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<document id="6E1A71C3908896E405105CEE11D59842" ID-CLB-Dataset="55466" ID-DOI="10.5281/zenodo.5169525" ID-GBIF-Dataset="4910761a-706e-4f53-a5a8-6c72379bdbc6" ID-ISSN="1942-1354" ID-Zenodo-Dep="5169525" ID-ZooBank="DA6CBFE5-927E-45B6-9D05-69AC97AF7B76" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628277966057" checkinUser="felipe" docAuthor="Paulsen, M. J." docDate="2017" docId="03DA133F7518B1384BC2F92F8EB36B2E" docLanguage="en" docName="InsectaMundi.0592.1-16.pdf" docOrigin="Insecta Mundi 2017 (592)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Ganelius madagascariensis" docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="FFE36B47751BB13D4B02FFA38A0B6E57" masterDocTitle="Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="2" updateTime="1699056540890" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="897867D732B33C173365B11C49470E04">Revision of the endemic Madagascan stag beetle genus Ganelius Benesh, and description of a new, related genus (Coleoptera: Lucanidae: Lucaninae: Figulini)</mods:title>
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<mods:namePart id="3F415B059A25C524900B40B8E831F1A5">Paulsen, M. J.</mods:namePart>
<mods:affiliation id="A1C02E5B9DA165460B68D4875FFAC0BC">Systematic Research Collections University of Nebraska State Museum W 436 Nebraska Hall Lincoln, NE 68588 - 0546</mods:affiliation>
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<mods:title id="05CCE9A7D2E5188BDA1326D51B312F10">Insecta Mundi</mods:title>
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<mods:part id="C82D4C3AA30F46A433D11F430FB6ABA8">
<mods:date id="6C9DC53A7F8DA99F1A32E0FC332B354D">2017</mods:date>
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<mods:number id="CF04B2AC4C2A071D703B2153875CF547">2017-12-29</mods:number>
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<mods:number id="9FC4CACA6C1CAB959167F78ACB5F4692">2017</mods:number>
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<treatment id="03DA133F7518B1384BC2F92F8EB36B2E" ID-DOI="http://doi.org/10.5281/zenodo.6567172" ID-GBIF-Taxon="195537735" ID-Zenodo-Dep="6567172" LSID="urn:lsid:plazi:treatment:03DA133F7518B1384BC2F92F8EB36B2E" httpUri="http://treatment.plazi.org/id/03DA133F7518B1384BC2F92F8EB36B2E" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
<subSubSection id="C369F1A27518B13E4BC2F92F88C368F3" box="[192,712,1676,1700]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="8BCCA2297518B13E4BC2F92F88C368F3" blockId="3.[192,712,1676,1700]" box="[192,712,1676,1700]" pageId="3" pageNumber="2">
<heading id="D08415457518B13E4BC2F92F88C368F3" bold="true" box="[192,712,1676,1700]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<taxonomicName id="4C73D9AA7518B13E4BC2F92F88C368F3" ID-CoL="3F8ZS" authority="(Laporte)" baseAuthorityName="Laporte" box="[192,712,1676,1700]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B7518B13E4BC2F92F88C368F3" bold="true" box="[192,712,1676,1700]" pageId="3" pageNumber="2">
<emphasis id="B9077E3B7518B13E4BC2F92F883668F3" bold="true" box="[192,573,1676,1700]" italics="true" pageId="3" pageNumber="2">Ganelius madagascariensis</emphasis>
(Laporte)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C369F1A27518B13E4BC2F96E8E0C68B2" box="[192,1031,1741,1765]" pageId="3" pageNumber="2" type="reference_group">
<paragraph id="8BCCA2297518B13E4BC2F96E8E0C68B2" blockId="3.[192,1031,1741,1765]" box="[192,1031,1741,1765]" pageId="3" pageNumber="2">
<treatmentCitationGroup id="AB6385077518B13E4BC2F96E8E0C68B2" box="[192,1031,1741,1765]" pageId="3" pageNumber="2">
