">Diagnosis.</emphasis>
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Crown (
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<figureCitation id="13742A0DFFFFFFC6FEF2D2EAFE673B30" box="[376,459,510,536]" captionStart="FIGURES 21–28" captionStartId="5.[151,264,1926,1950]" captionTargetBox="[162,1426,529,1903]" captionTargetId="figure-151@5.[162,1426,525,1903]" captionTargetPageId="5" captionText="FIGURES 21–28. Habitus, dorsal and lateral views, respectively. 21 and 22, Tichocoelidia marceloi sp. nov., holotype male. 23 and 24, T. vanessae sp. nov., holotype male. 25 and 26, T. clarkei Kramer, 1962, holotype male (USNM). 27 and 28, T. grandis Chiamolera & Cavichioli, 2003, holotype male (INPA)." figureDoi="http://doi.org/10.5281/zenodo.5511554" httpUri="https://zenodo.org/record/5511554/files/figure.png" pageId="2" pageNumber="419">Fig. 21</figureCitation>
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) with rounded anterior margin. Pygofer (
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<figureCitation id="13742A0DFFFFFFC6FC2ED2EAFC443B30" box="[932,1000,510,536]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 3</figureCitation>
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) with inner dorsal processes ventrally curved in apical third, not exceeding apex of pygofer. Subgenital plates, in lateral view (
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<figureCitation id="13742A0DFFFFFFC6FBFCD136FB153B14" box="[1142,1209,546,572]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 4</figureCitation>
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) with dorsal margin expanded in median third; in ventral view (
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<figureCitation id="13742A0DFFFFFFC6FDF5D152FD6F3B49" box="[639,707,582,609]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 5</figureCitation>
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), with lateral margins converging towards the apex, narrowed in apical third. Aedeagus (
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<figureCitation id="13742A0DFFFFFFC6FE28D17EFDAB3BAC" box="[418,519,618,644]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Figs 8–9</figureCitation>
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) with shaft asymmetrical and strongly flattened dorsoventrally, gonoduct distinct throughout length of shaft, apex slightly expanded. Anal tube (
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<figureCitation id="13742A0DFFFFFFC6FCD1D19AFC633B80" box="[859,975,654,680]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Figs 3, 10</figureCitation>
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) with short processes, with apices divergent and acute, outer margin with small denticles.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3556503FFFFFFC6FF4DD1C2FDD53C7C" pageId="2" pageNumber="419" type="description">
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<paragraph id="8BF03688FFFFFFC6FF4DD1C2FC903BD9" blockId="2.[151,1437,510,1689]" box="[199,828,726,753]" pageId="2" pageNumber="419">
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<emphasis id="B93BEA9AFFFFFFC6FF4DD1C2FE693BD8" bold="true" box="[199,453,726,752]" pageId="2" pageNumber="419">Measurements (mm).</emphasis>
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Male
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<typeStatus id="54F4882AFFFFFFC6FD87D1C2FDDF3BD8" box="[525,627,726,752]" pageId="2" pageNumber="419" type="holotype">holotype</typeStatus>
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: total length 7.25.