<treatmentCitation id="0AD284387518B13E4BC2F96E88ED68B2" author="Laporte, F." box="[192,742,1741,1765]" page="175" pageId="3" pageNumber="2" year="1840">
<taxonomicName id="4C73D9AA7518B13E4BC2F96E88ED68B2" ID-CoL="3C2M7" authority="Laporte 1840: 175" authorityName="Laporte" authorityPageNumber="175" authorityYear="1840" box="[192,742,1741,1765]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B7518B13E4BC2F96E880A68B2" box="[192,513,1741,1765]" italics="true" pageId="3" pageNumber="2">Eudora madagascariensis</emphasis>
<bibRefCitation id="EFE2DFD87518B13E490AF96E88ED68B2" author="Laporte, F." box="[520,742,1741,1765]" pageId="3" pageNumber="2" pagination="1 - 564" refId="ref7518" refString="Laporte, F. 1840. Histoire Naturelle des Insectes Coleopteres. Avec une introduction renfermant l'Anatomie et la Physiologie des Animaux Articules, par M. Brulle. P. Dumenil; Paris 2: 1 - 564." type="journal article" year="1840">Laporte 1840: 175</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original combination).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C369F1A27518B1384BC2F8AE8EBB6CF1" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="8BCCA2297518B1384BC2F8AE88C56F0A" blockId="3.[191,1440,1805,1926]" lastBlockId="5.[191,1441,194,349]" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
<materialsCitation id="3B1BA8747518B1384BC2F8AE8F976E8D" ID-GBIF-Occurrence="3776282318" collectionCode="MNHN" country="Madagascar" determinerName="M. J. Paulsen" lastPageId="5" lastPageNumber="4" location="Madagascar" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" typeStatus="lectotype">
<emphasis id="B9077E3B7518B13E4BC2F8AE8B846972" bold="true" box="[192,399,1805,1829]" pageId="3" pageNumber="2">Type material.</emphasis>
<typeStatus id="54C81C8B7518B13E4A98F8AD88186971" box="[410,531,1806,1830]" pageId="3" pageNumber="2" type="lectotype">Lectotype</typeStatus>
male of
<taxonomicName id="4C73D9AA7518B13E4985F8AE89C66972" authorityName="Laporte" authorityYear="1840" box="[647,973,1805,1829]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B7518B13E4985F8AE89C66972" box="[647,973,1805,1829]" italics="true" pageId="3" pageNumber="2">Eudora madagascariensis</emphasis>
</taxonomicName>
(
<figureCitation id="1348BEAC7518B13E48E3F8AE8E216972" box="[993,1066,1805,1829]" captionStart="Figures 14" captionStartId="4.[192,288,1875,1897]" captionTargetBox="[192,1407,299,1723]" captionTargetId="figure-48@4.[192,1440,299,1747]" captionTargetPageId="4" captionText="Figures 14. Ganelius madagascariensis (Laporte, 1840). 1) Male lectotype, dorsal habitus. 2) Female, dorsal habitus. 3) Lectotype labels. 4) Male genitalia, flagellum ~ 60 mm." figureDoi="http://doi.org/10.5281/zenodo.5169527" httpUri="https://zenodo.org/record/5169527/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
),
<collectionCode id="ED623AEC7518B13E4F4AF8AD8EAD6972" box="[1096,1190,1806,1829]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="3" pageNumber="2">MNHN</collectionCode>
,
<emphasis id="B9077E3B7518B13E4FB7F8AE8F936972" bold="true" box="[1205,1432,1805,1829]" pageId="3" pageNumber="2">here designated</emphasis>
, labeled (
<figureCitation id="1348BEAC7518B13E4A2AF88E8B656912" box="[296,366,1837,1861]" captionStart="Figures 14" captionStartId="4.[192,288,1875,1897]" captionTargetBox="[192,1407,299,1723]" captionTargetId="figure-48@4.[192,1440,299,1747]" captionTargetPageId="4" captionText="Figures 14. Ganelius madagascariensis (Laporte, 1840). 1) Male lectotype, dorsal habitus. 2) Female, dorsal habitus. 3) Lectotype labels. 4) Male genitalia, flagellum ~ 60 mm." figureDoi="http://doi.org/10.5281/zenodo.5169527" httpUri="https://zenodo.org/record/5169527/files/figure.png" pageId="3" pageNumber="2">Fig. 3</figureCitation>