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</paragraph>
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<paragraph id="8BF03688FFFFFFC6FF4DD1EEFCD03A74" blockId="2.[151,1437,510,1689]" pageId="2" pageNumber="419">
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<emphasis id="B93BEA9AFFFFFFC6FF4DD1EEFEE33A3C" bold="true" box="[199,335,762,788]" pageId="2" pageNumber="419">Coloration.</emphasis>
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Body yellow (
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<figureCitation id="13742A0DFFFFFFC6FE70D1EEFDD33A3C" box="[506,639,762,788]" captionStart="FIGURES 21–28" captionStartId="5.[151,264,1926,1950]" captionTargetBox="[162,1426,529,1903]" captionTargetId="figure-151@5.[162,1426,525,1903]" captionTargetPageId="5" captionText="FIGURES 21–28. Habitus, dorsal and lateral views, respectively. 21 and 22, Tichocoelidia marceloi sp. nov., holotype male. 23 and 24, T. vanessae sp. nov., holotype male. 25 and 26, T. clarkei Kramer, 1962, holotype male (USNM). 27 and 28, T. grandis Chiamolera & Cavichioli, 2003, holotype male (INPA)." figureDoi="http://doi.org/10.5281/zenodo.5511554" httpUri="https://zenodo.org/record/5511554/files/figure.png" pageId="2" pageNumber="419">Figs 21–22</figureCitation>
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). Crown yellow, without orange stripes. Face (
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<figureCitation id="13742A0DFFFFFFC6FB1FD1EEFB7B3A3C" box="[1173,1239,762,788]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 1</figureCitation>
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) without distinct marks. Pronotum (
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<figureCitation id="13742A0DFFFFFFC6FEECD00AFE1B3A10" box="[358,439,798,824]" captionStart="FIGURES 21–28" captionStartId="5.[151,264,1926,1950]" captionTargetBox="[162,1426,529,1903]" captionTargetId="figure-151@5.[162,1426,525,1903]" captionTargetPageId="5" captionText="FIGURES 21–28. Habitus, dorsal and lateral views, respectively. 21 and 22, Tichocoelidia marceloi sp. nov., holotype male. 23 and 24, T. vanessae sp. nov., holotype male. 25 and 26, T. clarkei Kramer, 1962, holotype male (USNM). 27 and 28, T. grandis Chiamolera & Cavichioli, 2003, holotype male (INPA)." figureDoi="http://doi.org/10.5281/zenodo.5511554" httpUri="https://zenodo.org/record/5511554/files/figure.png" pageId="2" pageNumber="419">Fig. 21</figureCitation>
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) with two dark maculae adjacent to posterior margin. Forewing (
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<figureCitation id="13742A0DFFFFFFC6FB02D00AFB673A10" box="[1160,1227,798,824]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 2</figureCitation>
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) yellow semi-hyaline; anal margin and apical portion smoky. Legs (
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<figureCitation id="13742A0DFFFFFFC6FD43D056FCB73A74" box="[713,795,834,860]" captionStart="FIGURES 21–28" captionStartId="5.[151,264,1926,1950]" captionTargetBox="[162,1426,529,1903]" captionTargetId="figure-151@5.[162,1426,525,1903]" captionTargetPageId="5" captionText="FIGURES 21–28. Habitus, dorsal and lateral views, respectively. 21 and 22, Tichocoelidia marceloi sp. nov., holotype male. 23 and 24, T. vanessae sp. nov., holotype male. 25 and 26, T. clarkei Kramer, 1962, holotype male (USNM). 27 and 28, T. grandis Chiamolera & Cavichioli, 2003, holotype male (INPA)." figureDoi="http://doi.org/10.5281/zenodo.5511554" httpUri="https://zenodo.org/record/5511554/files/figure.png" pageId="2" pageNumber="419">Fig. 22</figureCitation>
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) yellow.
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</paragraph>
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<paragraph id="8BF03688FFFFFFC6FF4DD072FBA73A8D" blockId="2.[151,1437,510,1689]" pageId="2" pageNumber="419">
|
||
<emphasis id="B93BEA9AFFFFFFC6FF4DD072FE613AA8" bold="true" box="[199,461,870,896]" pageId="2" pageNumber="419">External morphology.</emphasis>
|
||
Crown (
|
||
<figureCitation id="13742A0DFFFFFFC6FDBAD072FD2E3AA8" box="[560,642,870,896]" captionStart="FIGURES 21–28" captionStartId="5.[151,264,1926,1950]" captionTargetBox="[162,1426,529,1903]" captionTargetId="figure-151@5.[162,1426,525,1903]" captionTargetPageId="5" captionText="FIGURES 21–28. Habitus, dorsal and lateral views, respectively. 21 and 22, Tichocoelidia marceloi sp. nov., holotype male. 23 and 24, T. vanessae sp. nov., holotype male. 25 and 26, T. clarkei Kramer, 1962, holotype male (USNM). 27 and 28, T. grandis Chiamolera & Cavichioli, 2003, holotype male (INPA)." figureDoi="http://doi.org/10.5281/zenodo.5511554" httpUri="https://zenodo.org/record/5511554/files/figure.png" pageId="2" pageNumber="419">Fig. 24</figureCitation>
|
||
) subquadrangular, with rounded anterior margin; coronal maculae distinct near posterior margin. Other characteristics as in description of the genus.
|
||
</paragraph>
|
||
<paragraph id="8BF03688FFFFFFC6FF4DD0BAFCE73C18" blockId="2.[151,1437,510,1689]" pageId="2" pageNumber="419">
|
||
<emphasis id="B93BEA9AFFFFFFC6FF4DD0BAFE3D3AE0" bold="true" box="[199,401,942,968]" pageId="2" pageNumber="419">Male terminalia.</emphasis>
|
||
Pygofer, in lateral view (
|
||
<figureCitation id="13742A0DFFFFFFC6FD36D0BAFCAF3AE0" box="[700,771,942,968]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 3</figureCitation>
|
||
), subtriangular, high at base and progressively narrowing towards apex, approximately twice as long as high; macrosetae present on apical third; ventral tooth on basal third, apex rounded; inner dorsal processes ventrally curved in apical third, not exceeding apex of pygofer, ventral margin with small irregular and rounded apical teeth. Subgenital plates, in lateral view (
|
||
<figureCitation id="13742A0DFFFFFFC6FB80D70EFBE03D1C" box="[1034,1100,1050,1076]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 4</figureCitation>
|
||
) with dorsal margin expanded in median third forming a rounded lobe; in ventral view (
|
||
<figureCitation id="13742A0DFFFFFFC6FCA7D72AFCDE3D71" box="[813,882,1086,1113]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 5</figureCitation>
|
||
), with lateral margins converging towards apex, narrowed in apical third; outer margin with microsetae in apical third. Connective (
|
||
<figureCitation id="13742A0DFFFFFFC6FBBAD776FBDF3D55" box="[1072,1139,1122,1149]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 6</figureCitation>
|
||
) approximately two-thirds of length of style. Style (
|
||
<figureCitation id="13742A0DFFFFFFC6FE23D792FDA13D89" box="[425,525,1158,1185]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Figs 6–7</figureCitation>
|
||
) with base of apodeme bilobed, both lobes rounded and approximately same size; apophysis moderately curved dorsally; apical hook moderately developed. Aedeagus (
|
||
<figureCitation id="13742A0DFFFFFFC6FBE2D7BEFB623DEC" box="[1128,1230,1194,1220]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Figs 8–9</figureCitation>
|
||
) with shaft asymmetrical and strongly flattened dorsoventrally, slightly twisted; apex slightly expanded, with two small preapical lateral lobes; gonoduct distinct throughout length of the shaft. Anal tube (
|
||
<figureCitation id="13742A0DFFFFFFC6FC63D7E6FBCC3C24" box="[1001,1120,1266,1292]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Figs 3, 10</figureCitation>
|
||
) with short processes, with apices divergent and acute, outer margin with small denticles.
|
||
</paragraph>
|
||
<paragraph id="8BF03688FFFFFFC6FF4DD62EFDD53C7C" blockId="2.[151,1437,510,1689]" box="[199,633,1338,1364]" pageId="2" pageNumber="419">
|
||
<emphasis id="B93BEA9AFFFFFFC6FF4DD62EFE053C7C" bold="true" box="[199,425,1338,1364]" pageId="2" pageNumber="419">Female terminalia.</emphasis>
|
||
Female unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556503FFFFFFC6FF4DD64AFA2A3CB4" pageId="2" pageNumber="419" type="materials_examined">
|
||
<paragraph id="8BF03688FFFFFFC6FF4DD64AFA2A3CB4" blockId="2.[151,1437,510,1689]" pageId="2" pageNumber="419">
|
||
<emphasis id="B93BEA9AFFFFFFC6FF4DD64AFE1C3C50" bold="true" box="[199,432,1374,1400]" pageId="2" pageNumber="419">Material examined.</emphasis>
|
||
<materialsCitation id="3B273CD5FFFFFFC6FE3DD64AFA2A3CB4" collectingDate="2012-08-23" collectingDateMax="2012-08-31" collectingDateMin="2012-08-23" collectionCode="AMP, DM, MUSM" country="Peru" elevation="874" latitude="-13.336111" location="Rio Araza" longLatPrecision="21" longitude="-70.84917" pageId="2" pageNumber="419" specimenCount="1" stateProvince="Cusco" typeStatus="holotype">
|