): a) handwritten, “5440 /
<collectingCountry id="F364E2B97518B13E49B1F88E89406912" box="[691,843,1837,1861]" pageId="3" pageNumber="2">Madagascar</collectingCountry>
”; b) on underside of blue circular label, “54 | 40”; c) handwritten, “
<taxonomicName id="4C73D9AA7518B13E4A90F8ED8BF66931" box="[402,509,1870,1894]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E4A90F8ED8BF66931" box="[402,509,1870,1894]" italics="true" pageId="3" pageNumber="2">Nigidius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA7518B13E4914F8ED88F86931" authorityName="Laporte" authorityYear="1840" box="[534,755,1870,1894]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B7518B13E4914F8ED88F86931" box="[534,755,1870,1894]" italics="true" pageId="3" pageNumber="2">madagascariensis</emphasis>
</taxonomicName>
/ Laporte”; d) “Museum Paris / Madagascar”; e) on red paper “TYPE SBL 14”; f) on red paper “
<taxonomicName id="4C73D9AA7518B13E49A7F8CD890A69D1" authorityName="De Haan" authorityYear="1833" box="[677,769,1902,1926]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B7518B13E49A7F8CD890A69D1" box="[677,769,1902,1926]" italics="true" pageId="3" pageNumber="2">Eudora</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA7518B13E481AF8CD89FE69D1" authorityName="Laporte" authorityYear="1840" box="[792,1013,1902,1926]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B7518B13E481AF8CD89FE69D1" box="[792,1013,1902,1926]" italics="true" pageId="3" pageNumber="2">madagascariensis</emphasis>
</taxonomicName>
<specimenCount id="9D7569A07518B13E48FEF8CD8E1B69D2" box="[1020,1040,1902,1925]" pageId="3" pageNumber="2" type="male"></specimenCount>
/
<bibRefCitation id="EFE2DFD87518B13E4F25F8CD8ED969D1" author="Laporte, F." box="[1063,1234,1902,1926]" pageId="3" pageNumber="2" pagination="1 - 564" refId="ref7518" refString="Laporte, F. 1840. Histoire Naturelle des Insectes Coleopteres. Avec une introduction renfermant l'Anatomie et la Physiologie des Animaux Articules, par M. Brulle. P. Dumenil; Paris 2: 1 - 564." type="journal article" year="1840">Laporte, 1840</bibRefCitation>
/ LECTOTYPE / des. M.J. Paulsen 2017”; g) “
<taxonomicName id="4C73D9AA751EB1384920FF61889B6E8D" authorityName="Benesh" authorityYear="1955" box="[546,656,194,218]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB1384920FF61889B6E8D" box="[546,656,194,218]" italics="true" pageId="5" pageNumber="4">Ganelius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB13849AAFF61898E6E8D" authorityName="Laporte" authorityYear="1840" box="[680,901,194,218]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB13849AAFF61898E6E8D" box="[680,901,194,218]" italics="true" pageId="5" pageNumber="4">madagascariensis</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFE2DFD8751EB13848A7FF618E596E8D" author="Laporte, F." box="[933,1106,194,218]" pageId="5" pageNumber="4" pagination="1 - 564" refId="ref7518" refString="Laporte, F. 1840. Histoire Naturelle des Insectes Coleopteres. Avec une introduction renfermant l'Anatomie et la Physiologie des Animaux Articules, par M. Brulle. P. Dumenil; Paris 2: 1 - 564." type="journal article" year="1840">Laporte, 1840</bibRefCitation>
) / det.
<determinerName id="7E104EED751EB1384FAAFF618F416E8D" box="[1192,1354,194,218]" pageId="5" pageNumber="4">M.J. Paulsen</determinerName>
2017”
</materialsCitation>
.