||
<emphasis id="B93BEA9AFFFFFFC6FE3DD64AFDC13C50" bold="true" box="[439,621,1374,1400]" pageId="2" pageNumber="419">
|
||
<typeStatus id="54F4882AFFFFFFC6FE3DD64AFD883C50" box="[439,548,1374,1400]" pageId="2" pageNumber="419" type="holotype">Holotype</typeStatus>
|
||
male:
|
||
</emphasis>
|
||
“
|
||
<collectingCountry id="F3587618FFFFFFC6FDF4D64AFD1B3C50" box="[638,695,1374,1400]" name="Peru" pageId="2" pageNumber="419">Peru</collectingCountry>
|
||
,
|
||
<collectingRegion id="498BF86AFFFFFFC6FD4BD64AFCA63C50" box="[705,778,1374,1400]" country="Peru" name="Cusco" pageId="2" pageNumber="419">Cusco</collectingRegion>
|
||
, 19 rd Km W Quincemil,\
|
||
<location id="8E906053FFFFFFC6FBB8D64AFB0E3C50" LSID="urn:lsid:plazi:treatment:03E6879EFFFFFFC6FF1DD286FB2C3FB0:8E906053FFFFFFC6FBB8D64AFB0E3C50" box="[1074,1186,1374,1400]" country="Peru" latitude="-13.336111" longLatPrecision="21" longitude="-70.84917" name="Rio Araza" pageId="2" pageNumber="419" stateProvince="Cusco">Rio Araza</location>
|
||
tributary\
|
||
<geoCoordinate id="EE7B504FFFFFFFC6FA92D64AFA313C50" box="[1304,1437,1374,1400]" degrees="13" direction="south" minutes="20" orientation="latitude" pageId="2" pageNumber="419" precision="15" seconds="10" value="-13.336111">13°20’10”S</geoCoordinate>
|
||
<geoCoordinate id="EE7B504FFFFFFFC6FF1DD696FE8B3CB5" box="[151,295,1410,1437]" degrees="70" direction="west" minutes="50" orientation="longitude" pageId="2" pageNumber="419" precision="15" seconds="57" value="-70.84917">70°50’57”W</geoCoordinate>
|
||
,
|
||
<quantity id="4CB79B6DFFFFFFC6FEB8D696FEDD3CB4" box="[306,369,1410,1436]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="2" pageNumber="419" unit="m" value="874.0">
|
||
<elevation id="0062D1BBFFFFFFC6FEB8D696FEDD3CB4" box="[306,369,1410,1436]" metricMagnitude="2" metricUnit="m" metricValue="8.74" pageId="2" pageNumber="419" unit="m" value="874.0">874m</elevation>
|
||
</quantity>
|
||
\
|
||
<date id="FFF11048FFFFFFC6FE0BD696FD9C3CB4" box="[385,560,1410,1436]" pageId="2" pageNumber="419" value="2012-08-23" valueMax="2012-08-31" valueMin="2012-08-23">
|
||
<collectingDate id="EFB5E9A0FFFFFFC6FE0BD696FD9C3CB4" box="[385,560,1410,1436]" pageId="2" pageNumber="419" value="2012-08-23" valueMax="2012-08-31" valueMin="2012-08-23">23-31.viii.2012</collectingDate>
|
||
</date>
|
||
, Malaise, RR Cavichioli,\ JA Rafael,
|
||
<collectionCode id="ED5EAE4DFFFFFFC6FC59D696FBA33CB4" box="[979,1039,1410,1436]" country="Australia" httpUri="http://grbio.org/cool/90v0-7gw3" name="Australian Mycological Panel" pageId="2" pageNumber="419">AMP</collectionCode>
|
||
Santos &
|
||
<collectionCode id="ED5EAE4DFFFFFFC6FB0ED697FB1D3CB5" box="[1156,1201,1411,1437]" country="New Zealand" name="Dominion Museum" pageId="2" pageNumber="419">DM</collectionCode>
|
||
Takiya” (
|
||
<collectionCode id="ED5EAE4DFFFFFFC6FA95D696FAD73CB4" box="[1311,1403,1410,1436]" pageId="2" pageNumber="419">MUSM</collectionCode>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556503FFFFFFC6FF4DD6B2FBA53CCD" pageId="2" pageNumber="419" type="etymology">
|
||
<paragraph id="8BF03688FFFFFFC6FF4DD6B2FBA53CCD" blockId="2.[151,1437,510,1689]" pageId="2" pageNumber="419">
|
||
<emphasis id="B93BEA9AFFFFFFC6FF4DD6B2FEE23CE8" bold="true" box="[199,334,1446,1472]" pageId="2" pageNumber="419">Etymology.</emphasis>
|
||
The species name is a tribute to Marcelo Peixoto Gomes da Silva, a graduate student who started his studies on Neotropical leafhoppers advised by the first author of this paper.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3556503FFFFFFC6FF4DD6FAFB2C3FB0" pageId="2" pageNumber="419" type="discussion">
|
||
<paragraph id="8BF03688FFFFFFC6FF4DD6FAFB2C3FB0" blockId="2.[151,1437,510,1689]" pageId="2" pageNumber="419">
|
||
<emphasis id="B93BEA9AFFFFFFC6FF4DD6FAFE973F20" bold="true" box="[199,315,1518,1544]" pageId="2" pageNumber="419">Remarks.</emphasis>
|
||
<taxonomicName id="4C4F4D0BFFFFFFC6FECBD6FAFDEF3F20" authority="Gonçalves & Barbosa, 2021" authorityName="Gonçalves & Barbosa" authorityYear="2021" box="[321,579,1518,1544]" class="Insecta" family="Cicadellidae" genus="Tichocoelidia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="419" phylum="Arthropoda" rank="species" species="marceloi" status="sp. nov.">
|
||
<emphasis id="B93BEA9AFFFFFFC6FECBD6FAFDEF3F20" box="[321,579,1518,1544]" italics="true" pageId="2" pageNumber="419">Tichocoelidia marceloi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93BEA9AFFFFFFC6FDC2D6FAFD0D3F20" bold="true" box="[584,673,1518,1544]" pageId="2" pageNumber="419">
|
||
<taxonomicNameLabel id="A20857E1FFFFFFC6FDC2D6FAFD0D3F20" box="[584,673,1518,1544]" pageId="2" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="4C4F4D0BFFFFFFC6FC97D6FBFC293F20" authorityName="Kramer" authorityYear="1962" box="[797,901,1518,1544]" class="Insecta" family="Cicadellidae" genus="Tichocoelidia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="419" phylum="Arthropoda" rank="species" species="clarkei">
|
||
<emphasis id="B93BEA9AFFFFFFC6FC97D6FBFC293F20" box="[797,901,1518,1544]" italics="true" pageId="2" pageNumber="419">T. clarkei</emphasis>
|
||
</taxonomicName>
|
||
in the color pattern (
|
||
<figureCitation id="13742A0DFFFFFFC6FBE0D6FAFAEC3F21" box="[1130,1344,1518,1545]" captionStart="FIGURES 21–28" captionStartId="5.[151,264,1926,1950]" captionTargetBox="[162,1426,529,1903]" captionTargetId="figure-151@5.[162,1426,525,1903]" captionTargetPageId="5" captionText="FIGURES 21–28. Habitus, dorsal and lateral views, respectively. 21 and 22, Tichocoelidia marceloi sp. nov., holotype male. 23 and 24, T. vanessae sp. nov., holotype male. 25 and 26, T. clarkei Kramer, 1962, holotype male (USNM). 27 and 28, T. grandis Chiamolera & Cavichioli, 2003, holotype male (INPA)." figureDoi="http://doi.org/10.5281/zenodo.5511554" httpUri="https://zenodo.org/record/5511554/files/figure.png" pageId="2" pageNumber="419">Figs 21–22, 25–26</figureCitation>
|
||
), shapes of aedeagus and processes of anal tube. However,
|
||
<taxonomicName id="4C4F4D0BFFFFFFC6FD49D507FCEE3F04" authority="Gonçalves & Barbosa, 2021" authorityName="Gonçalves & Barbosa" authorityYear="2021" box="[707,834,1554,1581]" class="Insecta" family="Cicadellidae" genus="Tichocoelidia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="419" phylum="Arthropoda" rank="species" species="marceloi" status="sp. nov.">
|
||
<emphasis id="B93BEA9AFFFFFFC6FD49D507FD7F3F04" box="[707,723,1555,1580]" italics="true" pageId="2" pageNumber="419">T</emphasis>
|
||
.
|
||
<emphasis id="B93BEA9AFFFFFFC6FD55D506FCEE3F04" box="[735,834,1554,1580]" italics="true" pageId="2" pageNumber="419">marceloi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B93BEA9AFFFFFFC6FCC2D506FC0C3F04" bold="true" box="[840,928,1554,1580]" pageId="2" pageNumber="419">
|
||
<taxonomicNameLabel id="A20857E1FFFFFFC6FCC2D506FC0C3F04" box="[840,928,1554,1580]" pageId="2" pageNumber="419" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C4F4D0BFFFFFFC6FBBBD507FB303F04" authorityName="Kramer" authorityYear="1962" box="[1073,1180,1554,1581]" class="Insecta" family="Cicadellidae" genus="Tichocoelidia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="419" phylum="Arthropoda" rank="species" species="clarkei">
|
||
<emphasis id="B93BEA9AFFFFFFC6FBBBD507FBED3F04" box="[1073,1089,1555,1580]" italics="true" pageId="2" pageNumber="419">T</emphasis>
|
||
.