<materialsCitation id="3B1BA874751EB1384BC3FF4188C16F0A" ID-GBIF-Occurrence="3776282339" collectionCode="MNHN" country="Madagascar" determinerName="M. J. Paulsen" lastPageId="5" lastPageNumber="4" location="Madagascar" pageId="5" pageNumber="2" specimenCount="1" specimenCount-female="1" typeStatus="paralectotype">
<typeStatus id="54C81C8B751EB1384BC3FF418B606EAD" box="[193,363,226,250]" pageId="5" pageNumber="4" type="paralectotype">Paralectotype</typeStatus>
female,
<collectionCode id="ED623AEC751EB1384AD3FF4088246EAD" box="[465,559,227,250]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="4">MNHN</collectionCode>
, labeled: a) on underside of blue circular label, “54 | 40”; b) handwritten, “
<taxonomicName id="4C73D9AA751EB1384BC8FEA08B3E6F4C" box="[202,309,259,283]" class="Insecta" family="Lucanidae" genus="Nigidius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB1384BC8FEA08B3E6F4C" box="[202,309,259,283]" italics="true" pageId="5" pageNumber="4">Nigidius</emphasis>
</taxonomicName>
/
<emphasis id="B9077E3B751EB1384A4EFEA088036F4C" box="[332,520,259,283]" italics="true" pageId="5" pageNumber="4">madagascarien</emphasis>
/
<emphasis id="B9077E3B751EB138491DFEA788436F4C" box="[543,584,260,283]" italics="true" pageId="5" pageNumber="4">-sis</emphasis>
Laporte /
<collectingCountry id="F364E2B9751EB13849CAFEA0896B6F4C" box="[712,864,259,283]" pageId="5" pageNumber="4">Madagascar</collectingCountry>
”; c) on yellow paper “
<taxonomicName id="4C73D9AA751EB1384F6AFEA08ECF6F4C" authorityName="De Haan" authorityYear="1833" box="[1128,1220,259,283]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB1384F6AFEA08ECF6F4C" box="[1128,1220,259,283]" italics="true" pageId="5" pageNumber="4">Eudora</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB1384FD9FEA08AEA6F6B" authorityName="Laporte" authorityYear="1840" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB1384FD9FEA08AEA6F6B" italics="true" pageId="5" pageNumber="4">madagascariensis</emphasis>
</taxonomicName>
<specimenCount id="9D7569A0751EB1384BEAFE868AF76F6B" box="[232,252,293,316]" pageId="5" pageNumber="4" type="female"></specimenCount>
/
<bibRefCitation id="EFE2DFD8751EB1384A11FE868BB66F6B" author="Laporte, F." box="[275,445,292,316]" pageId="5" pageNumber="4" pagination="1 - 564" refId="ref7518" refString="Laporte, F. 1840. Histoire Naturelle des Insectes Coleopteres. Avec une introduction renfermant l'Anatomie et la Physiologie des Animaux Articules, par M. Brulle. P. Dumenil; Paris 2: 1 - 564." type="journal article" year="1840">Laporte, 1840</bibRefCitation>
/ PARALECTOTYPE / des. M.J. Paulsen 2017”; d) “
<taxonomicName id="4C73D9AA751EB1384F3DFE878EA66F6B" authorityName="Benesh" authorityYear="1955" box="[1087,1197,292,316]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="B9077E3B751EB1384F3DFE878EA66F6B" box="[1087,1197,292,316]" italics="true" pageId="5" pageNumber="4">Ganelius</emphasis>
</taxonomicName>
/
<taxonomicName id="4C73D9AA751EB1384FC1FE878FAB6F6B" authorityName="Laporte" authorityYear="1840" box="[1219,1440,292,316]" class="Insecta" family="Lucanidae" genus="Eudora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB1384FC1FE878FAB6F6B" box="[1219,1440,292,316]" italics="true" pageId="5" pageNumber="4">madagascariensis</emphasis>
</taxonomicName>
/ (
<bibRefCitation id="EFE2DFD8751EB1384BD5FEE68B886F0A" author="Laporte, F." box="[215,387,325,349]" pageId="5" pageNumber="4" pagination="1 - 564" refId="ref7518" refString="Laporte, F. 1840. Histoire Naturelle des Insectes Coleopteres. Avec une introduction renfermant l'Anatomie et la Physiologie des Animaux Articules, par M. Brulle. P. Dumenil; Paris 2: 1 - 564." type="journal article" year="1840">Laporte, 1840</bibRefCitation>
) / det.
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2017”
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<emphasis id="B9077E3B751FB1394BC2F8F08B5C693F" bold="true" box="[192,343,1875,1897]" pageId="4" pageNumber="3">Figures 14.</emphasis>
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<emphasis id="B9077E3B751FB1394854F8F08966693E" bold="true" box="[854,877,1875,1897]" pageId="4" pageNumber="3">1)</emphasis>
Male lectotype, dorsal habitus.
<emphasis id="B9077E3B751FB1394FD8F8F08EFA693E" bold="true" box="[1242,1265,1875,1897]" pageId="4" pageNumber="3">2)</emphasis>
Female, dorsal habitus.
<emphasis id="B9077E3B751FB1394A20F8CC8B3369D2" bold="true" box="[290,312,1903,1925]" pageId="4" pageNumber="3">3)</emphasis>
Lectotype labels.
<emphasis id="B9077E3B751FB1394902F8CC881C69D2" bold="true" box="[512,535,1903,1925]" pageId="4" pageNumber="3">4)</emphasis>
Male genitalia, flagellum ~ 60 mm.
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<emphasis id="B9077E3B751EB1384BC2FE248BB76FC9" bold="true" box="[192,444,390,415]" pageId="5" pageNumber="4">Non-type material</emphasis>
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(
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<collectingMunicipality id="6BA83853751EB1384FF5FDEF8FAB6C34" box="[1271,1440,588,611]" pageId="5" pageNumber="4">Maroantsetra</collectingMunicipality>
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(
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,
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)
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;
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<collectingMunicipality id="6BA83853751EB13849CAFDCE89476CD2" box="[712,844,621,645]" pageId="5" pageNumber="4">Ste. Marie</collectingMunicipality>
de
<taxonomicName id="4C73D9AA751EB138487CFDCE8E1F6CD2" authority="Paulsen, 2017" authorityName="Paulsen" authorityYear="2017" box="[894,1044,621,645]" genus="Madagascar" pageId="5" pageNumber="1" rank="genus" status="gen. nov.">Madagascar</taxonomicName>
(
<specimenCount id="9D7569A0751EB1384F29FDCE8E476CD2" box="[1067,1100,621,645]" pageId="5" pageNumber="4" type="female">1♀</specimenCount>
,
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).
<collectingRegion id="49B76CCB751EB1384FD0FDCE8F586CD2" box="[1234,1363,621,645]" country="Madagascar" name="Toliara" pageId="5" pageNumber="4">TOLIARA</collectingRegion>
:
<location id="8EACF4F2751EB1384E63FDCE8B7A6CF2" LSID="urn:lsid:plazi:treatment:03DA133F7518B1384BC2F92F8EB36B2E:8EACF4F2751EB1384E63FDCE8B7A6CF2" municipality="Ste. Marie" name="Cape Andrahomana" pageId="5" pageNumber="4" stateProvince="Toliara">Cape Andrahomana</location>
(
<specimenCount id="9D7569A0751EB1384A80FD2D8BAE6CF1" box="[386,421,654,678]" pageId="5" pageNumber="4" type="male">1♂</specimenCount>
,
<collectionCode id="ED623AEC751EB1384AB1FD2D88196CF1" box="[435,530,654,678]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="5" pageNumber="4">MNHN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1BA874751EB1384927FD2D8EA06CF1" ID-GBIF-Occurrence="3776282345" box="[549,1195,653,678]" collectionCode="MNHN" location="Taolagnaro" pageId="5" pageNumber="4" specimenCount="1" specimenCount-female="1" stateProvince="Toliara">
<locationDeviation id="9A7EBB0B751EB1384927FD2D89566CF2" box="[549,861,653,678]" location="Fort Dauphin" pageId="5" pageNumber="4">
<quantity id="4C8B0FCC751EB1384927FD2D88796CF2" box="[549,626,653,678]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="4" unit="km" value="20.0">20 km</quantity>
N of Fort Dauphin
</locationDeviation>
[
<location id="8EACF4F2751EB1384869FD2E89F06CF2" LSID="urn:lsid:plazi:treatment:03DA133F7518B1384BC2F92F8EB36B2E:8EACF4F2751EB1384869FD2E89F06CF2" box="[875,1019,653,677]" name="Taolagnaro" pageId="5" pageNumber="4" stateProvince="Toliara">Taolagnaro</location>
] (
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,
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C369F1A2751EB1384BC2FD6C88716DFB" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BCCA229751EB1384BC2FD6C88716DFB" blockId="5.[190,1440,719,940]" pageId="5" pageNumber="4">
<emphasis id="B9077E3B751EB1384BC2FD6C8B5B6CB0" bold="true" box="[192,336,719,743]" pageId="5" pageNumber="4">Diagnosis.</emphasis>
The male mandibles have an acute basal tooth on the dorsal ramus that is near the median denticle (
<figureCitation id="1348BEAC751EB1384A2DFD538B7F6D5F" box="[303,372,752,776]" captionStart="Figures 14" captionStartId="4.[192,288,1875,1897]" captionTargetBox="[192,1407,299,1723]" captionTargetId="figure-48@4.[192,1440,299,1747]" captionTargetPageId="4" captionText="Figures 14. Ganelius madagascariensis (Laporte, 1840). 1) Male lectotype, dorsal habitus. 2) Female, dorsal habitus. 3) Lectotype labels. 4) Male genitalia, flagellum ~ 60 mm." figureDoi="http://doi.org/10.5281/zenodo.5169527" httpUri="https://zenodo.org/record/5169527/files/figure.png" pageId="5" pageNumber="4">Fig. 1</figureCitation>
,
<figureCitation id="1348BEAC751EB1384A83FD538BAA6D5F" box="[385,417,752,776]" captionStart="Figures 2627" captionStartId="16.[192,298,837,861]" captionTargetBox="[540,1123,396,805]" captionTargetId="figure-47@16.[509,1123,386,805]" captionTargetPageId="16" captionText="Figures 2627. Comparison of male mandibles. Arrows indicate relative position of dorsal basal teeth and median denticles. 26) Ganelius madagascariensis. 27) G. oberndorferi." figureDoi="http://doi.org/10.5281/zenodo.5169537" httpUri="https://zenodo.org/record/5169537/files/figure.png" pageId="5" pageNumber="4">26</figureCitation>
), and this tooth remains subacute in smaller specimens. In males of
<taxonomicName id="4C73D9AA751EB1384FDCFD538F9C6D5F" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[1246,1431,752,776]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB1384FDCFD538F9C6D5F" box="[1246,1431,752,776]" italics="true" pageId="5" pageNumber="4">G. oberndorferi</emphasis>
</taxonomicName>
, the homologous tooth is poorly developed and obtuse (
<figureCitation id="1348BEAC751EB1384858FCB289A96D7E" box="[858,930,785,809]" captionStart="Figures 58" captionStartId="6.[192,288,1875,1897]" captionTargetBox="[208,1419,355,1726]" captionTargetId="figure-48@6.[191,1439,311,1741]" captionTargetPageId="6" captionText="Figures 58. Ganelius oberndorferi (Nonfried, 1892). 5) Male neotype, dorsal habitus. 6) Female, dorsal habitus. 7) Neotype labels. 8) Male genitalia, flagellum, 41 mm." figureDoi="http://doi.org/10.5281/zenodo.5169529" httpUri="https://zenodo.org/record/5169529/files/figure.png" pageId="5" pageNumber="4">Fig. 5</figureCitation>
,
<figureCitation id="1348BEAC751EB13848ADFCB289DB6D7E" box="[943,976,785,809]" captionStart="Figures 2627" captionStartId="16.[192,298,837,861]" captionTargetBox="[540,1123,396,805]" captionTargetId="figure-47@16.[509,1123,386,805]" captionTargetPageId="16" captionText="Figures 2627. Comparison of male mandibles. Arrows indicate relative position of dorsal basal teeth and median denticles. 26) Ganelius madagascariensis. 27) G. oberndorferi." figureDoi="http://doi.org/10.5281/zenodo.5169537" httpUri="https://zenodo.org/record/5169537/files/figure.png" pageId="5" pageNumber="4">27</figureCitation>
). Females (
<figureCitation id="1348BEAC751EB1384F67FCB28EA76D7E" box="[1125,1196,785,809]" captionStart="Figures 14" captionStartId="4.[192,288,1875,1897]" captionTargetBox="[192,1407,299,1723]" captionTargetId="figure-48@4.[192,1440,299,1747]" captionTargetPageId="4" captionText="Figures 14. Ganelius madagascariensis (Laporte, 1840). 1) Male lectotype, dorsal habitus. 2) Female, dorsal habitus. 3) Lectotype labels. 4) Male genitalia, flagellum ~ 60 mm." figureDoi="http://doi.org/10.5281/zenodo.5169527" httpUri="https://zenodo.org/record/5169527/files/figure.png" pageId="5" pageNumber="4">Fig. 2</figureCitation>
) are difficult to distinguish from those of
<taxonomicName id="4C73D9AA751EB1384AD7FC9188846D1D" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[469,655,818,842]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB1384AD7FC9188846D1D" box="[469,655,818,842]" italics="true" pageId="5" pageNumber="4">G. oberndorferi</emphasis>
</taxonomicName>
, but the ocular canthus of female
<taxonomicName id="4C73D9AA751EB1384F30FC918F396D1D" box="[1074,1330,818,842]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB1384F30FC918F396D1D" box="[1074,1330,818,842]" italics="true" pageId="5" pageNumber="4">G. madagascariensis</emphasis>
</taxonomicName>
(
<figureCitation id="1348BEAC751EB1384E40FC918F9C6D1D" box="[1346,1431,818,842]" captionStart="Figures 2832" captionStartId="16.[192,302,1675,1699]" captionTargetBox="[230,1426,1030,1622]" captionTargetId="figure-48@16.[192,1440,1020,1634]" captionTargetPageId="16" captionText="Figures 2832. Comparison of ocular canthi, females. 28) Ganelius madagascariensis. 29) G. oberndorferi. 30) G. gnamptus. 31) G. zombi. 32) Agnelius nageli." figureDoi="http://doi.org/10.5281/zenodo.5169539" httpUri="https://zenodo.org/record/5169539/files/figure.png" pageId="5" pageNumber="4">Fig. 28</figureCitation>
) is straight to weakly concave, while that of
<taxonomicName id="4C73D9AA751EB13849D2FCF089816D3C" baseAuthorityName="Nonfried" baseAuthorityYear="1892" box="[720,906,851,875]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="oberndorferi">
<emphasis id="B9077E3B751EB13849D2FCF089816D3C" box="[720,906,851,875]" italics="true" pageId="5" pageNumber="4">G. oberndorferi</emphasis>
</taxonomicName>
is distinctly concave, and the suprantennal tubercle is less elevated in
<taxonomicName id="4C73D9AA751EB138490DFCD789046DDC" box="[527,783,883,907]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB138490DFCD789046DDC" box="[527,783,883,907]" italics="true" pageId="5" pageNumber="4">G. madagascariensis</emphasis>
</taxonomicName>
. The male genitalia possess a flagellum that is much longer than the insect itself (
<figureCitation id="1348BEAC751EB1384926FC3788676DFB" box="[548,620,916,940]" captionStart="Figures 14" captionStartId="4.[192,288,1875,1897]" captionTargetBox="[192,1407,299,1723]" captionTargetId="figure-48@4.[192,1440,299,1747]" captionTargetPageId="4" captionText="Figures 14. Ganelius madagascariensis (Laporte, 1840). 1) Male lectotype, dorsal habitus. 2) Female, dorsal habitus. 3) Lectotype labels. 4) Male genitalia, flagellum ~ 60 mm." figureDoi="http://doi.org/10.5281/zenodo.5169527" httpUri="https://zenodo.org/record/5169527/files/figure.png" pageId="5" pageNumber="4">Fig. 4</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C369F1A2751EB1384BC2FC758EB36B2E" pageId="5" pageNumber="4" type="discussion">
<paragraph id="8BCCA229751EB1384BC2FC7589816B42" blockId="5.[190,1441,982,1401]" pageId="5" pageNumber="4">
<emphasis id="B9077E3B751EB1384BC2FC758B486DB9" bold="true" box="[192,323,982,1006]" pageId="5" pageNumber="4">Remarks.</emphasis>
<bibRefCitation id="EFE2DFD8751EB1384A45FC748BF26DB9" author="Laporte, F." box="[327,505,982,1006]" pageId="5" pageNumber="4" pagination="1 - 564" refId="ref7518" refString="Laporte, F. 1840. Histoire Naturelle des Insectes Coleopteres. Avec une introduction renfermant l'Anatomie et la Physiologie des Animaux Articules, par M. Brulle. P. Dumenil; Paris 2: 1 - 564." type="journal article" year="1840">Laporte (1840)</bibRefCitation>
described this species from an unknown number of specimens. Two specimens from MNHN, a male and female of nearly identical size, bear labels corresponding to the accession log entry for “Séries 1854-40”, which translates to 60 insect specimens from
<taxonomicName id="4C73D9AA751EB1384F3BFBBB8EC46A67" authority="Paulsen, 2017" authorityName="Paulsen" authorityYear="2017" box="[1081,1231,1048,1072]" genus="Madagascar" pageId="5" pageNumber="1" rank="genus" status="gen. nov.">Madagascar</taxonomicName>
bought from Mr. Deyrolle for 339 francs. The
<typeStatus id="54C81C8B751EB1384925FB9A889D6A06" box="[551,662,1081,1105]" pageId="5" pageNumber="4" type="lectotype">lectotype</typeStatus>
male chosen bears a type label added by S. Boucher (MNHN, pers. comm.) who selected it as certainly having been studied by Laporte while in the Deyrolle collection before accession. The other specimen is a female that bears similar labels and therefore must be treated as a
<typeStatus id="54C81C8B751EB1384A62FB3888006AE4" box="[352,523,1179,1203]" pageId="5" pageNumber="4" type="paralectotype">paralectotype</typeStatus>
. The
<typeStatus id="54C81C8B751EB1384953FB3888CB6AE4" box="[593,704,1179,1203]" pageId="5" pageNumber="4" type="lectotype">lectotype</typeStatus>
lacks specific locality data, however it is indistinguishable from males from the locality of Ambodivoangy and others around Antongil Bay. Although somewhat variable in mandibular form, additional specimens from farther south along the entire eastern side of the island cannot be separated from
<taxonomicName id="4C73D9AA751EB1384980FB5D89886B41" box="[642,899,1278,1302]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB1384980FB5D89886B41" box="[642,899,1278,1302]" italics="true" pageId="5" pageNumber="4">G. madagascariensis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BCCA229751EB1384BF2FABC8EB36B2E" blockId="5.[190,1441,982,1401]" pageId="5" pageNumber="4">
Two female specimens of
<taxonomicName id="4C73D9AA751EB1384933FABC89386B61" box="[561,819,1310,1334]" class="Insecta" family="Lucanidae" genus="Ganelius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B9077E3B751EB1384933FABC89386B61" box="[561,819,1310,1334]" italics="true" pageId="5" pageNumber="4">G. madagascariensis</emphasis>
</taxonomicName>
are imaged and listed as types on the website of IRSNB. I have examined the
<specimenCount id="9D7569A0751EB138492BFAE388D46B00" box="[553,735,1343,1367]" pageId="5" pageNumber="4" type="generic">two specimens</specimenCount>
, which appear to be more recent Oberthür specimens and determined that they are not part of the
<typeStatus id="54C81C8B751EB13849B7FAC288E16B2E" box="[693,746,1377,1401]" pageId="5" pageNumber="4">type</typeStatus>
series. They are not labeled as
<typeStatus id="54C81C8B751EB1384F6DFAC28EBF6B2E" box="[1135,1204,1377,1401]" pageId="5" pageNumber="4">types</typeStatus>
.
</paragraph>
</subSubSection>
</treatment>
</document>