|
||
<emphasis id="B93BEA9AFFFFFFC6FBC4D506FB303F04" box="[1102,1180,1554,1580]" italics="true" pageId="2" pageNumber="419">clarkei</emphasis>
|
||
</taxonomicName>
|
||
by the shape of the pygofer. In the new species the ventral tooth arises on the basal third of the ventral margin and the inner processes are thin and curved ventrally (
|
||
<figureCitation id="13742A0DFFFFFFC6FE37D54EFDAC3F5C" box="[445,512,1626,1652]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Fig. 3</figureCitation>
|
||
), while in
|
||
<taxonomicName id="4C4F4D0BFFFFFFC6FDFDD54FFD4E3F5C" authorityName="Kramer" authorityYear="1962" box="[631,738,1626,1653]" class="Insecta" family="Cicadellidae" genus="Tichocoelidia" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="419" phylum="Arthropoda" rank="species" species="clarkei">
|
||
<emphasis id="B93BEA9AFFFFFFC6FDFDD54FFD2B3F5C" box="[631,647,1627,1652]" italics="true" pageId="2" pageNumber="419">T</emphasis>
|
||
.
|
||
<emphasis id="B93BEA9AFFFFFFC6FD1ED54EFD4E3F5C" box="[660,738,1626,1652]" italics="true" pageId="2" pageNumber="419">clarkei</emphasis>
|
||
</taxonomicName>
|
||
the ventral tooth is on the middle third and the inner processes are robust and strongly curved internally with crossed apices (
|
||
<bibRefCitation id="EFDE4B79FFFFFFC6FCC7D56AFC4B3FB0" author="Kramer, J. P." box="[845,999,1662,1688]" pageId="2" pageNumber="419" pagination="101 - 105" refId="ref4303" refString="Kramer, J. P. (1962) A synopsis of Biza and a new allied genus (Homoptera: Cicadellidae: Neocoelidiinae). Proceedings of the Biological Society of Washington, 75, 101 - 105." type="journal article" year="1962">Kramer 1962</bibRefCitation>
|
||
;
|
||
<figureCitation id="13742A0DFFFFFFC6FC78D56AFBE43FB0" box="[1010,1096,1662,1688]" captionStart="FIGURES 1–10" captionStartId="3.[151,264,1860,1884]" captionTargetBox="[159,1414,179,1793]" captionTargetId="figure-19@3.[151,1435,179,1803]" captionTargetPageId="3" captionText="FIGURES 1–10. Tichocoelidia marceloi sp. nov., holotype male: 1, head, frontal view; 2, forewing; 3, pygofer and anal tube, lateral view; 4, valve and subgenital plates, lateral view; 5, valve and subgenital plates, ventral view; 6, connective and styles, dorsal view; 7, style, lateral view; 8, aedeagus, lateral view; 9, aedeagus, caudal view; 10 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511548" httpUri="https://zenodo.org/record/5511548/files/figure.png" pageId="2" pageNumber="419">Figs 10</figureCitation>
|
||
,
|
||
<figureCitation id="13742A0DFFFFFFC6FBDED56AFBDF3FB0" box="[1108,1139,1662,1688]" captionStart="FIGURES 11–20" captionStartId="4.[151,264,1830,1854]" captionTargetBox="[154,1418,181,1793]" captionTargetId="figure-21@4.[151,1435,181,1805]" captionTargetPageId="4" captionText="FIGURES 11–20. Tichocoelidia vanessae sp. nov., holotype male: 11, head, frontal view; 12, forewing; 13, pygofer and anal tube, lateral view; 14, valve and subgenital plates, lateral view; 15, valve and subgenital plates, ventral view; 16, connective and style, dorsal view; 17, style, lateral view; 18, aedeagus, lateral view; 19, aedeagus, caudal view; 20 processes of anal tube, caudal view." figureDoi="http://doi.org/10.5281/zenodo.5511552" httpUri="https://zenodo.org/record/5511552/files/figure.png" pageId="2" pageNumber="419">12</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |